Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Zapotecanillus montanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 52miles N of Oaxaca, Ber.202, sink litter, 17 May 1971 S.B.Peck collector (CMNH)
Scale bar = 0.5mm
Copyright © Igor Sokolov

Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro

Typhlocharis laurentii Magrini, 2000 Paratype
Unsexed
España, Almeria, Sierra Almagrera ; carretera N332, 30.XI.98, leg. Fancello (MNHN, Antoine Mantilleri, 2016)
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Nesamblyops sp.
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Pelodiaetus sulcatipennis Jeannel, 1937
Male adult
New Zealand, Otago, Outram
Copyright © Igor Sokolov

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Anillus florentinus florentinus Dieck, 1869
Unsexed
Emilia Romagna, Sasseleone (BO), 26.3.2002 350m, A.Degiovanni leg. (cAA)
Copyright © Alexander Anichtchenko

Megastylulus pivai Giachino & Sciaky, 2003 Holotype
Male adult
leg. E. Piva - Isl. St.Lucia Caraibi
Copyright © Vittorino Monzini

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
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Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Philochthus) pallidiveste Carret, 1905
Male adult
Volgogrda Area, Pallasovka Distr., near Elton Vill. 4.V.2009 leg. K.Makarov
L = 3.8
Copyright © Kirill Makarov

Bembidion (Hirmoplataphus) nigrum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphanes) adygorum Belousov & Sokolov, 1996
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Zecillenus) albescens (Bates 1878)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Notaphemphanes) ephippium (Marsham, 1802)
Female adult

Copyright © Kirill Makarov

Bembidion (Terminophanes) kaschmirense Netolitzky, 1920
Female adult
India, Uttarakhand, 5 km NE Govind Ghat, road to Gangaria, 9-10.6.2011, Anichtchenko A. leg. det. J. Schmidt 2015 8 mm
Copyright © Alexander Anichtchenko

Neolovricia ozimeci Lakota, Jazlic & Moravec, 2009 Holotype
Female adult
Spilja u Radinovcima, Dedici, Rastovac, Biokovo, 25.10.2006 Coleoptera (1) Leg: Roman Ozimec. Deposited in the Croatian Natural History Museum, Zagreb. Image from Lakota, Jazlic & Moravec, 2009
Copyright © - - -

Bembidion (Bembidionetolitzkya) lhai Schmidt, 2018 Paratype
Male adult
“TIBET South Centr. 17-20.VI.07 / Budha Vall. N of Yangpachem / ca. 30°10’56N 90°29’21E / 5000-5200m, leg. J. Schmidt” (CSCHM).
Copyright © Alexander Anichtchenko

Odontium (Odontium) striatum (Fabricius, 1792)
Female adult
Latvia, Dunava
Copyright © Alexander Anichtchenko

Ocys reuteri Toledano & Wrase, 2016 Holotype
Female adult
“N-Iraq, S Rawanduz/ XI.2007 – III.2008 / 36°30’N, 44°36’E / 1400m, leg. C. Reuter” // Ocys reuteri n.sp./ Toledano & Wrase des. 2016” (CWBE).
Copyright © Luca Toledano

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platidiolus vandykei Kurnakov, 1960
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Patrobus cinctus Motschulsky, 1860
Female adult
Jewish Aut. Reg., Obluchie Distr., 3-5 km upper Radde Vill. on Lagar Riv., forest 24.07.2004 I.Melnik leg.
L=9.5 mm
Copyright © Kirill Makarov

Deltomerus (Deltomerus) pseudoplatynus pseudoplatynus s.str.
Female adult
NW-Caucasus,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
long - 11mm.
Copyright © Ruslan Panin

Deltomerus (Deltomerus) tatricus L. Miller, 1859: 306
Male adult
N Slovakia; Vysoké Tatry Mts., 10mm
Copyright © Aleš Sedláček

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Platidiolus cf. borealis Zamotajlov, 2001
Female adult
Russia, Khabarovsk reg., Verhnebureinski distr. Bureya river valley, Ussomah river, 22.7.2006 (anal sternite with 3 pairs of setae). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Platidiolus rufus Chaudoir, 1878
Male adult
Russia, Hamar-Daban Mt. rng., Komarinskiy Mt., upper flow of Slyudyanka riv. 15.7.2013, A. Shavrin leg. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Deltomerodes conaensis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Elaphropus) latissimus (Motschulsky, 1851)
Female adult
Kunashiri Isl., Calder of Golovnina Volcano, S coast od Goryachee Lake 43°52'19"N 145°29'43"E 12.VIII.2008 leg.K.Makarov, A.Zaitzev
2.0 mm
Copyright © Kirill Makarov

Tachyura (Amaurotachys) notaphoida (Bates, 1886)
Male adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
Copyright © Roman Dudko

Pericompsus (Upocompsus) habitans (Sloane, 1896) Syntypes
Male adult

Copyright © Museums Victoria

Setitachys macrops Baehr, 2016 Paratype
Female adult
Qld. Bentinck Is. Rokoti, 16 June 1963. P. Aitken, N. B. Tindale
Copyright © Martin Baehr

Elaphropus (Nototachys) occidentalis Boyd & Erwin, 2016 Paratype
Male adult
Perú, Madre de Dios, Pakitza
Copyright © Terry Erwin

Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Polyderis) laeva (Say, 1823)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Paratachyta) coracina (Putzeys, 1875)
Male adult
Malaysia, Langkawi, environment of Hotel Berjaya, leg. U.Schmidt, 7.-22.XI.2009; det. M.Baehr, 2010; Photo: U.Schmidt, 2021; 2.1mm
Copyright © Udo Schmidt

Tachyta (Paratachyta) quadrinotata Baehr, 2016 Holotype
Female adult
320 Ilam Distr., betw. Hodia Khola Valley to Sktia, 250–500 m, Shorea forest, dry, 7 Apr 1988, MARTENS & SCHAWALLER leg. / Nepal-Expeditionen JOCHEN MARTENS (SMNS).
Copyright © Martin Baehr

Tachyura (Tachylopha) ovatus (Motschulsky, 1851)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. leg & det. (cAA). 2.2 mm
Copyright © Alexander Anichtchenko

Trechus (Trechus) matejkai Vsetecka, 1938: 81
Female adult
SK, C Slovakia, Nízké Tatry Mts.; 3mm
Copyright © Aleš Sedláček

Trechus (Trechus) arnoldii Belousov, 1987: 594
Male adult

Copyright © Kirill Makarov

Xuedytes bellus Tian & Huang, 2017 Holotype
Male adult
Cave II, southeastern Du’an Yao Autonomous County, Hechi Shi (=Prefecture), northern Guangxi Zhuang Autonomous Region, southern China, VIII-08-2017, leg. Mingyi Tian, Sunbin Huang, Dianmei Wang and Mengzhen Chen. Deposited in the insect collection of South China Agricultural University, Guangzhou. Image from Tian, Huang & Wang, 2017.
Copyright © - - -

Geotrechus orpheus Dieck, 1869
Male adult
France, Ariege, Aven de Santhe Catherine, VIII.1981. Leg. Sciaky (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Rakantrechus (Paratrechiama) tofaceus Ueno, 1970b: 106
Male adult
Japan, Kumamoto Ken
Copyright © Vittorino Monzini

Orotrechus jamae C. Etonti et M. Etonti, 1979: 27
Male adult
NE Italy, Belluno, Prealps, Cansiglio plateau, Bus dell'Ors, 19.V.2013, L=6.2mm
Copyright © Denis Khrapov

Dracotrechus fontanus Ueno, 2010
Male adult

Copyright © Vittorino Monzini

Trechus (Trechus) mattisi Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Bale Mts, Harenna Forest, E Rira, alt. 3150 m, 06°44′51″ N, 39°46′40″ E, Feb. 2007
Copyright © Joachim Schmidt

Afrotrechus (Abyssiniopsis) bunae Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Oromia province, Belleta forest (degraded forest, 25 km WSW of Jimma), 7˚31’50’’N, 36˚31’46’’E, 1800 m a.s.l., VIII.2011, E. Quéinnec & C. Reeb leg.
Copyright © Eric Quéinnec

Aphaenops (Geotrechidius) gallicus (Delarouzee, 1857)
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -

Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
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Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -