Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Hypotyphlus degiovannii Magrini, 2013 Holotype
Male adult
Italia, Lazio, Montagna di Scauri (Latina), 50 m s.l.m., 1.V.2008, P. Magrini leg. (CM).
Copyright © Paolo Magrini

Nesamblyops sp.
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Anillus florentinus florentinus Dieck, 1869
Unsexed
Emilia Romagna, Sasseleone (BO), 26.3.2002 350m, A.Degiovanni leg. (cAA)
Copyright © Alexander Anichtchenko

Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
Copyright © - - -

Stylulus sp.
Unsexed
"US Virgin Islands, St. Croix, 12.vi.1972, W.B. Muchmore Leg." Florida State Collection of Arthropods. "Stylulus sp." det. C.W. Harden 2023.
Copyright © Curt Harden

Megastylulus pivai Giachino & Sciaky, 2003 Holotype
Male adult
leg. E. Piva - Isl. St.Lucia Caraibi
Copyright © Vittorino Monzini

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Mioptachys flavicauda (Say, 1823)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Ocydromus) zolotarewi Reitter, 1910b: 313
Male adult
Russia, NW Caucas, Krasnodar reg., Tuapse distr., Podchrebetovoe vicinity
(NW Кавказ, Краснодарский край, Туапсинский р-н, окр пос. Подхребeтовое 19.V.2012 leg. М. Смирнов)
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Sinechostictus (Sinechostictus) nordmanni (Chaudoir, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Ecuadion) andersoni Toledano, 2008
Male adult
ECUADOR: Pichincha: Reserva Yanacocha, start Andean Snipe Trail, DRM voucher V100655. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Orzolina thalassophila Machado, 1987
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Sloanephila) tahitiense Liebherr & Maddison, 2013 Holotype
Male adult
(MNHN): French Polynesia: Tahiti Nui / Mt. Mauru lava tube / el. 705 m 18-IX-2006 lot 01 / 17˚38.017´S 149˚21.284´W / pyr. fog moss/liverworts / along stream J.K. Liebherr // HOLOTYPE / Bembidion / tahitiense / Liebherr & / Maddison 2012 (black-bordered red label).
Copyright © David Maddison

Bembidion (Chlorodium) difforme (Motschulsky, 1844)
Female adult
Russia, Burjatia, Muhor-Shibirsk distr., 5km N Nikolsk vill., salt marsh, 27.8.1998, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Erwiniana irisculptilis (Erwin, 1994)
Unsexed
ECUADOR, Parque Nacional Yasunì
Copyright © Gianni Allegro

Bembidion (Euperyphus) scapulare lomnickii Netolitzky, 1916: 260
Male adult
W-Ukraine, Transcarpathian region, Vynogradiv distr., bank of Tisza river near Mala Kopanya vill., 5-7.VI.2018, Panin.R leg., L=5,8 mm
Copyright © Denis Khrapov

Bembidion (Hirmoplataphus) hirmocaelum Chaudoir, 1850
Male adult
Primorski reg., Kedrovaya riv. det. Sundukov, 2023
Copyright © Kirill Makarov

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Deltomerus (Deltomerus) tatricus L. Miller, 1859: 306
Male adult
N Slovakia; Vysoké Tatry Mts., 10mm
Copyright © Aleš Sedláček

Platidiolus rufus Chaudoir, 1878
Male adult
Russia, Hamar-Daban Mt. rng., Komarinskiy Mt., upper flow of Slyudyanka riv. 15.7.2013, A. Shavrin leg. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
Copyright © - - -

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Patrobus septentrionis volgensis Zamotajlov & Isaev, 2006
Male adult

Copyright © Igor Solodovnikov

Deltomerodes conaensis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Diplous (Diplous) sibiricus atratus Habu, 1951
Male adult

Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Tachyura (Amaurotachys) notaphoida (Bates, 1886)
Male adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
Copyright © Roman Dudko

Lymnastis galilaeus Piochard de la Brullerie, 1876
Male adult

Copyright © Gianni Allegro

Tachyura ascendens (Alluaud, 1917)
Female adult
TANZANIA, Kilimanjaro, Nkuansa, grassland,1660m, 3.7.2011, R. Felix det. as Elaphropus (Tachyura) exiguus (BOHEMAN 1848)
Copyright © Peter Schüle

Tachyxysta howdenorum Boyd & Erwin, 2016 Holotype
Male adult
México, Chiapas, El Aguacero.
Copyright © Terry Erwin

Tachys (Polyderis) laeva (Say, 1823)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) fulvicollis (Dejean, 1831)
Female adult
NW-Ukraine, Volhynia, Turiysk env ., into the light, 6.VII.2015, Voytko P. leg./ R.Panin det., L=3,3 мм
Copyright © Denis Khrapov

Elaphropus (Elaphropus) globulus (Dejean, 1831)
Male adult
Spain, Granada, rio Darro, barranco de Teatino, 20.V.2005, Anichtchenko A. leg & det. (cAA). 1.8 mm
Copyright © Alexander Anichtchenko

Tachyura (Tachyura) thoracicus (Kolenati, 1845)
Female adult
India, Uttarakhand, 14 km E Uttarkashi, 1500m, 10-12.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr as octostriata (Netolitzky, 1929)
Copyright © Alexander Anichtchenko

Tachyura (Tachylopha) ovatus (Motschulsky, 1851)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. leg & det. (cAA). 2.2 mm
Copyright © Alexander Anichtchenko

Queinnectrechus (Gaoligongtrechus) balli Deuve and Kavanaugh, 2016 Holotype
Male adult
CHINA, Yunnan Province, Gaoligong Shan, Nujiang Prefecture, Nujiang State Nature Reserve, Dulong/Gongshan Yakou [= Qiqi/Dulong divide] area, 21 airkm w of Gongshan,”/ “N27.69655°/ E98.45389°, 3300-3680m, 16-17 July 2000, Stop#00-24C, D.H. Kavanaugh, C.E. Griswold, Liang H.-B., D. Ubick, & Dong D.-Z. collectors (IOZ). Image from Deuve et al, 2016
Copyright © - - -

Doderotrechus ghilianii ghilianii Fairmaire, 1859a: 26
Male adult
Piemonte, Valle Po, Crissolo (TO), Gr. Buco di Valenza, 5.V.1995 (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Pheggomisetes globiceps Buresch, 1925
Male adult
Bulgaria, Ponor plannina, Dusnica Pestera cave, 30/7/2004, Vasiljev leg.
Copyright © Vittorino Monzini

Orcusiella prokletijensis Pavićević, Lohaj & Popović, 2020 Holotype Paratype
Male+Female adult
Montenegro, Prokletije Mts., Bjelič massif, Maja Kolata, “Jama Kolektor” pit, 2052 m a.s.l., 20. 07. 2014 - 28. 07. 2015, traps, leg. M. Popović. HOLOTYPUS Orcusiella gen. nov. prokletijensis sp. nov., D. Pavićević, R. Lohaj & M. Popović des. 2020 (CDP). Image from Pavićević, Lohaj & Popović, 2020
Copyright © - - -

Zhijinaphaenops zhaofeii Tian, Cheng & Huang, 2021
Male adult
Guizhou, Guiyang, Xifeng, Jiuzhuang, Changtu Dong cave (贵州省贵阳市息烽县九庄镇张口洞), 27°11'N, 106°29'E, 1008 m, 2019–VI–08, leg. Jingli Cheng & Mingyi Tian.
Copyright © Mingyi Tian

Trechus (Trechus) minitrechus Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Oromia, Mt Enkuolo NE slope, alt. 3200–3300 m, 07°23′45″ N, 39°22′27″ E, 8 Dec. 2016
Copyright © Joachim Schmidt

Afrotrechus (Abyssiniopsis) bunae Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Oromia province, Belleta forest (degraded forest, 25 km WSW of Jimma), 7˚31’50’’N, 36˚31’46’’E, 1800 m a.s.l., VIII.2011, E. Quéinnec & C. Reeb leg.
Copyright © Eric Quéinnec

Tianzhuaphaenops jinshanensis D. Zhao & Tian, 2016 Holotype
Male adult
Cave Jinshan Rongdong, Luihe Vallige, Bangdong Town, Tianzhu County, Qiandongnan Miao and Dong Autonomous Prefecture, Guizhou Province, 26°57'42" N/109°12'24" E, 460 m, VIII-21-2015, Mingyi Tian & Jingli Cheng leg., deposited in the insect collections of South China Agricultural University, Guangzhou (SCAU)
Copyright © Mingyi Tian

Panotrechus pilosus Tian & Zhou, 2023 Holotype
Male adult
Wojiao Dong Cave, Baotian Zhen, Panzhou Shi, Luipanshui Shi, western Guizhou, 25.40° N / 104.74° E, 1593 m, 7.VII.2021, Jingli Cheng & Mingyi Tian leg. (SCAU).
Copyright © Mingyi Tian

Cimmeritodes (Shimenrites) shimenensis Deuve & Tian, 2017 Holotype
Male adult
Chine, Hunan, Shimen, Xiannu Dong, 280 mètres, 29°38’55’’N -111°04’01’’E (Tian M. & Chen J., 6.VIII.2016), in coll. South China Agricultural University, Canton, Chine.
Copyright © Thierry Deuve

Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
Copyright © - - -

Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -