Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Zapotecanillus montanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 52miles N of Oaxaca, Ber.202, sink litter, 17 May 1971 S.B.Peck collector (CMNH)
Scale bar = 0.5mm
Copyright © Igor Sokolov

Anillinus langdoni Sokolov & Carlton, 2004
Male adult
USA-TN, Sevier Co, GSMNP, secondary site, 490 m, ex leaf et log litter, 35°43.157' N, 83°23.309' W, 19.04.2011, J.-S. Park det. I. Sokolov, 2013
Copyright © Roman Dudko

Illaphanus pteridophyticus Giachino, 2005 Holotype
Male adult

Copyright © Alexander Anichtchenko

Pelodiaetodes (Pelodiaetodes) constricticollis Sokolov, 2015 Paratype
Male adult
NZ, Northland, Radar Bush. Image from Sokolov, 2015
Copyright © - - -

Pelodiaetus sulcatipennis Jeannel, 1937
Male adult
New Zealand, Otago, Outram
Copyright © Igor Sokolov

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
Copyright © - - -

Stylulus sp.
Unsexed
"US Virgin Islands, St. Croix, 12.vi.1972, W.B. Muchmore Leg." Florida State Collection of Arthropods. "Stylulus sp." det. C.W. Harden 2023.
Copyright © Curt Harden

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Eupetedromus) dentellum (Thunberg, 1787)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Nothonepha) tetrapholeon Maddison, 2014 Paratype
Male adult
Argentina: Neuquén: Arroyo Queñi at Lago Queñi, 830m, 40.1575°S, 71.721°W, / 10–11.ii.2007. DRM 07.035. D.R. Maddison, S.A.Roig. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Terminophanes) pulcherrimum (Motschulsky, 1850)
Male adult
Caucasus, Elbrus, Itkol (CTVR). Image from Toledano L. & Gueorguiev B., 2015
Copyright © - - -

Bembidion (Testedium) laetum Brulle, 1836: pl.2
Female adult
Morocco, 65 km N Essaouria, 5 km N Jamaa Shoim, 23-25.III.2011, A. Gusakov. Фотография с сайта zin.ru\Animalia\Coleoptera.
Copyright © Maksim Smirnov

Bembidion (Pamirium) platypterum Solsky, 1874: 120
Female adult
Kazakhstan, Alma-Ata, 2.06.1990. det. Anichtchenko A. 2016
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) transhimalayanum Schmidt, 2018 Paratype
Female adult
“S TIBET, ca. 40km SW of / Yangpachem, 13.VII.2010 / Pijin moraine 5000-5500 m / 29°45’36N 90°17’54E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) dhaulagiricum Schmidt, 2018 Holotype
Male adult
“NEPAL W Dhaulagiri / below Dhule 26-2700m / 7.IX.12 leg. J. Schmidt / 28°41‘07N 82°56‘31E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Emphanes) latiplaga latiplaga Chaudoir, 1850
Female adult
NW-Ukraine, Volhynia, Turiysk env., 17-26.VI.2016, to light, Voytko P. leg./ Panin R. det., L=3.1 mm
Copyright © Denis Khrapov

Bembidion (Notaphus) mormon Hayward, 1897
Male adult

Copyright © David Maddison

Odontium (Bracteon) conicolle Motschulsky, 1844: 273
Female adult
Burjatia, Barguzin depr., Alla riv., 8.8.1997, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Patrobus cinctus Motschulsky, 1860
Female adult
Jewish Aut. Reg., Obluchie Distr., 3-5 km upper Radde Vill. on Lagar Riv., forest 24.07.2004 I.Melnik leg.
L=9.5 mm
Copyright © Kirill Makarov

Deltomerus (Deltomerus) pseudoplatynus pseudoplatynus s.str.
Female adult
NW-Caucasus,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
long - 11mm.
Copyright © Ruslan Panin

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Dimorphopatrobus ludmilae Casale et Sciaky, 1994: 47
Female adult
Tibet, Motuo, Hanmi, 8.2013. det. A. Anichtchenko, 2015 17 mm
Copyright © Alexander Anichtchenko

Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Deltomerodes ovicollis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Penetretus andalusicus (Reitter, 1897)
Male adult
Spain, Sierra Nevada, Lagunillos de la Virgen, 2850m, 18.5.2005, A. Anichtchenko leg. & det (cAA)
Copyright © Alexander Anichtchenko

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Pericompsus (Pericompsus) elegantulus (Laferte, 1848)
Male adult
Puerto Rico<br /> photo I. M. Sokolov
Copyright © Igor Sokolov

Elaphropus (Barytachys) tripunctatus (Say, 1830)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Tachyura (Amaurotachys) rhombophorus (Andrewes, 1925)
Male adult
India, Uttarakhand, 8 km NNE of Ramnagar, valley of Pipaliasot River, 420 m, 29°28,5' N, 79°10' E, 20.04.2012, R. Dudko, I. Lyubechanskii<br /> det. R. Dudko 2012
Copyright © Roman Dudko

Porotachys bisulcatus (Nicolai, 1822)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Elaphropus (Idiotachys) acutifrons Boyd & Erwin, 2016 Holotype
Female adult
Brazil, Santarem
Copyright © Terry Erwin

Polyderidius sp.
Male adult
México, San Luis Potosí
Copyright © Terry Erwin

Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) arcanicola Blackburn, 1878 Paralectotypes
Female adult
Oahu: Honolulu (Bates collection, box 389). Image from Liebherr, 2021
Copyright © James K. Liebherr

Eotachys (Jeannelys) euryodes (Bates, 1892)
Unsexed
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort leg. U.Schmidt, 14.XI.2007; det. M.Baehr, 2008; Photo: U.Schmidt, 2021
Copyright © Udo Schmidt

Costitachys inusitatus Erwin, 1974 Holotype
Male adult
Santarem, Para, Brazil, IV.27.1963 FG Werner (MCZ)
Copyright © - - -

Typhlotrechus bilimekii hacquenii Sturm, 1853
Male adult
Slovenia, Ljubljana, Gorni lg.Jama, Velika Basica, IV-IX.2003, M. Egger leg. L=7.5 mm
Copyright © Ruslan Panin

Huoyanodytes tujiaphilus Tian & Huang, 2016 Holotype
Female adult
Cave Tujiamei Dong, Huoyan Karst, Huoyan Xiang, Wulongshan Geopark, Longshan County, NW Hunan Province, China, 29°12'20.11"N/109°18'37.21"E, 427 m in altitude, VII-3-2014, leg. Mingyi Tian, Weixin Liu, Haomin Yin, Sunbin Huang & Xinhui Wang, deposited in SCAU. Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Kozlovites major Belousov et Kabak, 2016 Paratype
Male adult
pass 25 km SE Dequin
Copyright © - - -

Dongoblemus kemadongicus Deuve & Tian, 2016 Holotype
Female adult
Chine, Yunnan, Zhenxiong Xian, Chishuiyuan, grotte Kema Dong, 1730 m, 27°39’13’’N - 104°51’10’’E (Tian Mingyi, Liu Weixin, Wang Xinhui, Tang Mingruo, 23.VI.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -

Omalodera discoidalis discoidalis
Unsexed
Image from Chilean Coleoptera Biodiversity
Copyright © - - -

Anchotrechus punctipennis Jeannel, 1927
Male adult
Tenerife, Anaga, Barranco de Inguana, III.1984, leg. Sciaky (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Pheggomisetes globiceps Buresch, 1925
Male adult
Bulgaria, Ponor plannina, Dusnica Pestera cave, 30/7/2004, Vasiljev leg.
Copyright © Vittorino Monzini

Duvalius (Paraduvalius) joannidisi Casale, Giachino et M. Etonti, 1990
Female adult
Macedonia
Copyright © Vittorino Monzini

Agonotrechus sinotroglophilus Deuve, 2000
Male adult
Hanwang Dong, Zhongbai, Zengjia, Chaotian, Guangyuan, Sichuan (四川省广元市朝天区曾家镇中柏村汉王洞), 32.577297°N, 106.106979°E, 1210 m, 2018-IV-06, leg. Li He (SCAU). Image from Tian & He, 2020
Copyright © - - -

Trechus (Trechus) mattisi Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Bale Mts, Harenna Forest, E Rira, alt. 3150 m, 06°44′51″ N, 39°46′40″ E, Feb. 2007
Copyright © Joachim Schmidt

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -