Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Illaphanus pteridophyticus Giachino, 2005 Holotype
Male adult

Copyright © Alexander Anichtchenko

Typhlocharis laurentii Magrini, 2000 Paratype
Unsexed
España, Almeria, Sierra Almagrera ; carretera N332, 30.XI.98, leg. Fancello (MNHN, Antoine Mantilleri, 2016)
Copyright © - - -

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Medusapyga hehalis LaBonte, 2023 Paratype
Female adult
WASHINGTON, Thurston Co., 11.3 km NW Littlerock, elev. ~400 m, D.J. Cox and J.R. LaBonte, 6 May 2007, 46.9600°N 123.13450°W. Image from LaBonte & Maddison, 2023
Copyright © - - -

Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
Copyright © - - -

Stylulus sp.
Unsexed
"US Virgin Islands, St. Croix, 12.vi.1972, W.B. Muchmore Leg." Florida State Collection of Arthropods. "Stylulus sp." det. C.W. Harden 2023.
Copyright © Curt Harden

Anillinus alleni Sokolov & Carlton, 2017
Male adult
USA: Arkansas, Stone Co., Ozark National Forest, Blanchard Springs Rec. Area. 27 September 2024. C.W. Harden & K.D. Karns leg. Under rock during rain. / Det. C.W. Harden 2024.
Copyright © Curt Harden

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Mioptachys flavicauda (Say, 1823)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Kunashiri Isl., mouth of Medny Spring 44°16'31"N 145°56'29"E 27.VI.2008 leg. K.Makarov
L=4.3
Copyright © Kirill Makarov

Bembidion (Peryphanes) lacunarium (Zimmermann, 1869)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Nothonepha) eburneonigrum Germain, 1906
Male adult
Argentina: Neuquén: Rio Neuquén at Chos Malal, DRM voucher V100785. Image from Maddison D.R., 2014.
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Bembidion (Zeplataphus) tairuense Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Leuchydrium) tigrinum Leconte, 1879
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Plataphus) coelestinum (Motschulsky, 1844)
Female adult
N Mongolia, Tuul-Gol R., Engin Aimak Tev, 14.8.2013, I. Enuschenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) saxatile morsum Netolitzky, 1930
Male adult
Tajikistan, Khoja Obi Garm, 15.9.2011, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Hydrium) mimbres Maddison, 2020 Paratype
Male adult
USA: New Mexico: Grant Co., Gila River, Billings Vista, 1320 m, 32.8163°N, 108.6032°W, 11.viii.2005. DRM 05.043. D.R. & J.H.A. Maddison
Copyright © David Maddison

Bembidion (Armatocillenus) sauteri (Jedlicka, 1958) Holotype
Female adult
Formosa, Sauter // Anping, 1909.X. // Typus [red] // Armatocillenus sauteri n.sp. [handwritten], det. ING. JEDLIČKA [pink] (HNHM).
Copyright © - - -

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Platidiolus vandykei Kurnakov, 1960
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
Copyright © - - -

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Patrobus septentrionis volgensis Zamotajlov & Isaev, 2006
Male adult

Copyright © Igor Solodovnikov

Deltomerus (Deltomerus) corax Peyerimhoff, 1922
Male adult
Morocco, Grand Atlas, 2200-2300m, N31.13.09.5 W07.49.29.7 16-17.6.2010 Anichtchenko A. leg. & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Penetretus andalusicus (Reitter, 1897)
Male adult
Spain, Sierra Nevada, Lagunillos de la Virgen, 2850m, 18.5.2005, A. Anichtchenko leg. & det (cAA)
Copyright © Alexander Anichtchenko

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Tachyura) ocellatus (Bates, 1892)
Female adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko

Lymnastis galilaeus Piochard de la Brullerie, 1876
Male adult

Copyright © Gianni Allegro

Polyderidius sp.
Male adult
México, San Luis Potosí
Copyright © Terry Erwin

Tachyura (Tachyura) nitens (Andrewes, 1925)
Female adult
Indonesia E, W New Guinea, Doberai Peninsula, Ayamaru vill. 27 km SE Uter (Aitinyo) lakeside, 1*23'01S 132*23'09E, 175-177m, 4.IX.2015, muddy bank, between lakeside vegetation, leg. D. Telnov. 2.8 mm. det. M. Baehr, 2016
Copyright © Alexander Anichtchenko

Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) scitulus Leconte, 1848
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Paratachyta) coracina (Putzeys, 1875)
Male adult
Malaysia, Langkawi, environment of Hotel Berjaya, leg. U.Schmidt, 7.-22.XI.2009; det. M.Baehr, 2010; Photo: U.Schmidt, 2021; 2.1mm
Copyright © Udo Schmidt

Liotachys antennatus Bates, 1871
Female adult
Bolivia: Santa Cruz Prov. Andrez Ibanez Potrerillo de Guenda, Terebinto. / 3-4 October 1994 / leg. P. Bettella / det. R. Davidson. Carnegie Museum of Natural History, Pittsburg, PA, USA
Copyright © Curt Harden

Aepiblemus caeculus Belousov et Kabak, 1993b: 141
Male adult

Copyright © Kirill Makarov

Eotrechodes larisae Ueno, Lafer et Sundukov, 1995: 114
Male adult

Copyright © Kirill Makarov

Kozlovites major Belousov et Kabak, 2016 Paratype
Male adult
pass 25 km SE Dequin
Copyright © - - -

Wanoblemus wui Tian & Fang, 2016 Holotype
Male adult

Copyright © - - -

Mamesdytes glacialis Trezzi, 2007
Male adult
Guatemala
Copyright © Vittorino Monzini

Trichaphaenops (Trichaphaenops) gounellei annae Jeannel, 1949a: 30
Male adult
France, Gouffre Berger, Isere- Engins, Vercors 4.VIII.1981, leg. Regalin (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Aphaenopidius kamnikensis kamnikensis Drovenik, 1987: 53 Paratype
Male adult

Copyright © Vittorino Monzini

Paratrechus hypogeus Deuve & Moret, 2017 Holotype
Male adult
Equateur, Prov. Tungurahua, Parc national de Llanganatis, P13, Cerro Jaramillo, 4150 mètres, 1°09’40.6’’S - 78°21’23.5’’W, P. Moret, 14 février 2010. (MNHN).
Copyright © Thierry Deuve

Qianaphaenops (Qiandongaphaenops) variabilis Tian, Huang & Wang, 2015 Holotype Paratype
Male adult
cave Wanfuchangcheng Dong, in Jiangjun Shan, Pingzhuang Xiang, Cengong County, Guizhou Province, 27°24’50.91”N, 108°35’43.0”E, 830 m, 2014-VI-29, leg. Mingyi Tian, Weixin Liu, Haomin Yin, Sunbin Huang and Xinhui Wang (SCAU). Image from Tian, Huang & Wang, 2015
Copyright © Mingyi Tian

Aphaenops (Nafarroaphaenops) giraudi J. Ochs, 1938
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -