Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Austranillus macleayi (Lea, 1906) Syntypes
Female adult

Copyright © Museums Victoria

Typhlocharis laurentii Magrini, 2000 Paratype
Unsexed
España, Almeria, Sierra Almagrera ; carretera N332, 30.XI.98, leg. Fancello (MNHN, Antoine Mantilleri, 2016)
Copyright © - - -

Nesamblyops sp.
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Winklerites stevanovici Hlavac & Margini, 2017 Paratype
Male adult
Srbija [p], Dimitrovgrad, Petrlaš, Džemanska prop[ast], 02.05.2002 [h], M. Stevanović leg. Image from HLAVÁČ & MAGRINI, 2017
Copyright © - - -

Scotodipnus (Binaghites) armellinii (Ganglbauer, 1900)
Male adult
Veneto Italia
Copyright © Erminio Piva

Pelodiaetus sulcatipennis Jeannel, 1937
Male adult
New Zealand, Otago, Outram
Copyright © Igor Sokolov

Anillus florentinus andreinii Jeannel, 1937
Unsexed
Italy, Toscana, Giuncarico 8.12.96 leg Degiovanni A. (cAA)
Copyright © Alexander Anichtchenko

Stylulus sp.
Unsexed
"US Virgin Islands, St. Croix, 12.vi.1972, W.B. Muchmore Leg." Florida State Collection of Arthropods. "Stylulus sp." det. C.W. Harden 2023.
Copyright © Curt Harden

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Kunashiri Isl., mouth of Medny Spring 44°16'31"N 145°56'29"E 27.VI.2008 leg. K.Makarov
L=4.3
Copyright © Kirill Makarov

Bembidion (Neoemphanes) shimoyamai Habu, 1978
Male adult
Far East, khanka Lake env., Bol.Usachi Riv. near Novokachalinsk 5.VII.2004 R.Filimonov leg.
L=3.5 mm
Copyright © Kirill Makarov

Bembidion (Antiperyphanes) mandibulare (Solier, 1849)
Male adult
Chile: Reg. X, Chiloé: Cucao, DRM voucher V100789. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Ecuadion) chimborazonum (Bates, 1891)
Male adult
ECUADOR: Pichincha: Paso de la Virgen, DRM voucher V100793. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Zecillenus) chalmeri (Broun, 1886)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Xystosomus laevis Erwin, 1973 Holotype
Male adult
Brasilien, Nova Teutonia
Copyright © - - -

Bembidion (Omotaphus) alsium Coquerel, 1866
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Phyla) phoeniceum Huber & Marggi, 1997: 778
Female adult
Turkey, Kirish distr., Camuyuva riv. 28.9.2014, I. Enischenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) bactrianum K. Daniel, 1902
Male adult
Tadzhikistan, Varzob, Khodja obi Garm, 2000m 21-24.3.2015 (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) atlanticum atlanticum Wollaston, 1854
Male adult
Uzbekistan, Chatkal mt. rng., 6 km NW Irtash, Iertash riv., 41.10.46N 70.17.28E, 1900m, 3.8.2021, Anichtchenko A. leg. (cAA). det. L. Toledano, 2026
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platidiolus vandykei Kurnakov, 1960
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Patrobus cinctus Motschulsky, 1860
Female adult
Jewish Aut. Reg., Obluchie Distr., 3-5 km upper Radde Vill. on Lagar Riv., forest 24.07.2004 I.Melnik leg.
L=9.5 mm
Copyright © Kirill Makarov

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Platidiolus rufus Chaudoir, 1878
Male adult
Russia, Hamar-Daban Mt. rng., Komarinskiy Mt., upper flow of Slyudyanka riv. 15.7.2013, A. Shavrin leg. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Patrobus septentrionis volgensis Zamotajlov & Isaev, 2006
Male adult

Copyright © Igor Solodovnikov

Deltomerodes ovicollis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Pogonus (Pogonus) syriacus Chaudoir, 1871
Female adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 7 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Elaphropus) latissimus (Motschulsky, 1851)
Female adult
Kunashiri Isl., Calder of Golovnina Volcano, S coast od Goryachee Lake 43°52'19"N 145°29'43"E 12.VIII.2008 leg.K.Makarov, A.Zaitzev
2.0 mm
Copyright © Kirill Makarov

Elaphropus (Tachyuropsis) exaratus (Bates, 1873)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Elaphropus (Tachylopha) spenceri (Sloane, 1896) Paratype
Male adult

Copyright © Museums Victoria

Elaphropus (Idiotachys) acutifrons Boyd & Erwin, 2016 Holotype
Female adult
Brazil, Santarem
Copyright © Terry Erwin

Stigmatachys uvea Boyd & Erwin, 2016 Holotype
Female adult
Perú, Loreto, Campamento San Jacinto
Copyright © Terry Erwin

Tachyta (Paratachyta) quadrinotata Baehr, 2016 Holotype
Female adult
320 Ilam Distr., betw. Hodia Khola Valley to Sktia, 250–500 m, Shorea forest, dry, 7 Apr 1988, MARTENS & SCHAWALLER leg. / Nepal-Expeditionen JOCHEN MARTENS (SMNS).
Copyright © Martin Baehr

Tachys (Paratachys) frischi Coulon & Felix, 2009
Female adult
Iran, Khuzestan, 30 km E Shushtar, 5.5.2007, A. Anichtchenko leg. (cAA). det. J. Muilwijk, 2021
Copyright © Alexander Anichtchenko

Tachyura (Tachyura) stevensi (Andrewes, 1925)
Female adult
India, Uttarakhand, 5 km SW Rishikesh, left bank of Ganga riv., 9.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko

Tachyura (Tachylopha) ovatus (Motschulsky, 1851)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. leg & det. (cAA). 2.2 mm
Copyright © Alexander Anichtchenko

Blemus alexandrovi (Lutshnik, 1915)
Male adult
Jewish Aut. Reg., Obluchie Distr., env. Radde Vill., bank of Amur Riv. between mouth of Lagar and Belaja Riv. 26.07.2004 I.Melnik leg.<br /> L=4.0 mm
Copyright © Kirill Makarov

Guiaphaenops deuvei Tian, Feng & Wei, 2017 Holotype
Male adult
cave near Yangli Cun, Jiayou Zhen, Lingyun Xian, Baise Shi, Guangxi, 24°28'39"N, 106°37'52"E, 643 m, VII-25-2012, Mingyi Tian, Weixin Liu, Feifei Sun & Haomin Yin leg., in South China Agricultural University, Guangzhou, China (SCAU). Image from Tian, Feng & Wei, 2017
Copyright © - - -

Allegrettia comottii Monguzzi, 2011 Paratype
Male adult
10 mm
Copyright © Vittorino Monzini

Pseudaphaenops tauricus Winkler, 1912a: 134
Male adult
Crimea, Dolgorukovskaya Yalla, yama Tshokrak gully, 13/5/2007, Turbanov leg.
Copyright © Vittorino Monzini

Trechus (Trechus) humboldti Vandyke, 1945
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Panaphaenops guixicus Tian, Huang & Ma, 2021 Holotype
Male adult
the cave Zimu Dong, Danxiashan, Danxia Town, Panzhou, Liupanshui City, Guizhou Province, 25°40'0.0"N 104°37'3.6";E, 1791 m in altitude, 24.VIII.2020, leg. Yunchun Li & Yucheng Lin.
Copyright © Mingyi Tian

Casaleaphaenops liyunchuni Tian, Huang & Ma, 2021 Holotype
Male adult
cave Hei Dong, Longjiang Cun, Shenxi Town, Honghuagang District, Zunyi City, northern Guizhou Province, 2736°3.2“N 10658°4.8“E, altitude 822 m, 22.VIII.2020, leg. Yunchun Li & Yucheng Lin.
Copyright © Mingyi Tian

Uenotrechus nandanensis Deuve & Tian, 2010 Holotype
Male adult
Cave Dadongqiao Dong, Encun , Nandan County, Guangxi, XII-27-2008, leg. Zhihong Xue, Wei Lin and Lei Gao
Copyright © Mingyi Tian

Baehria separata Schmidt & Faille, 2023 Paratype
Male+Female adult
Ethiopia, Amhara, Mt. Choke, crater valley, alt. 3780–3900 m, 10°42'12"N, 37°50'58"E, 27.II.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W.
Copyright © Joachim Schmidt

Sinaphaenops (Dongaphaenops) xuxiakei Deuve & Tian, 2014 Holotype
Male adult
Chine, Guizhou, Panxian, au sud-ouest de Gucheng, Biyun Dong (= Shui Dong) (Tian Mingyi & Huang Sunbin leg., 9.VII.2013), in coll. South China Agricultural University, à Canton (Chine).
Copyright © Thierry Deuve

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -