Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Hypotyphlus degiovannii Magrini, 2013 Holotype
Male adult
Italia, Lazio, Montagna di Scauri (Latina), 50 m s.l.m., 1.V.2008, P. Magrini leg. (CM).
Copyright © Paolo Magrini

Scotodipnus alpinus Baudi di Selve, 1871
Female adult
Piemonte, Valle Po, Crissolo (CN), 27.V.2000, 1250m, fagus, leg. F. Angelini
Copyright © Alexander Anichtchenko

Winklerites stevanovici Hlavac & Margini, 2017 Paratype
Male adult
Srbija [p], Dimitrovgrad, Petrlaš, Džemanska prop[ast], 02.05.2002 [h], M. Stevanović leg. Image from HLAVÁČ & MAGRINI, 2017
Copyright © - - -

Pelodiaetus sulcatipennis Jeannel, 1937
Male adult
New Zealand, Otago, Outram
Copyright © Igor Sokolov

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Microtyphlus saxatilis Magrini, Leo & Fancello, 2004
Male adult
Italia, Sardegna, Perda Longa (NU) Baunei.
Copyright © Augusto Degiovanni

Stylulus sp.
Unsexed
"US Virgin Islands, St. Croix, 12.vi.1972, W.B. Muchmore Leg." Florida State Collection of Arthropods. "Stylulus sp." det. C.W. Harden 2023.
Copyright © Curt Harden

Anillinus alleni Sokolov & Carlton, 2017
Male adult
USA: Arkansas, Stone Co., Ozark National Forest, Blanchard Springs Rec. Area. 27 September 2024. C.W. Harden & K.D. Karns leg. Under rock during rain. / Det. C.W. Harden 2024.
Copyright © Curt Harden

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Philochthus) inoptatum Schaum, 1857a: 149
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) olegleonidovici Fassati, 1990: 169
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Bracteon) stenoderum stenoderum Bates, 1873a: 300
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Eupetedromus) dentellum (Thunberg, 1787)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum dalmatinum Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014 Holotype
Male adult
Northern Kunashir, 2.5 km NW of Cape Nelyudimyi, N44°29.433', E146°11.783', 4.VIII.2013, leg. K. Makarov & Y. Sundukov” and “HOLOTYPE Bembidion (Nipponobembidion) ruruy Makarov & Sundukov 2014 [printed on red paper]
Copyright © Kirill Makarov

Bembidion (Zecillenus) chalmeri (Broun, 1886)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Sinechostictus (Pseudolimnaeum) alesmetana Toledano, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Omoperyphus) hypocrita Dejean, 1831
Male adult
Spain, Jaen prov., Arroyo de la piedra del Aguila, 37.683885, -3.485062, 3-4.5.2014, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Nesocidium) corticarium Sharp, 1903
Female adult
Halalea For. Res., Namolokama Mtn., 1295-1340 m el.. Image from Liebherr, 2008
Copyright © - - -

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Patrobus cinctus Motschulsky, 1860
Female adult
Jewish Aut. Reg., Obluchie Distr., 3-5 km upper Radde Vill. on Lagar Riv., forest 24.07.2004 I.Melnik leg.
L=9.5 mm
Copyright © Kirill Makarov

Deltomerus (Deltomerus) pseudoplatynus pseudoplatynus s.str.
Female adult
NW-Caucasus,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
long - 11mm.
Copyright © Ruslan Panin

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček

Platidiolus rufus Chaudoir, 1878
Male adult
Russia, Hamar-Daban Mt. rng., Komarinskiy Mt., upper flow of Slyudyanka riv. 15.7.2013, A. Shavrin leg. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Patrobus septentrionis volgensis Zamotajlov & Isaev, 2006
Male adult

Copyright © Igor Solodovnikov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) syriacus Chaudoir, 1871
Female adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 7 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Tachyura) ocellatus (Bates, 1892)
Female adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko

Tachyura (Amaurotachys) rhombophorus (Andrewes, 1925)
Male adult
India, Uttarakhand, 8 km NNE of Ramnagar, valley of Pipaliasot River, 420 m, 29°28,5' N, 79°10' E, 20.04.2012, R. Dudko, I. Lyubechanskii<br /> det. R. Dudko 2012
Copyright © Roman Dudko

Porotachys bisulcatus (Nicolai, 1822)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Eotachys (Eotachys) enormis Dostal, 2015 Holotype
Male adult
Madagascar, Andasibe National Park, Lokato, near Andasibe, 10.X.2007, leg. Zd. Mracek (in coll. Dostal, Vienna). Scale bar 1 mm
Copyright © Alexander Dostal

Pericompsus (Pericompsus) laetulus LeConte, 1851
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) terryli Liebherr, 2021 Paratype
Female adult
Kauai: Halelea F. R.: Namolokama Mtn., Waioli Str., litter, sift, ohia/ferns, 22°08.00'N, 159°29.85'W, 1340 m el., 21-v-2005, lot 1 Liebherr
Copyright © James K. Liebherr

Elaphropus (Elaphropus) globulus (Dejean, 1831)
Male adult
Spain, Granada, rio Darro, barranco de Teatino, 20.V.2005, Anichtchenko A. leg & det. (cAA). 1.8 mm
Copyright © Alexander Anichtchenko

Micratopus anaisae Lemaire, 2021 Holotype
Female adult
Saint-Félix, commune du Gosier, Guadeloupe, 16.210 N, 61.461 O, J. M. Lemaire leg. 15/04/2016. (MNHN). Image from Lemaire, 2021
Copyright © - - -

Tachyura (Tachyura) thoracicus (Kolenati, 1845)
Female adult
India, Uttarakhand, 14 km E Uttarkashi, 1500m, 10-12.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr as octostriata (Netolitzky, 1929)
Copyright © Alexander Anichtchenko

Tachyura (Tachylopha) ovatus (Motschulsky, 1851)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. leg & det. (cAA). 2.2 mm
Copyright © Alexander Anichtchenko

Meganophthalmus medvedevi Belousov & Koval, 2009 Paratype
Male adult
W Cauc., Abkhazia, Otshamtshira Distr., Otap Vill., Golova Otapa (= Uatapakhy) Cave, middle part, 22.VIII. 2009. A.G. Koval leg.<br /> Photo of Belousov et Koval.
Copyright © - - -

Shiqianaphaenops majusculus Ueno, 1999c: 47 Holotype
Male adult
Cave Feng Dong, Tangshan, Shiqian County, 27°29'10"N/108°15'23"E, 700 m in altitude, Mingyi Tian & Jingli Cheng leg.Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Minutotrechus minutus (Ueno, 1997) Paratype
Female adult
China, Yunnan, Gaoligong Shan, Tengchong County, Dabei, 2430 m (IOZ). Image from Deuve et al, 2016
Copyright © - - -

Oarotrechus gracilentus Townsend, 2010 Holotype
Male adult
Image from Lincoln University Entomology Research Collection
Copyright © - - -

Ckacus youcha Tian & Huang, 2021 Holotype
Male adult
Luoshui Dong, a vertical cave in Anlong Xian, southwestern most of Guizhou Province, Southwest China
Copyright © Mingyi Tian

Duvaliomimus (Duvaliomimus) watti Britton, 1958 Holotype
Male adult

Copyright © - - -

Birmaphaenops brehieri Deuve, 2017 Holotype
Male adult
Birmanie, État Shan, environs d’Ywangan, grotte Kyauk Khaung (“Stone Cave”), 21°11’28,2’’N - 96°33’9,4’’E (Franck Bréhier, 30.XI.2015, lot n°MY15-17/04), in coll. Muséum national d’Histoire naturelle, Paris.
Copyright © Thierry Deuve

Yunaphaenops graciliphallus Tian & He, 2023 Holotype
Male adult
Fengwei Dong Cave, Wenliu Xiang, Qiubei Xian, Wenshan Zhuang & Miao Autonomous Prefecture, eastern Yunnan, 24.33° N / 104.29° E, 1302 m, 3.IX.2020, leg. Chenghai Fu (SCAU).
Copyright © Mingyi Tian

Baehria separata Schmidt & Faille, 2023 Paratype
Male+Female adult
Ethiopia, Amhara, Mt. Choke, crater valley, alt. 3780–3900 m, 10°42'12"N, 37°50'58"E, 27.II.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W.
Copyright © Joachim Schmidt

Duvalius (Biharotrechus) durmitorensis bulbosus Janák, 2025 Holotype
Male adult
“MONTENEGRO, Bjelasica Mts: Δ Bendovac (1774 m) 1550-1600m (forest zone) P. Vonička lgt. 6.6.2003” [42°54’3.647”N, 19°37’0.657”E], “HOLOTYPUS Duvalius (Biharotrechus) durmitorensis bulbosus ssp. nov. J. Janák det. 2025”, (NMP). Image from Janak, 2025
Copyright © - - -

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -