Carabidae image gallery
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Anillini taxon page (59 photos)
Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov
Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov
Anillinus langdoni Sokolov & Carlton, 2004
Male adult
USA-TN, Sevier Co, GSMNP, secondary site, 490 m, ex leaf et log litter, 35°43.157' N, 83°23.309' W, 19.04.2011, J.-S. Park det. I. Sokolov, 2013
Copyright © Roman Dudko
Pelodiaetodes (Pelodiaetodes) constricticollis Sokolov, 2015 Paratype
Male adult
NZ, Northland, Radar Bush. Image from Sokolov, 2015
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Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini
Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
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Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
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Stylulus (Stylulus) isabelae Giachino & Sciaky, 2003 Holotype
Male adult
Copyright © Vittorino Monzini -
Bembidarenini taxon page (14 photos)
Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
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Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison
Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin -
Bembidiini taxon page (1016 photos)
Asaphidion substriatum Andrewes, 1925a: 56
Male adult
India, Uttarakhand, 2-3 km SW of Sonprayag, 2000-2200 m asl, 30°37' N, 78°59' E, 8-9.04.2012, R. Dudko, I. Lyubechanskii
R. Dudko det. 2012
Copyright © Roman Dudko
Sinechostictus (Sinechostictus) lederi (Reitter, 1888)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Ecuadion) chimborazonum (Bates, 1891)
Male adult
ECUADOR: Pichincha: Paso de la Virgen, DRM voucher V100793. Image from Maddison D.R., 2014.
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Bembidion (Cillenus) palosverdes Kavanaugh and Erwin, 1992
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Omotaphus) alsium Coquerel, 1866
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0
Bembidion (Nepha) genei trinacriae (Bonavita & Vigna-Taglianti, 2010)
Female adult
S Spain, Sierra de las Nieves, Los Quejigales 6-8.4.2010, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko
Bembidion (Testedium) laetum Brulle, 1836
Male adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 3.9 mm
Copyright © Kirill Makarov
Bembidion (Omotaphus) mellissii Wollaston, 1869
Female adult
RSA, 3km NW Nieuwoudtville, Willems riv. 27.11.2023 (cAA). det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko -
Horologionini taxon page (2 photos)
Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden -
Lissopogonini incertae sedis taxon page (7 photos)
Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov
Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko
Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko -
Patrobini taxon page (75 photos)
Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov
Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko
Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček
Dimorphopatrobus ludmilae Casale et Sciaky, 1994: 47
Female adult
Tibet, Motuo, Hanmi, 8.2013. det. A. Anichtchenko, 2015 17 mm
Copyright © Alexander Anichtchenko
Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk
Platidiolus rufus Chaudoir, 1878
Male adult
Russia, Hamar-Daban Mt. rng., Komarinskiy Mt., upper flow of Slyudyanka riv. 15.7.2013, A. Shavrin leg. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko
Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky
Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky
Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
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Pogonini taxon page (50 photos)
Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov
Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt
Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
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Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
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Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko
Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko
Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko
Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko -
Sinozolini taxon page (1 photos)
Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison -
Tachyini taxon page (170 photos)
Elaphropus (Tachyuropsis) exaratus (Bates, 1873)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Tachyura ascendens (Alluaud, 1917)
Female adult
TANZANIA, Kilimanjaro, Nkuansa, grassland,1660m, 3.7.2011, R. Felix det. as Elaphropus (Tachyura) exiguus (BOHEMAN 1848)
Copyright © Peter Schüle
Eotachys (Eotachys) enormis Dostal, 2015 Holotype
Male adult
Madagascar, Andasibe National Park, Lokato, near Andasibe, 10.X.2007, leg. Zd. Mracek (in coll. Dostal, Vienna). Scale bar 1 mm
Copyright © Alexander Dostal
Tachys (Paratachys) scitulus Leconte, 1848
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tachys (incertae) mulwalensis Sloane, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tachyura (Tachylopha) leleupi (Basilewsky In Basilewsky & Straneo, 1950)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tachysbembix sirena Erwin, 2004 Holotype
Male adult
COSTA RICA: Osa, P.N. Corcovado, Llorona, 20km N Estación Sirena, 1–100 m, 08º 37’ N 083º 42’ W, LS 270500,508300, January (R.W. Matthews & J.R. Matthews)(INBio: ADP103930).
Copyright © Terry Erwin
Liotachys antennatus Bates, 1871
Female adult
Bolivia: Santa Cruz Prov. Andrez Ibanez Potrerillo de Guenda, Terebinto. / 3-4 October 1994 / leg. P. Bettella / det. R. Davidson. Carnegie Museum of Natural History, Pittsburg, PA, USA
Copyright © Curt Harden
Tachyura (Tachylopha) ovatus (Motschulsky, 1851)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. leg & det. (cAA). 2.2 mm
Copyright © Alexander Anichtchenko -
Trechini taxon page (824 photos)
Trechus (Minitrechus) gypaeti Vigna Taglianti & Magrini, 2009 Holotype
Male adult
Etiopia - Arussi - loc. 33 bis / M. Encuolò, vetta / m 4000 circa - 5.XI.1973 / A. Vigna leg. (CVT). Image from Vigna Taglianti & Magrini, 2009.
Copyright © - - -
Pseudaphaenops tauricus Winkler, 1912a: 134
Male adult
Crimea, Dolgorukovskaya Yalla, yama Tshokrak gully, 13/5/2007, Turbanov leg.
Copyright © Vittorino Monzini
Trechus (Trechus) kricheldorffi Wagner, 1913
Female adult
Spain, Mt. Cantabricos, Picos de Europa, 2200m, 3km NW mirador Fuente De, 22.07.2019, A. Anichtchenko & R. Sciaky leg.; det. M. Toribio, 2019
Copyright © Alexander Anichtchenko
Uenoites reuteri Belousov, Kabak & Schmidt, 2019 Holotype
Male adult
MYANMAR: 1 male, with label data “N Burma, leg. Reuter, ca. 75 km NW Putao, 3700 m NN, 10.08.2006 ” (coll. J. Schmidt).
Copyright © Joachim Schmidt
Haixiaphaenops jinxiaohongae Tian, Cheng & Huang, 2021 Paratype
Male adult
Guizhou, Qingzhen, Anliu, Ximi, Changtu Dong cave (贵州省清镇市暗流镇长土洞), 26°51‘N, 106°21‘E, 1249 m, 2020–VI–27, leg. Chenggang Wang, Xiaohong Jing, Guangyuan Cheng, Yi Zhao & Mingyi Tian, SCAU.
Copyright © Mingyi Tian
Pangaphaenops seraphicus Tian, Huang & Jia, 2023 Holotype
Male adult
Cave I, Suolue Town, Bama Yao Autonomous County, Hechi Shi, Guangxi Zhuang Autonomous Region, Southern China, 113.36°E, 23.16°N, 532 m, 2022-VI-21, leg. Jingli Cheng, in SCAU.
Copyright © Mingyi Tian
Aspidaphaenops qujingensis achaetus Deuve & Tian, 2020 Holotype
Male adult
Chine, Yunnan, Préfecture de Qujing, Shizong, grotte Xinzhuangke Dong, 1110 mètres, 24°39’4’’N 104°15’57’’E (Tian M. Y., Lui W. X., Wang X. H., Tang M. R. leg., 17.VI.2015), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve
Perileptus (Parablemus) cf cameroni Jeannel, 1923
Male adult
India, Uttarakhand, 8km NW Jaspur, lake shore, 22-23.4.2012, Anichtchenko A. leg. & det. (cAA). 2.3 mm
Copyright © Alexander Anichtchenko -
Zolini taxon page (32 photos)
Zolus helmsi (Sharp, 1886)
Female adult
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Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
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Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
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Maungazolus parvulus (Broun, 1903)
Male adult
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Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
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Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
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Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult
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