Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro

Illaphanus pteridophyticus Giachino, 2005 Holotype
Male adult

Copyright © Alexander Anichtchenko

Typhlocharis laurentii Magrini, 2000 Paratype
Unsexed
España, Almeria, Sierra Almagrera ; carretera N332, 30.XI.98, leg. Fancello (MNHN, Antoine Mantilleri, 2016)
Copyright © - - -

Scotodipnus alpinus Baudi di Selve, 1871
Female adult
Piemonte, Valle Po, Crissolo (CN), 27.V.2000, 1250m, fagus, leg. F. Angelini
Copyright © Alexander Anichtchenko

Scotodipnus (Binaghites) affinis affinis (Baudi di Selva, 1871)
Male adult
Italy, Emilia, M. Penna (PR), 1400m, 24.5.1999, leg. F. Angelini
Copyright © Alexander Anichtchenko

Scotodipnus (Binaghites) armellinii (Ganglbauer, 1900)
Male adult
Veneto Italia
Copyright © Erminio Piva

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Anillus florentinus florentinus Dieck, 1869
Unsexed
Emilia Romagna, Sasseleone (BO), 26.3.2002 350m, A.Degiovanni leg. (cAA)
Copyright © Alexander Anichtchenko

Stylulus (Stylulus) isabelae Giachino & Sciaky, 2003 Holotype
Male adult

Copyright © Vittorino Monzini

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Erwiniana punctisculptilis (Erwin, 1994)
Male adult
Ecuador, Yasuni
Photo of Sokolov I. M.
Copyright © Igor Sokolov

Bembidion (Notaphus) semipunctatum (Danovan, 1806)
Female adult

Copyright © Kirill Makarov

Bembidion (Notholopha) scitulum (Erichson, 1834)
Male adult
CHILE: Reg. VII: Los Niches E of Curicó, DRM voucher V100790; Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Peryphus) bracculatum Bates, 1889b: 212
Female adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A.; 5.5 mm, det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Lionepha) disjunctum Lindroth, 1963 Holotype
Male adult

Copyright © - - -

Bembidion (Bembidionetolitzkya) bactrianum K. Daniel, 1902
Male adult
Tadzhikistan, Varzob, Khodja obi Garm, 2000m 21-24.3.2015 (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Euperyphus) scapulare lomnickii Netolitzky, 1916: 260
Female adult
W Ukraine, NE Carpathians, Lviv reg., near Rozgirche vill., bank of Stryi river, 11.V.2014, Khrapov D. leg./det.Panin R., L= 5.4 mm
Copyright © Denis Khrapov

Bembidion (Liocosmius) cachagua Maddison, 2023 Holotype
Male adult
CAS: “U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln.”, “C.A.S. Accession 29-V-48 Cachagua Creek”, “David R. Maddison DNA3842 DNA Voucher”
Copyright © David Maddison

Orzolina kratkyi Machado, 2025 Holotype
Male adult
SPAIN, Canary Islands, Tenerife, La Guancha, Charco del Viento, -1 m. (28º24'03"N 16º40'26.5"W) 3-11-2024 leg. A. Machado. Coll. TFMC-EN 4592. Image from Machado, 2025
Copyright © - - -

Bembidion (Bembidionetolitzkya) peliopterum Chaudoir, 1850b: 182
Male adult
Uzbekistan, Chatkal Mt. rng., Yangiobod, left conf of Dukentsay riv, 41.07.28N 70.06.21E 1300m 2.8.2021 Anichtchenko A. leg. (cAA). det. L. Toledano, 2026; 5.8 mm
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Deltomerodes ovicollis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Deltomerodes conaensis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Diplous (Diplous) sibiricus atratus Habu, 1951
Male adult

Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) syriacus Chaudoir, 1871
Female adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 7 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Elaphropus (Tachylopha) spenceri (Sloane, 1896) Paratype
Male adult

Copyright © Museums Victoria

Tachyura (Tachyura) nitens (Andrewes, 1925)
Female adult
Indonesia E, W New Guinea, Doberai Peninsula, Ayamaru vill. 27 km SE Uter (Aitinyo) lakeside, 1*23'01S 132*23'09E, 175-177m, 4.IX.2015, muddy bank, between lakeside vegetation, leg. D. Telnov. 2.8 mm. det. M. Baehr, 2016
Copyright © Alexander Anichtchenko

Tachyura (Sphaerotachys) curticollis (Sloane, 1896)
Male adult
Indonesia E, W New Guinea, Doberai Peninsula, Ayamaru vill. 27 km SE Uter (Aitinyo) lakeside, 1*23'01S 132*23'09E, 175-177m, 4.IX.2015, muddy bank, between lakeside vegetation, leg. D. Telnov. 2.2 mm. det. M. Baehr, 2016
Copyright © Alexander Anichtchenko

Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Tachys) corax Leconte, 1852
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Eotachys (Jeannelys) euryodes (Bates, 1892)
Unsexed
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort leg. U.Schmidt, 14.XI.2007; det. M.Baehr, 2008; Photo: U.Schmidt, 2021
Copyright © Udo Schmidt

Tachyta (Eurytachyta) laticollis Baehr, 2016 Holotype
Female adult
Borneo, Sabah, Kinabalu NP, HQ vic., 1550 m, 22.–25.V.2005, R. GRIMM (CBM).
Copyright © Martin Baehr

Costitachys inusitatus Erwin, 1974 Holotype
Male adult
Santarem, Para, Brazil, IV.27.1963 FG Werner (MCZ)
Copyright © - - -

Liotachys antennatus Bates, 1871
Female adult
Bolivia: Santa Cruz Prov. Andrez Ibanez Potrerillo de Guenda, Terebinto. / 3-4 October 1994 / leg. P. Bettella / det. R. Davidson. Carnegie Museum of Natural History, Pittsburg, PA, USA
Copyright © Curt Harden

Queinnectrechus micrangulus Belousov & Kabak, 2003 Paratype Paratype
Male adult
China, Sichuan, SSW of Shimian, E slope of Mt "4977" W of Lijipin 3000-3900m
5.07.2000 Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Dianotrechus gueorguievi Tian, 2016 Holotype
Female adult
China, Yunnan Province, Ermu Vill., Kunming District, Dashi Dong (Big Rock Cave), 24°49'13"N/102°27'56"E, 1940 m in altitude, XI-8- 2011, B. Petrov leg.” (SCAU). Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Paratrechodes macleayi (Sloane, 1920)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Pseudanophthalmus engelhardti (Barber, 1928)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Cimmeritodes (Zhecimmerites) zhejiangensis Deuve & Tian, 2015 Holotype
Male adult
Chine, Zhejiang occidental, Quzhou City, Changshan Xian, Xinjiao Xiang, Dongling Cun, grotte Niujiao Dong, 546 mètres, 29°09’44,86’’N - 118°40’26,18’’E, Sunbin Huang & Xinhui Wang leg., 4.X.2014, in coll. South China Agricultural University, à Canton (Chine). Image from Deuve & Tian, 2015
Copyright © Mingyi Tian

Neotrechus hilfi (Reitter, 1903)
Male adult

Copyright © Vittorino Monzini

Boldoriella (Insubrites) chiarae Monguzzi, 1982b: 211
Male adult
Mt. Guglielmo Brescia Italia
Copyright © Vittorino Monzini

Deuveopsis (Deuveopsis) lobeliae Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Amhara, Mount Choke, station I (afro-alpine belt, rocks below Lobelia), 10˚41’54’’N, 037˚50’21’’E, 3950 m a.s.l., 21.VIII.2011, E. Quéinnec & C. Reeb leg.
Copyright © Eric Quéinnec

Aphaenops (Riberaphaenops) holcartensis Genest, 1977: 281
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -

Alanorites enigmaticus Belousov, 1998: 139
Female adult
Russia, Karachay-Cherkessia, Zelenchuksky Distr., Arkhyz, “Romantik”, 43°34'28.45"N, 41°12'18.27"E, h ~ 2730 m, 02.07.2026, Ya.N. Kovalenko leg L=2.6 mm
Copyright © Kirill Makarov

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
Copyright © - - -

Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
Copyright © - - -

Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -