Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Anillinus affabilis (Brues, 1902) Lectotype
Male adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm.
Copyright © Igor Sokolov

Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Rhegmatobius quadricollis (Ehlers, 1883)
Unsexed
Majella, Abruzzo, Italy
Copyright © Vittorino Monzini

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Medusapyga hehalis LaBonte, 2023 Paratype
Female adult
WASHINGTON, Thurston Co., 11.3 km NW Littlerock, elev. ~400 m, D.J. Cox and J.R. LaBonte, 6 May 2007, 46.9600°N 123.13450°W. Image from LaBonte & Maddison, 2023
Copyright © - - -

Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
Copyright © - - -

Stylulus sp.
Unsexed
"US Virgin Islands, St. Croix, 12.vi.1972, W.B. Muchmore Leg." Florida State Collection of Arthropods. "Stylulus sp." det. C.W. Harden 2023.
Copyright © Curt Harden

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Talanes) subfasciatum Chaudoir, 1850b: 187
Female adult
Krasnodar Reg., Taman Peninsula, Tuzlo Lake 6.V.2009
2.7 mm
Copyright © Kirill Makarov

Bembidion (Philochthus) cf. neresheimeri J. Muller, 1929
Female adult
Ивановская обл., г. Иваново, парк, 1905 г., 29.05.2014, М. Смирнов leg.; det. K.Makarov. Image from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Xystosomus laevis Erwin, 1973 Holotype
Male adult
Brasilien, Nova Teutonia
Copyright © - - -

Bembidion (Peryphus) bracculatum Bates, 1889b: 212
Female adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A.; 5.5 mm, det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) mckinleyi scandicum Lindroth, 1943
Female adult
E Siberia, Lake Baikal, Malyi Kochergat vill., Goloustnaya riv., 16.VII.1991, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Sloanephila) jacksoniense Guerin Meneville, 1830
Female adult
Wiluna W.A. / ix 25 1931 //Australia / Harvard Exp. / Darlington (MCZ). Scale bars 0.5 mm.
Copyright © David Maddison

Bembidion (Chlorodium) difforme (Motschulsky, 1844)
Female adult
Russia, Burjatia, Muhor-Shibirsk distr., 5km N Nikolsk vill., salt marsh, 27.8.1998, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Bembidion (Peryphanes) milleri carpathicum J. Muller, 1918b: 88
Male adult
W Ukraine, W Podolian Upland, Rogatyn Opillia, Zaluzhzhya vill, 49°24'33"N 24°35'36"E, clay quarry, 30-31.VIII.2018, Panin R leg., L=4.6mm
Copyright © Denis Khrapov

Odontium (Bracteon) conicolle Motschulsky, 1844: 273
Female adult
Burjatia, Barguzin depr., Alla riv., 8.8.1997, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Bembidion (Nepha) genei trinacriae (Bonavita & Vigna-Taglianti, 2010)
Female adult
S Spain, Sierra de las Nieves, Los Quejigales 6-8.4.2010, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Deltomerus (Deltomerus) tatricus L. Miller, 1859: 306
Male adult
N Slovakia; Vysoké Tatry Mts., 10mm
Copyright © Aleš Sedláček

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Platidiolus cf. borealis Zamotajlov, 2001
Female adult
Russia, Khabarovsk reg., Verhnebureinski distr. Bureya river valley, Ussomah river, 22.7.2006 (anal sternite with 3 pairs of setae). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Deltomerodes ovicollis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Deltomerodes conaensis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Elaphropus (Barytachys) tripunctatus (Say, 1830)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Micratopus sp.
Male adult
Panama, Darien Nat. Park, Rancho Frio, 10.I.2017, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Setitachys macrops Baehr, 2016 Paratype
Female adult
Qld. Bentinck Is. Rokoti, 16 June 1963. P. Aitken, N. B. Tindale
Copyright © Martin Baehr

Tachys (Polyderis) laeva (Say, 1823)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Tachys) corax Leconte, 1852
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachysbembix sirena Erwin, 2004 Holotype
Male adult
COSTA RICA: Osa, P.N. Corcovado, Llorona, 20km N Estación Sirena, 1–100 m, 08º 37’ N 083º 42’ W, LS 270500,508300, January (R.W. Matthews & J.R. Matthews)(INBio: ADP103930).
Copyright © Terry Erwin

Tachys (Paratachys) terryli Liebherr, 2021 Paratype
Female adult
Kauai: Halelea F. R.: Namolokama Mtn., Waioli Str., litter, sift, ohia/ferns, 22°08.00'N, 159°29.85'W, 1340 m el., 21-v-2005, lot 1 Liebherr
Copyright © James K. Liebherr

Eotachys (Jeannelys) euryodes (Bates, 1892)
Unsexed
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort leg. U.Schmidt, 14.XI.2007; det. M.Baehr, 2008; Photo: U.Schmidt, 2021
Copyright © Udo Schmidt

Costitachys inusitatus Erwin, 1974 Holotype
Male adult
Santarem, Para, Brazil, IV.27.1963 FG Werner (MCZ)
Copyright © - - -

Eotrechodes larisae Ueno, Lafer et Sundukov, 1995: 114
Male adult

Copyright © Kirill Makarov

Huoyanodytes tujiaphilus Tian & Huang, 2016 Holotype
Female adult
Cave Tujiamei Dong, Huoyan Karst, Huoyan Xiang, Wulongshan Geopark, Longshan County, NW Hunan Province, China, 29°12'20.11"N/109°18'37.21"E, 427 m in altitude, VII-3-2014, leg. Mingyi Tian, Weixin Liu, Haomin Yin, Sunbin Huang & Xinhui Wang, deposited in SCAU. Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Pseudanophthalmus engelhardti (Barber, 1928)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Yalongaphaenops erwini Belousov & Kabak, 2021 Holotype
Male adult
China, Sichuan Province, SW Mianning Town; 28°15'27"N, 101°43'51"E; H = 3805 m a.s.l.; 11.07.2011; Belousov & Kabak leg. (ZIN).
Copyright © Ilya Kabak

Orotrechus jamae C. Etonti et M. Etonti, 1979: 27
Male adult
NE Italy, Belluno, Prealps, Cansiglio plateau, Bus dell'Ors, 19.V.2013, L=6.2mm
Copyright © Denis Khrapov

Velebitaphaenops giganteus Casale & Jalzic, 2012
Female adult
Image from cavebiology.net
Copyright © Creative Commons

Deuveopsis (Deuveopsis) lobeliae Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Amhara, Mount Choke, station I (afro-alpine belt, rocks below Lobelia), 10˚41’54’’N, 037˚50’21’’E, 3950 m a.s.l., 21.VIII.2011, E. Quéinnec & C. Reeb leg.
Copyright © Eric Quéinnec

Troglocimmerites zapparolii Casale & Magrini, 2023 Holotype
Female adult
Turchia, Anatolia nord-orientale, vilayet Giresun, strada per Çanakci, 70-80 m s.l.m., 5,8 km dal bivio per Çanakci, 6.V.1987, leg. M. Zapparoli (AVT). Image from Casale & Magrini, 2023
Copyright © - - -

Neoblemus championi Jeannel, 1923
Male adult
India, Uttarakhand, 18-20.4.2012, 15km SW New Tehri, 869m, Anichtchenko A. leg. & det. (cAA).
Copyright © Alexander Anichtchenko

Tianaphaenops peijiui Deuve & Li, 2026 Holotype
Male adult
Chine, Hunan, partie orientale de la Préfecture de Huaihua (collecteur local, juin-août 2025), in coll. Li Aihuan, à Chengdu, Sichuan, Chine.
Copyright © Thierry Deuve

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
Copyright © - - -

Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -