Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Anillinus langdoni Sokolov & Carlton, 2004
Male adult
USA-TN, Sevier Co, GSMNP, secondary site, 490 m, ex leaf et log litter, 35°43.157' N, 83°23.309' W, 19.04.2011, J.-S. Park det. I. Sokolov, 2013
Copyright © Roman Dudko

Anillinus affabilis (Brues, 1902) Lectotype
Male adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm.
Copyright © Igor Sokolov

Illaphanus pteridophyticus Giachino, 2005 Holotype
Male adult

Copyright © Alexander Anichtchenko

Winklerites stevanovici Hlavac & Margini, 2017 Paratype
Male adult
Srbija [p], Dimitrovgrad, Petrlaš, Džemanska prop[ast], 02.05.2002 [h], M. Stevanović leg. Image from HLAVÁČ & MAGRINI, 2017
Copyright © - - -

Scotodipnus (Binaghites) armellinii (Ganglbauer, 1900)
Male adult
Veneto Italia
Copyright © Erminio Piva

Rhegmatobius quadricollis (Ehlers, 1883)
Unsexed
Majella, Abruzzo, Italy
Copyright © Vittorino Monzini

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Nesamblyops townsendi Sokolov, 2023
Male adult
Tennyson Inlet, Marlborough Sounds, SO
Copyright © Igor Sokolov

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Talanes) subfasciatum Chaudoir, 1850b: 187
Female adult
Krasnodar Reg., Taman Peninsula, Tuzlo Lake 6.V.2009
2.7 mm
Copyright © Kirill Makarov

Amerizus (Amerizus) teles Belousov & Dudko, 2010 Paratype
Male adult
NE Altay Mts., 35-40 km SSE Artybash, Evrechala Mt., 1850-2100 м, tundra, 51°28' N, 87°26' E, 20-22.05.2003, R.Yu.Dudko & I.I.Lubechanskii leg.
Copyright © Igor Belousov

Bembidion (Microserrullula) splendens Andrewes, 1923
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 23-26.04.2012 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Cillenus (Cillenus) lateralis Samouelle, 1819
Male adult
Netherlands, Bruinisse, leg. Krift, 1976; det. U.Schmidt, 1982
Photo: U.Schmidt, 2008
Copyright © Udo Schmidt

Bembidion (Blepharoplataphus) hastii C.R. Sahlberg, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Antiperyphanes) hirtipes (Jeannel, 1962)
Male adult
Argentina, Mendoza, Pampa Palauco. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Zecillenus) albescens (Bates 1878)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Sloanephila) tahitiense Liebherr & Maddison, 2013 Holotype
Male adult
(MNHN): French Polynesia: Tahiti Nui / Mt. Mauru lava tube / el. 705 m 18-IX-2006 lot 01 / 17˚38.017´S 149˚21.284´W / pyr. fog moss/liverworts / along stream J.K. Liebherr // HOLOTYPE / Bembidion / tahitiense / Liebherr & / Maddison 2012 (black-bordered red label).
Copyright © David Maddison

Bembidion (Pseudophilochthus) evanescens (Wollaston, 1877)
Female adult

Copyright © David Maddison

Bembidion (Ocyturanes) marginipenne Solsky, 1874: 130
Male adult
Uzbekistan, Chatkal Mt. rng., Yangiobod, left conf of Dukentsay riv, 41.07.28N 70.06.21E 1300m 2.8.2021 Anichtchenko A. leg. (cAA). det. L. Toledano, 2026; 3.8 mm
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Penetretus nebrioides (Vuillefroy, 1866)
Male adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Patrobus septentrionis volgensis Zamotajlov & Isaev, 2006
Male adult

Copyright © Igor Solodovnikov

Deltomerodes conaensis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Deltomerus (Deltomerus) corax Peyerimhoff, 1922
Male adult
Morocco, Grand Atlas, 2200-2300m, N31.13.09.5 W07.49.29.7 16-17.6.2010 Anichtchenko A. leg. & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Elaphropus) latissimus (Motschulsky, 1851)
Female adult
Kunashiri Isl., Calder of Golovnina Volcano, S coast od Goryachee Lake 43°52'19"N 145°29'43"E 12.VIII.2008 leg.K.Makarov, A.Zaitzev
2.0 mm
Copyright © Kirill Makarov

Elaphropus (Tachyura) bombycinus (Andrewes, 1925)
Male adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko

Tachyura (Amaurotachys) rhombophorus (Andrewes, 1925)
Male adult
India, Uttarakhand, 8 km NNE of Ramnagar, valley of Pipaliasot River, 420 m, 29°28,5' N, 79°10' E, 20.04.2012, R. Dudko, I. Lyubechanskii<br /> det. R. Dudko 2012
Copyright © Roman Dudko

Elaphropus (Tachyuropsis) exaratus (Bates, 1873)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pericompsus (Upocompsus) habitans (Sloane, 1896) Syntypes
Male adult

Copyright © Museums Victoria

Tachyura (Sphaerotachys) curticollis (Sloane, 1896)
Male adult
Indonesia E, W New Guinea, Doberai Peninsula, Ayamaru vill. 27 km SE Uter (Aitinyo) lakeside, 1*23'01S 132*23'09E, 175-177m, 4.IX.2015, muddy bank, between lakeside vegetation, leg. D. Telnov. 2.2 mm. det. M. Baehr, 2016
Copyright © Alexander Anichtchenko

Tachyura (Tachylopha) leleupi (Basilewsky In Basilewsky & Straneo, 1950)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) terryli Liebherr, 2021 Paratype
Female adult
Kauai: Halelea F. R.: Namolokama Mtn., Waioli Str., litter, sift, ohia/ferns, 22°08.00'N, 159°29.85'W, 1340 m el., 21-v-2005, lot 1 Liebherr
Copyright © James K. Liebherr

Tachyta (Paratachyta) coracina (Putzeys, 1875)
Male adult
Malaysia, Langkawi, environment of Hotel Berjaya, leg. U.Schmidt, 7.-22.XI.2009; det. M.Baehr, 2010; Photo: U.Schmidt, 2021; 2.1mm
Copyright © Udo Schmidt

Tachyura (Tachyura) thoracicus (Kolenati, 1845)
Female adult
India, Uttarakhand, 14 km E Uttarkashi, 1500m, 10-12.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr as octostriata (Netolitzky, 1929)
Copyright © Alexander Anichtchenko

Meganophthalmus medvedevi Belousov & Koval, 2009 Paratype
Male adult
W Cauc., Abkhazia, Otshamtshira Distr., Otap Vill., Golova Otapa (= Uatapakhy) Cave, middle part, 22.VIII. 2009. A.G. Koval leg.<br /> Photo of Belousov et Koval.
Copyright © - - -

Trechus (Epaphius) rivularis (Gyllenhal, 1810)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Queinnectrechus (Gaoligongtrechus) balli Deuve and Kavanaugh, 2016 Holotype
Male adult
CHINA, Yunnan Province, Gaoligong Shan, Nujiang Prefecture, Nujiang State Nature Reserve, Dulong/Gongshan Yakou [= Qiqi/Dulong divide] area, 21 airkm w of Gongshan,”/ “N27.69655°/ E98.45389°, 3300-3680m, 16-17 July 2000, Stop#00-24C, D.H. Kavanaugh, C.E. Griswold, Liang H.-B., D. Ubick, & Dong D.-Z. collectors (IOZ). Image from Deuve et al, 2016
Copyright © - - -

Rakantrechus (Paratrechiama) tofaceus Ueno, 1970b: 106
Male adult
Japan, Kumamoto Ken
Copyright © Vittorino Monzini

Boldoriella (Insubrites) grignensis Monzini, 1987: 13 Paratype
Male adult
Italy, Grigna Mt., leg V. Monzini
Copyright © Vittorino Monzini

Pheggomisetes globiceps Buresch, 1925
Male adult
Bulgaria, Ponor plannina, Dusnica Pestera cave, 30/7/2004, Vasiljev leg.
Copyright © Vittorino Monzini

Sinotroglodytes hefengensis Tian & Li, 2021 Holotype
Male adult
the limestone cave Qizimei Dong, Qizimeishan, Chunmuyun, Hefeng County, Enshi Tujia & Miao Autonomous Prefecture, southwestern Hubei Province, Central China, 30°02’30”N /109°47’09”E, 1820 m in altitude, 2020-VI-18, leg. Chenliang Li (SCAU).
Copyright © Mingyi Tian

Boldoriella (Insubrites) pesarinii Sciaky, 1982: 25
Male adult
Mt. Pora, 1800 m., Bratto BG Italia
Copyright © Vittorino Monzini

Allotrechiama (Allotrechiama) iriei Ueno, 1970b: 95
Male adult
Japan
Copyright © Augusto Degiovanni

Duvalius (Biharotrechus) durmitorensis bulbosus Janák, 2025 Holotype
Male adult
“MONTENEGRO, Bjelasica Mts: Δ Bendovac (1774 m) 1550-1600m (forest zone) P. Vonička lgt. 6.6.2003” [42°54’3.647”N, 19°37’0.657”E], “HOLOTYPUS Duvalius (Biharotrechus) durmitorensis bulbosus ssp. nov. J. Janák det. 2025”, (NMP). Image from Janak, 2025
Copyright © - - -

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -