Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Hypotyphlus degiovannii Magrini, 2013 Holotype
Male adult
Italia, Lazio, Montagna di Scauri (Latina), 50 m s.l.m., 1.V.2008, P. Magrini leg. (CM).
Copyright © Paolo Magrini

Illaphanus pteridophyticus Giachino, 2005 Holotype
Male adult

Copyright © Alexander Anichtchenko

Austranillus macleayi (Lea, 1906) Syntypes
Female adult

Copyright © Museums Victoria

Winklerites stevanovici Hlavac & Margini, 2017 Paratype
Male adult
Srbija [p], Dimitrovgrad, Petrlaš, Džemanska prop[ast], 02.05.2002 [h], M. Stevanović leg. Image from HLAVÁČ & MAGRINI, 2017
Copyright © - - -

Pelodiaetus nunni Sokolov, 2019 Paratype
Male adult
NZ, Canterbury, Ahuriri Scenic Reserve. 1.27 mm
Copyright © Igor Sokolov

Nesamblyops townsendi Sokolov, 2023
Male adult
Tennyson Inlet, Marlborough Sounds, SO
Copyright © Igor Sokolov

Anillus florentinus andreinii Jeannel, 1937
Unsexed
Italy, Toscana, Giuncarico 8.12.96 leg Degiovanni A. (cAA)
Copyright © Alexander Anichtchenko

Stylulus sp.
Unsexed
"US Virgin Islands, St. Croix, 12.vi.1972, W.B. Muchmore Leg." Florida State Collection of Arthropods. "Stylulus sp." det. C.W. Harden 2023.
Copyright © Curt Harden

Stylulus (Stylulus) isabelae Giachino & Sciaky, 2003 Holotype
Male adult

Copyright © Vittorino Monzini

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Plataphus) prasinum (Duftschmid, 1812)
Female adult

Copyright © Kirill Makarov

Asaphidion substriatum Andrewes, 1925a: 56
Male adult
India, Uttarakhand, 2-3 km SW of Sonprayag, 2000-2200 m asl, 30°37' N, 78°59' E, 8-9.04.2012, R. Dudko, I. Lyubechanskii
R. Dudko det. 2012
Copyright © Roman Dudko

Sinechostictus (Sinechostictus) frederici (G. Muller, 1918)
Female adult
Spain, Jaen prov., Arroyo de la Piedra del Aguila; 3-4.05.2014, 37.683885, -3.485062, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Antiperyphus) philippii Germain, 1906
Male adult
Argentina: Neuquén: Rio Collón Curá ca 13 km S La Rinconada, DRM voucher V100787. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Peryphanes) dalmatinum dalmatinum Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion () eutherum Andrewes, 1923b: 96
Male adult
India, Uttarakhand, 15 km SW New Tehri, 869m, 18-20.4.2012, Anichtchenko A. leg.; det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) reibnitzi Schmidt, 2018 Paratype
Male adult
“S TIBET Kyi Chu river / betw. Lhasa and airport / ca. 3650 m, 22.IX.2010 / leg. J. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Asioperyphus) notatum Andrewes, 1922b: 175
Female adult
India, Uttarakhand, 8 km NW Jaspur, lake shore, 22-23.4.2012, Anichtchenko A. leg. (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Euperyphus) giganteum giganteum J.R. Sahlberg, 1900
Female adult
S Kazakhstan, N slope Talas Mt.rng., Dzhabagly riv., 1100m 28.7.2004, A.Matalin leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Trepanedoris) kieranae Maddison & Sproul, 2026 Paratype
Male adult
“USA: Oregon: Lane Co., SW Eugene, 288m 44.0031°N 123.1299°W, 5.vi.2015. DRM 15.069. Maddison, Kieran, Mendez, Zeleznik”, “David R. Maddison DNA4576 DNA Voucher” (OSAC)
Copyright © David Maddison

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Deltomerus (Deltomerus) pseudoplatynus pseudoplatynus s.str.
Female adult
NW-Caucasus,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
long - 11mm.
Copyright © Ruslan Panin

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Deltomerus (Deltomerus) tatricus L. Miller, 1859: 306
Male adult
N Slovakia; Vysoké Tatry Mts., 10mm
Copyright © Aleš Sedláček

Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček

Penetretus temporalis Bedel, 1909
Female adult
Spain, Cadiz prov., Algeciras vic., h=50m, 36.080429, -5.428613, 11-12.05.2014, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
Copyright © - - -

Deltomerodes conaensis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Tachyura) ocellatus (Bates, 1892)
Female adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko

Meotachys (Hylotachys) ballorum Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Tapurucuara
Copyright © Terry Erwin

Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) terryli Liebherr, 2021 Paratype
Female adult
Kauai: Halelea F. R.: Namolokama Mtn., Waioli Str., litter, sift, ohia/ferns, 22°08.00'N, 159°29.85'W, 1340 m el., 21-v-2005, lot 1 Liebherr
Copyright © James K. Liebherr

Eotachys (Jeannelys) euryodes (Bates, 1892)
Unsexed
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort leg. U.Schmidt, 14.XI.2007; det. M.Baehr, 2008; Photo: U.Schmidt, 2021
Copyright © Udo Schmidt

Tachyta (Paratachyta) coracina (Putzeys, 1875)
Male adult
Malaysia, Langkawi, environment of Hotel Berjaya, leg. U.Schmidt, 7.-22.XI.2009; det. M.Baehr, 2010; Photo: U.Schmidt, 2021; 2.1mm
Copyright © Udo Schmidt

Tachyta (Eurytachyta) laticollis Baehr, 2016 Holotype
Female adult
Borneo, Sabah, Kinabalu NP, HQ vic., 1550 m, 22.–25.V.2005, R. GRIMM (CBM).
Copyright © Martin Baehr

Tachyura (Tachyura) yuezhi Kabak, 2021 Holotype
Male adult
China, Xinjiang, Ruoqiang River, Ruoqiang (= Charklik) Town, 39°00ʹ59ʺN / 88°09ʹ24ʺE, H = 820 m, 19.07.2015, Kabak I.I. leg.
Copyright © Ilya Kabak

Tachyura (Tachylopha) ovatus (Motschulsky, 1851)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. leg & det. (cAA). 2.2 mm
Copyright © Alexander Anichtchenko

Trechus (Trechus) schyberosiae Szallies & Schüle, 2011 Holotype
Male adult
Kanton Luzern, Schrattenfluh, unterhalb Hengst, 2100 m, 14.6.2009, leg. A. Szallies (NMBE).
Drawing Peter Schuele
Copyright © Peter Schüle

Epaphiopsis (Epaphiama) jacobsoni Sokolov et Shilenkov, 1987: 10
Male adult
Altai Republic: Holzun mt.rng., Bannaya riv., 1300-1600 m,12-14.06.1999, R.Yu & A.Yu. Dudko leg. & det.
Copyright © Roman Dudko

Cimmeritodes (Xiangcimmerites) zhongfangensis Deuve & Tian, 2016 Holotype
Female adult
Chine, Hunan, Huaihua Shi, Zhongfang Xian, Huangyan Xiang, Fujia Cun, grotte Jinji Dong, 890 m, 27°28’20’’N - 110°4’7’’E (Tian Mingyi, Liu Weixin, Wang Xinhui, Tang Mingruo, 1.VII.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -

Cathaiaphaenops enshiensis Deuve & Tian, 2016 Holotype
Male adult
Chine, Hubei, Enshi, Banqiao, grotte Huangjingping Dadong, 1710 m, 30°30’42’’N - 109°18’38’’E (Tian Mingyi, Huang Sunbin, Wang Xinhui, Tang Mingruo, 8.XI.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -

Cnides cf jeanneli Ueno, 1985
Female adult
Creative Commons CC-BY 3.0. det. Anichtchenko A. 2018
Copyright © David Maddison

Qianaphaenops (Qiandongaphaenops) variabilis Tian, Huang & Wang, 2015 Holotype Paratype
Male adult
cave Wanfuchangcheng Dong, in Jiangjun Shan, Pingzhuang Xiang, Cengong County, Guizhou Province, 27°24’50.91”N, 108°35’43.0”E, 830 m, 2014-VI-29, leg. Mingyi Tian, Weixin Liu, Haomin Yin, Sunbin Huang and Xinhui Wang (SCAU). Image from Tian, Huang & Wang, 2015
Copyright © Mingyi Tian

Zhijinaphaenops liuae Deuve & Tian, 2015 Holotype
Male adult
Chine, Guizhou, Guiyang, Xifeng Xian, Shidong Zhen, grotte Hejia Dong, 1198 mètres, 27°2’30,89’’N - 106°31’40,38’’E, Weixin Liu, 13.VII.2013, in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2015
Copyright © Mingyi Tian

Velebitaphaenops giganteus Casale & Jalzic, 2012
Female adult
Image from cavebiology.net
Copyright © Creative Commons

Chiapadytes bolivari Vigna Taglianti, 1977
Female adult
Mexico, Chiapas, San Cristobal de Las Casas, Las Piedrecitas, Cueva de la Planta I, 25.III.1982, leg. A. Gobetti e J.F. Pittet (TT ♀, MSNG). Image from Casale & Magrini, 2021
Copyright © - - -

Trechiama (Leptepaphiama) sichotanus Lafer, 1989: 143
Male adult
South Primorye, Przhevalskogo Mt.R., 7 km NNW of Benevskoye vill., middle flow of Elamovsky Spring, h~380, 43°14'07"N 133°44'01"E 23.V.2026, leg. K.Makarov L=6.1 mm
Copyright © Kirill Makarov

Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -