Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Austranillus macleayi (Lea, 1906) Syntypes
Female adult

Copyright © Museums Victoria

Typhlocharis laurentii Magrini, 2000 Paratype
Unsexed
España, Almeria, Sierra Almagrera ; carretera N332, 30.XI.98, leg. Fancello (MNHN, Antoine Mantilleri, 2016)
Copyright © - - -

Scotodipnus (Binaghites) armellinii (Ganglbauer, 1900)
Male adult
Veneto Italia
Copyright © Erminio Piva

Pelodiaetus sulcatipennis Jeannel, 1937
Male adult
New Zealand, Otago, Outram
Copyright © Igor Sokolov

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
Copyright © - - -

Megastylulus pivai Giachino & Sciaky, 2003 Holotype
Male adult
leg. E. Piva - Isl. St.Lucia Caraibi
Copyright © Vittorino Monzini

Stylulus (Stylulus) isabelae Giachino & Sciaky, 2003 Holotype
Male adult

Copyright © Vittorino Monzini

Rhegmatobius strictus (Baudi di Selva, 1891)
Male adult
Sardegna, Gennargentu Mte., Spada Mt., 1000, 27.4.2003 A. Degiovanni leg. (cAA).
Copyright © Alexander Anichtchenko

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Hirmoplataphus) nigrum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Chlorodium) splendidum Sturm, 1825: 145
Female adult

Copyright © Kirill Makarov

Bembidion (Plataphus) infuscatipenne Netolitzky, 1938
Male adult
Russia, SE Primorie, Lazo Nat. res., ~10 км SW Sokolchi vill., Korpad' vicinity 43°15'N / 134°7'E, 23-27.06.10, leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Microserrullula) luniferum Andrewes, 1924
Female adult
N INDIA, Uttarakhand, 22-23.04.2012, 8 km NW Jaspur, lake shore, 29°21'10.54", 78°48' 30.99", Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Bracteon) stenoderum stenoderum Bates, 1873a: 300
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Trichoplataphus) kara Andrewes, 1921d: 250
Female adult
India, Uttarakhand, 15 km SW New Tehri, 869 m, N30*15'874" E 078*21'587", 18-20.4.2012, A. Anichtchenko leg.; 6 mm; det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) persicum Menetries, 1832
Male adult
Tajikistan, Khoja Obi, Garm, 15.9.2011, Barsevskis A. leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) fontinale (Raffray, 1885)
Male adult
Ethiopia Amhara, Mt. Abuna yosef, n-slope, 3800-3950 m, 12°07’52”N 39°11’39”E, 4.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Bembidion) humerale humerale Sturm, 1825: 176
Female adult
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha” NW part, 17.IV-10.VII.2018, Panin R. leg., L =2.95 mm
Copyright © Denis Khrapov

Bembidion (Princidium) punctulatum Drapiez, 1820
Male adult
Pechoro-Ilychskii N.P. (Печеро-Илычский зап)
Copyright © Kirill Makarov

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platidiolus vandykei Kurnakov, 1960
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Deltomerus (Deltomerus) tatricus L. Miller, 1859: 306
Male adult
N Slovakia; Vysoké Tatry Mts., 10mm
Copyright © Aleš Sedláček

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Platidiolus cf. borealis Zamotajlov, 2001
Female adult
Russia, Khabarovsk reg., Verhnebureinski distr. Bureya river valley, Ussomah river, 22.7.2006 (anal sternite with 3 pairs of setae). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Penetretus nebrioides (Vuillefroy, 1866)
Male adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Diplous (Diplous) sibiricus atratus Habu, 1951
Male adult

Copyright © Kirill Makarov

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Pericompsus (Upocompsus) habitans (Sloane, 1896) Syntypes
Male adult

Copyright © Museums Victoria

Meotachys (Scolistichus) riparius Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Rio Solimões, Ilha de Marchantaria
Copyright © Terry Erwin

Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) scitulus Leconte, 1848
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Tachys) corax Leconte, 1852
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyura (Tachylopha) leleupi (Basilewsky In Basilewsky & Straneo, 1950)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Aenigmatachys rubens Baehr, 2020 Holotype
Male adult
“Australia Black B . . . (unreadable) / G.C. Champion Coll. B.M. 1927-409” (NHM).
Copyright © Martin Baehr

Micratopus anaisae Lemaire, 2021 Holotype
Female adult
Saint-Félix, commune du Gosier, Guadeloupe, 16.210 N, 61.461 O, J. M. Lemaire leg. 15/04/2016. (MNHN). Image from Lemaire, 2021
Copyright © - - -

Tachyura (Tachyura) stevensi (Andrewes, 1925)
Female adult
India, Uttarakhand, 5 km SW Rishikesh, left bank of Ganga riv., 9.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko

Dianotrechus gueorguievi Tian, 2016 Holotype
Female adult
China, Yunnan Province, Ermu Vill., Kunming District, Dashi Dong (Big Rock Cave), 24°49'13"N/102°27'56"E, 1940 m in altitude, XI-8- 2011, B. Petrov leg.” (SCAU). Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Eocnides fragilis Ueno, 1989a: 14
Male adult
Bingzhongluo Township (Niwaluo He, just below Nu Jiang Road, 1630 m, N28.05140°/ E98.59319°, 8 October 2002, D.H. Kavanaugh, P.E. Marek and D.Z. Dong collectors. Image from Deuve et al, 2016
Copyright © - - -

Trechinotus flavocinctus Jeannel, 1962
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanorites magnus Moore, 1972
Male adult
Australia Tas. m1450 Ben Lemond N.P. Hamilton Crags. Meadow Vale
Copyright © Vittorino Monzini

Xiangxius jiangi Tian & X. Huang, 2021 Holotype
Male adult
cave Tangle Dong, suburb of Jishou, Xiangxi Tujia & Miao Autonomous Prefecture, western Hunan Province, 28°18’21”N / 109°39’15”E, 270 m in altitude, 2018-I-15, leg. Xinglong Huang (SCAU)
Copyright © Mingyi Tian

Duvalius (Duvalius) nezelensis Muilwijk & Malek Hosseini, 2021 Holotype
Male adult
Iran Kohgiluyeh Va Boyer Ahmad, Zagros/ Nezel cave, 12.vi.2018 / M. Porebrahim, S. Azizi, Z. Rozbehi, S. Rahideh, & J. Muilwijk leg.”
Copyright © Jan Muilwijk

Deuveopsis (Deuveopsis) lobeliae Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Amhara, Mount Choke, station I (afro-alpine belt, rocks below Lobelia), 10˚41’54’’N, 037˚50’21’’E, 3950 m a.s.l., 21.VIII.2011, E. Quéinnec & C. Reeb leg.
Copyright © Eric Quéinnec

Dongodytes (Dongodytes) mingruoae Tian, Huang & Jia, 2023 Holotype
Male adult
Cave I, Jiudu Xiang, Du’an Yao Autonomous County, Hechi Shi, Guangxi Zhuang Autonomous Region, southern China, 24.27°N / 108.23°E, 333 m in altitude, 2018-IX-25, leg. Mingyi Tian, Sunbin Huang & Mengzhen Chen, in SCAU
Copyright © Mingyi Tian

Trechus (Abunetrechus) bipartitus Raffray, 1880
Male adult
Ethiopia, Amhara, Mt. Abuna Yosef, N-slope, 3800–3950 m, 12°07'52"N, 39°11'39"E, 4.III.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W
Copyright © Joachim Schmidt

Darlingtonea kentuckensis Valentine, 1952
Male adult
Kentucky, Pulaski Co., 10mi NE Somerset, stab, Taylor Cave, west bank Buck Creek 10.11.1949, Valentine, Jones & Diamond (cAA).
Copyright © Alexander Anichtchenko

Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -