Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Anillinus affabilis (Brues, 1902) Lectotype
Male adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm.
Copyright © Igor Sokolov

Winklerites stevanovici Hlavac & Margini, 2017 Paratype
Male adult
Srbija [p], Dimitrovgrad, Petrlaš, Džemanska prop[ast], 02.05.2002 [h], M. Stevanović leg. Image from HLAVÁČ & MAGRINI, 2017
Copyright © - - -

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Rhegmatobius bastianinii Magrini & Casale, 2015 Paratype
Male adult
Italia, Lazio (LT) M.te S. Angelo, Formia, 1200m.
Copyright © Augusto Degiovanni

Stylulus sp.
Unsexed
"US Virgin Islands, St. Croix, 12.vi.1972, W.B. Muchmore Leg." Florida State Collection of Arthropods. "Stylulus sp." det. C.W. Harden 2023.
Copyright © Curt Harden

Stylulus (Stylulus) isabelae Giachino & Sciaky, 2003 Holotype
Male adult

Copyright © Vittorino Monzini

Anillinus alleni Sokolov & Carlton, 2017
Male adult
USA: Arkansas, Stone Co., Ozark National Forest, Blanchard Springs Rec. Area. 27 September 2024. C.W. Harden & K.D. Karns leg. Under rock during rain. / Det. C.W. Harden 2024.
Copyright © Curt Harden

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Furcacampa) versicolor (Leconte, 1848)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Plataphus) prasinum (Duftschmid, 1812)
Female adult

Copyright © Kirill Makarov

Sinechostictus (Sinechostictus) nordmanni (Chaudoir, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Bracteon) stenoderum stenoderum Bates, 1873a: 300
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Sinechostictus (Sinechostictus) ruficornis Sturm, 1825
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) yukonum Fall, 1926: 131
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) transhimalayanum Schmidt, 2018 Paratype
Female adult
“S TIBET, ca. 40km SW of / Yangpachem, 13.VII.2010 / Pijin moraine 5000-5500 m / 29°45’36N 90°17’54E” (CSCHM).
Copyright © Joachim Schmidt

Erwiniana irisculptilis (Erwin, 1994)
Unsexed
ECUADOR, Parque Nacional Yasunì
Copyright © Gianni Allegro

Bembidion (Ecuadion) agonoides Vigna-Taglianti & Toledano, 2008
Female adult
Ecuador, Provincia Cotopaxi, Otonga Nature Reserve, 0°25’15” S, 79°00’23” W, 1800-2000 m
Copyright © Pierre Moret

Bembidion (Ocydromus) atlanticum atlanticum Wollaston, 1854
Male adult
Uzbekistan, Chatkal mt. rng., 6 km NW Irtash, Iertash riv., 41.10.46N 70.17.28E, 1900m, 3.8.2021, Anichtchenko A. leg. (cAA). det. L. Toledano, 2026
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Platidiolus cf. borealis Zamotajlov, 2001
Female adult
Russia, Khabarovsk reg., Verhnebureinski distr. Bureya river valley, Ussomah river, 22.7.2006 (anal sternite with 3 pairs of setae). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
Copyright © - - -

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Deltomerodes ovicollis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Deltomerus (Deltomerus) corax Peyerimhoff, 1922
Male adult
Morocco, Grand Atlas, 2200-2300m, N31.13.09.5 W07.49.29.7 16-17.6.2010 Anichtchenko A. leg. & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Diplous (Diplous) sibiricus atratus Habu, 1951
Male adult

Copyright © Kirill Makarov

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Tachys (Paratachys) micros (Fischer von Waldheim, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pericompsus (Upocompsus) habitans (Sloane, 1896) Syntypes
Male adult

Copyright © Museums Victoria

Elaphropus (Tachylopha) spenceri (Sloane, 1896) Paratype
Male adult

Copyright © Museums Victoria

Elaphropus (Ammotachys) marchantarius Boyd & Erwin, 2016 Paratype
Male adult
Venezuela, San Carlos de Rio Negro
Copyright © Terry Erwin

Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Kiwitachys cf. antarcticus Bates, 1874
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Polyderis) laeva (Say, 1823)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Paratachyta) quadrinotata Baehr, 2016 Holotype
Female adult
320 Ilam Distr., betw. Hodia Khola Valley to Sktia, 250–500 m, Shorea forest, dry, 7 Apr 1988, MARTENS & SCHAWALLER leg. / Nepal-Expeditionen JOCHEN MARTENS (SMNS).
Copyright © Martin Baehr

Tachys (Paratachys) frischi Coulon & Felix, 2009
Female adult
Iran, Khuzestan, 30 km E Shushtar, 5.5.2007, A. Anichtchenko leg. (cAA). det. J. Muilwijk, 2021
Copyright © Alexander Anichtchenko

Epaphiopsis (Epaphiama) sinuata Belousov & Kabak, 2003 Paratype
Male adult
China, Sichuan, right bank of Lanhegou Riv., NW of Mt. Ubaoshan,
E of Ji..i, 2800-3500m 30.6.2000 Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Trechus (Trechus) carrilloi Toribio et Rodriguez, 1997: 283
Male adult

Copyright © Marcos Toribio

Trechus (Epaphius) rivularis (Gyllenhal, 1810)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Typhlotrechus bilimekii hacquenii Sturm, 1853
Male adult
Slovenia, Ljubljana, Gorni lg.Jama, Velika Basica, IV-IX.2003, M. Egger leg. L=7.5 mm
Copyright © Ruslan Panin

Huoyanodytes tujiaphilus Tian & Huang, 2016 Holotype
Female adult
Cave Tujiamei Dong, Huoyan Karst, Huoyan Xiang, Wulongshan Geopark, Longshan County, NW Hunan Province, China, 29°12'20.11"N/109°18'37.21"E, 427 m in altitude, VII-3-2014, leg. Mingyi Tian, Weixin Liu, Haomin Yin, Sunbin Huang & Xinhui Wang, deposited in SCAU. Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Kosswigia insularis Jeannel, 1947
Male adult
Turkey - Konya, Beyseyr Gölü, Cave on the island Aci Akif, 13.VII.1988, leg. Zoia (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Allegrettia boldorii Jeannel, 1928 (as zavattarii Ghidini, 1934)
Male adult
Prealpi Bresciane, Italia
Copyright © Vittorino Monzini

Zhijinaphaenops liuae Deuve & Tian, 2015 Holotype
Male adult
Chine, Guizhou, Guiyang, Xifeng Xian, Shidong Zhen, grotte Hejia Dong, 1198 mètres, 27°2’30,89’’N - 106°31’40,38’’E, Weixin Liu, 13.VII.2013, in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2015
Copyright © Mingyi Tian

Minosaphaenops ollivieri Queinnec, 2008
Male adult

Copyright © Vittorino Monzini

Troglocimmerites zapparolii Casale & Magrini, 2023 Holotype
Female adult
Turchia, Anatolia nord-orientale, vilayet Giresun, strada per Çanakci, 70-80 m s.l.m., 5,8 km dal bivio per Çanakci, 6.V.1987, leg. M. Zapparoli (AVT). Image from Casale & Magrini, 2023
Copyright © - - -

Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons