Anisodactylus (Pseudanisodactylus) sadoensis Schauberger, 1931
Endophallus
Collection ground Sekikawa mura, NiigataPref. 35942
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) punctatipennis A.Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Endophallus
Collection ground Mukawa cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Anisodactylus) karennius (Bates, 1892)
Endophallus
N Vietnam, 40 km WNW of Lao Cai, env. Y Ty, Bat Xat Nat.Park 22°37'36"N 103°37'32"E 4-14.VI.2019 h~1850 leg. D. Fedorenko
Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Collection ground Ashikaga city,Tochigi Pref.Ashikaga～shi 36079
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) lucidus (A. Morawitz, 1863)
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) longitarsis A. Morawitz, 1863: 65
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) lewisi Schauberger, 1936: 9
Endophallus
Collection ground Kikonai cho, Hokkaido Pref. 34925
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) pseudophonoides Schauberger, 1930
Endophallus
Collection ground Tanagura cho, Fukushima pref. 37900
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) pastor niigatanus Schauberger, 1929
Endophallus
Collection ground Himeji city, Hyogo Pref. 32778
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) ussuriensis vicarius Harold, 1878
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Endophallus
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) sinicus Hope, 1845
Endophallus
Collection ground Hokkaido Pref. Kikonai～cho 34925
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Nipponoharpalus discrepans (A.Morawitz, 1862)
Endophallus
Collection ground Chitose city, Hokkaido Pref. pref. 34168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Endophallus
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Endophallus
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883: 239
Endophallus
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Endophallus
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) chalcentus Bates, 1873a: 263
Endophallus
Collection ground Minobu cho, Yamanashi Pref. 35617
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Loboharpalus) platynotus Bates, 1873
Endophallus
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Endophallus
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Carterus (Pristocarterus) angustus (Menetries, 1832)
Endophallus
Iran, Lorestan prov., near Dorud, 5 km SW Chabdar vill. 33.501061°N 49.003441°E 14-15.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Harpalus (Harpalus) semipunctatus Dejean, 1829
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) nevadensis K. Daniel & J. Daniel, 1898d: 65
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) ebeninus Heyden, 1870
Endophallus

Copyright © Alexander Anichtchenko

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Scale bar: 0.6 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Trichotichnus (Bottchrus) nanus Habu, 1954c: 233
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Machozetus lehmanni Menetries, 1848
Endophallus

Copyright © Kirill Makarov

Stenolophus (Stenolophus) agonoides Bates, 1883
Endophallus
Collection ground Kobe city, HyogoPref. 38129
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Stenolophus (Stenolophus) propinquus A.Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Stenolophus (Egadroma) difficilis (Hope, 1845)
Endophallus
Collection ground Kamogawa city, ChibaPref. 34922
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Stenolophus (Egadroma) quinquepustulatus (Wiedemann, 1823)
Endophallus
Collection ground Toride city, Ibaraki pref. 35685
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Bradycellus (Tachycellus) subditus (Lewis, 1879)
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39249
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Bradycellus (Tachycellus) grandiceps (Bates, 1873)
Endophallus
Collection ground Zushi city, Kanagawa pref. 33314
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Psychristus (Nipponobradycellus) lewisi (Schauberger, 1933)
Endophallus
Collection ground Sakauchi mura, GifuPref. 37801
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Acupalpus (Setacupalpus) sobosanus Habu, 1954
Endophallus
Collection ground Minoo city, Oosaka pref. 37075
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Acupalpus (Palcuapus) inornatus Bates, 1873
Endophallus
Collection ground Kamogawa city, ChibaPref. 34922
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Acupalpus (Setacupalpus) hilaris Tschitscherine, 1899
Endophallus
Russia, Evreiskaya AO, Birobidzhan vic., 3.11.2013. leg. Budilov. P.
Copyright © Pavel Budilov

Bradycellus (Tachycellus) glabratus Reitter, 1894d: 125
Endophallus
Russia, Evreiskaya AO, Birobidzhan vic., 3.11.2013. leg. Budilov. P.
Copyright © Pavel Budilov

Anthracus furvinus (Darlington, 1968)
Endophallus

Copyright © Bernd Jaeger

Anthracus derogatus (Walker, 1858)
Endophallus

Copyright © Bernd Jaeger

Anthracus wrasei Jaeger, 2012
Endophallus

Copyright © Bernd Jaeger

Anthracus latus Jaeger, 2012 Holotype Paratype
Endophallus

Copyright © Bernd Jaeger

Anthracus skalei Jaeger, 2015
Endophallus
16 - HT; 17 - India, Kurseong; 18, 21, 23 - Bhutan, Samchi; 19, 22 - India, Cherrapunjee; 20 - India, Tura.
Copyright © Bernd Jaeger

Anthracus weigeli Jaeger, 2015
Endophallus
7, 11 - Nepal, Kathmandu; 8, 10, 12, 15 - India, Similipal; 13 - HT; 9, 14 - Nepal, Nepalganj.
Copyright © Bernd Jaeger

Anthracus anichtchenkoi Jaeger, 2015
Endophallus

Copyright © Bernd Jaeger

Anthracus philippinensis Jaeger, 2015
Endophallus

Copyright © Bernd Jaeger

Acupalpus (Acupalpus) inouyei (Habu, 1980)
Endophallus
Kunashir Island, west coast of Lake Peshchanoe 29.IX.2014 leg. Yu. Sundukov
Copyright © Kirill Makarov

Dicheirotrichus (Cardiostenus) medvedevi (Kabak & Kataev, 1993)
Endophallus

Copyright © Kirill Makarov

Acupalpus (Acupalpus) orszuliki Wrase & Jaeger, 2017 Holotype Paratype
Endophallus
Median lobe of aedeagus, dorsal and lateral aspect. (1-2) Paratype, Syria, Homs env.; Dorsal (3) Holotype.
Copyright © Bernd Jaeger