Pterostichus (Eosteropus) orientalis antiquus (Motschulsky, 1860)
Endophallus
Collection ground Fukushima cho, Hokkaido Pref. 31290
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Rhagadus) thorectes Bates, 1873a: 287
Endophallus
Collection ground Taka cho, Hyogo Pref. 39536
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Eosteropus) japonicus (Motschulsky, 1860)
Endophallus
Collection ground Tomakomai, Hokkaido Pref. 30926
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Endophallus
Collection ground Rebun Is. Hokkaido Pref. 27191
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Lyrothorax) yoritomus Bates, 1873
Endophallus
Collection ground Maizuru city, Kyoto 33744
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Nialoe) sumondakensis Sasakawa, 2005 Holotype
Endophallus
Mt. Sumon, 1100 m, Niigata Pref., Japan, 4. IX. 1969, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-270
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Poecilus (Poecilus) festivus (Chaudoir, 1868)
Endophallus

Copyright © Kirill Makarov

Pterostichus (Morphnosoma) chechcirensis Lafer, 1979: 28 Paratype
Endophallus

Copyright © Pavel Budilov

Pterostichus (Phonias) kutensis Poppius, 1905
Endophallus
Хабаровский край, Верхнебуреинский район, окр. пос. Софийск. Лиственничная тайга, h~1000м, 4.09.2013 (Habarovsk prov., Verhnebureinsk distr., Sofiisk vill.)
Copyright © Pavel Budilov

Pterostichus (Eosteropus) orientalis nigromontanus Lafer & Budilov, 2015 Holotype
Endophallus
Россия: Приморский край, Хасанский р-н, заповедник «Кедровая Падь», 27.VIII 2004, В.Н. Кузнецов
Copyright © Pavel Budilov

Pterostichus (Vietosteropus) dalatensis Fedorenko, 2017
Endophallus

Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) kontumensis Fedorenko, 2017
Endophallus

Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) annamita s.str. (Straneo, 1939)
Endophallus

Copyright © Dmitri Fedorenko

Poecilus (Ancholeus) baeticus baeticus
Endophallus
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Poecilus (Poecilus) crenulatus crenulatus (Dejean, 1828)
Endophallus
Spain, Leon, Qiuntanilla de Samoza, 3.IV.2004, J.L. Zapata leg.
Copyright © Alexander Anichtchenko

Poecilus (Parapedius) decipiens decipiens
Endophallus
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Lesticus (Triplogenius) insignis Gestro, 1883
Endophallus
N Borneo, Sabah, Keningau distr., Mt. Trus Madi, 7-8.4.2013, 1250m.
Copyright © Alexander Anichtchenko

Pterostichus (Pseudomaseus) fuscicornis (Reiche & Saulcy, 1855)
Endophallus
Iran, East Azerbaijan prov., near Kandovan vill. 37.748092°N 46.286895°E 21-22.V.2017 leg.D.Kasatkin
Copyright © Kirill Makarov

Trigonotoma nitidicollis Chaudoir, 1868
Endophallus
Thailand, Lampang Wang Nuea, 10.V.2011
Copyright © Alexander Anichtchenko

Lesticus (Triplogenius) gregori Kuntzen, 1911
Endophallus
Philippines, Mt. Polis, Mt Province, N Luzon.
Copyright © Alexander Anichtchenko

Lesticus (Triplogenius) xiaodongi Zhu, Shi & Liang, 2018
Endophallus
Image from Zhu, Shi & Liang, 2018
Copyright © - - -

Lesticus (Triplogenius) violaceous Zhu, Shi & Liang, 2018
Endophallus
Image from Zhu, Shi & Liang, 2018
Copyright © - - -

Lesticus (Lesticus) bii Zhu, Shi & Liang, 2018
Endophallus
Image from Zhu, Shi & Liang, 2018
Copyright © - - -

Lesticus (Triplogenius) auripennis Zhu, Shi & Liang, 2018
Endophallus
Image from Zhu, Shi & Liang, 2018
Copyright © - - -

Lesticus (Lesticus) solidus Roux & Shi, 2011
Endophallus
Image from Zhu, Shi & Liang, 2018
Copyright © - - -

Trigonotoma digitata Zhu, Shi & Liang, 2020
Endophallus
Image from Zhu, Shi & Liang, 2020
Copyright © - - -

Trigonotoma constricta Zhu, Shi & Liang, 2020
Endophallus
Image from Zhu, Shi & Liang, 2020
Copyright © - - -

Pterostichus (Morphnosoma) silvestris Sun & Shi, 2018 Paratype
Endophallus
Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) maryseae Sun & Shi, 2018 Paratype
Endophallus
Image from Sun et al, 2018
Copyright © Hongliang Shi

Poecilus (Poecilus) cupreus (Linne, 1758)
Endophallus
Turkije : Ardahan; Ardahan 7KM E Karst 13-7-1992 1900m. Heinz W. leg.
Copyright © Jan Muilwijk

Poecilus (Poecilus) cupreus erythropus Dejean, 1828
Endophallus
Libanon J.Sannin Sud 16-5-2000 1300 I. Rapuzzi leg.
Copyright © Jan Muilwijk

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Endophallus
Agri, Balık Gölü, Taşlıçay, 2250 m, 17.07.76; W. Heinz leg.
Copyright © Jan Muilwijk

Orthomus (Orthomus) perezii (Martinez y Saez, 1873)
Endophallus
Spain, Cuenca, Montes Universales, pto del Cubillo 1620m, 23.5.2006, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Pterostichus (Badistrinus) neglectus A.Morawitz, 1862
Endophallus
Khabarovsk Territory, Khawar Stream below cordon Bichi, 24.VII.2021, Yu. Sundukov & L. Sundukova leg.
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930
Endophallus
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930
Endophallus
Primorye Territory, Khasansky district, 10.7 km east of Khasan, Cape Ostrovok Falshivy, 42°26’54”N 130°46’47”E 23.V.2022, K. Makarov leg.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
South Primorye, Kedrovaya Pad nature reserve, mouth of Kedrovaya River, swamp 43°04'47"N 131°36'42"E 1.VI.2019 leg. K. Makarov L= 7.1
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
Hirotuka Kanagawa-Ken 23.jul.1958 S.Nomura L= 7.4 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.2 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Amara (Zezea) plebeja (Gyllenhal, 1810)
Endophallus
Collection ground Chitose city, Hokkaido Pref. pref. 34882
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Bradytus) sinuaticollis A. Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Bradytus) simplicidens A. Morawitz, 1863: 60
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Bradytus) majuscula (Chaudoir, 1850)
Endophallus
Collection ground Monbetsu cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) macronota Solsky, 1875
Endophallus
Collection ground Tomakomai city, Hokkaido Pref. 37172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) hiogoensis (Bates, 1873)
Endophallus
Collection ground Tsushima, NagasakiPref. 35695
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) gigantea (Motschulsky, 1844)
Endophallus
Collection ground Kagoshima city, Kagoshima Pref. 38979
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Reductocelia) lucens Baliani, 1943: 44
Endophallus
Collection ground Sadamitsu cho, Tokushima Pref. 38262
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Reductocelia) chalcophaea Bates, 1873a: 292
Endophallus
Collection ground Tomakomai city, Hokkaido Pref. Tomakomai～shi 37172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) ussuriensis Lutshnik, 1935a: 260
Endophallus
Collection ground Otobe cho, Hokkaido Pref. 35273
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) obscuripes Bates, 1873a: 294
Endophallus
Collection ground Shibecha cho, Hokkaido Pref. 38493
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) familiaris (Duftschmid, 1812)
Endophallus
Collection ground Kiyosato cho, Hokkaido Pref. pref. 38785
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) congrua A. Morawitz, 1862b: 326
Endophallus
Collection ground Ise city, Mie Pref. 37125
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) chalcites Dejean, 1828: 462
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Zezea) davidi Tschitscherine, 1897b: 67
Endophallus
Median lobe of aedeagus of male China, Gaoligong Shan. CASENT8125447 Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Amara (Amara) silvestrii Baliani, 1937: 179
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Amara (Amara) shaanxiensis Hieke, 2002
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Amara (Pseudoamara) birmana Baliani, 1934c: 189
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Amara (Amara) communis (Panzer, 1797)
Endophallus
det. K. Makarov
Copyright © Kirill Makarov

Amara (Bradytus) elegantula Tschitscherine, 1899
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Amara (Bradytus) dissimilis Tschitscherine, 1894a: 404
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Amara (Bradytus) chalciope (Bates, 1891)
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Amara (Xenocelia) sikkimensis Andrewes, 1930: 24
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Zabrus (Pelor) kraatzi Andujar & Serrano, 2000
Endophallus
Turkey, Abant, 1.6.1991 Skoupy leg. det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Zabrus (Pelor) spinipes rugosus Menetries, 1832
Endophallus
Iran, W.Azerbaijan vill. Urmia lake distr. near Rajan vill. 23 05 2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Zabrus (Pelor) spinipes rugosus Menetries, 1832
Endophallus
Iran, W.Azerbaijan vill. Urmia lake distr. near Rajan vill. 23 05 2017 leg. D.Kasatkin. det. Anichtchenko A. 2018
Copyright © Kirill Makarov

Amara (Curtonotus) gigantea (Motschulsky, 1844)
Endophallus
Russia, Primorskii reg.
Copyright © Alexander Anichtchenko

Amara (Amara) pulpani Kult, 1949
Endophallus
W Ukraine, NE Carpathians, Lviv reg., Skole Beskids Mts, Grabnyk Range h=670 m,
Copyright © Denis Khrapov

Amara (Leironotus) rotundicollis (Schaufuss, 1862)
Endophallus
Spain, Sierra Guadarrama, 2 km E Navacerrada 1900m, 1.9.2002, Anichtchenko A. leg. & det. (cAA).
Copyright © Alexander Anichtchenko