Carterus (Pristocarterus) angustus (Menetries, 1832)
Endophallus
Iran, Lorestan prov., near Dorud, 5 km SW Chabdar vill. 33.501061°N 49.003441°E 14-15.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Machozetus lehmanni Menetries, 1848
Endophallus

Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Collection ground Ashikaga city,Tochigi Pref.Ashikaga～shi 36079
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) lucidus (A. Morawitz, 1863)
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) longitarsis A. Morawitz, 1863: 65
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) lewisi Schauberger, 1936: 9
Endophallus
Collection ground Kikonai cho, Hokkaido Pref. 34925
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) pseudophonoides Schauberger, 1930
Endophallus
Collection ground Tanagura cho, Fukushima pref. 37900
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) pastor niigatanus Schauberger, 1929
Endophallus
Collection ground Himeji city, Hyogo Pref. 32778
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) ussuriensis vicarius Harold, 1878
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Endophallus
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) sinicus Hope, 1845
Endophallus
Collection ground Hokkaido Pref. Kikonai～cho 34925
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Nipponoharpalus discrepans (A.Morawitz, 1862)
Endophallus
Collection ground Chitose city, Hokkaido Pref. pref. 34168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Endophallus
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Endophallus
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883: 239
Endophallus
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Endophallus
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) chalcentus Bates, 1873a: 263
Endophallus
Collection ground Minobu cho, Yamanashi Pref. 35617
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Loboharpalus) platynotus Bates, 1873
Endophallus
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Endophallus
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) semipunctatus Dejean, 1829
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) nevadensis K. Daniel & J. Daniel, 1898d: 65
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) ebeninus Heyden, 1870
Endophallus

Copyright © Alexander Anichtchenko

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Scale bar: 0.6 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Trichotichnus (Bottchrus) nanus Habu, 1954c: 233
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim