Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro

Typhlocharis laurentii Magrini, 2000 Paratype
Unsexed
España, Almeria, Sierra Almagrera ; carretera N332, 30.XI.98, leg. Fancello (MNHN, Antoine Mantilleri, 2016)
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Scotodipnus (Binaghites) affinis affinis (Baudi di Selva, 1871)
Male adult
Italy, Emilia, M. Penna (PR), 1400m, 24.5.1999, leg. F. Angelini
Copyright © Alexander Anichtchenko

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Anillus florentinus florentinus Dieck, 1869
Unsexed
Emilia Romagna, Sasseleone (BO), 26.3.2002 350m, A.Degiovanni leg. (cAA)
Copyright © Alexander Anichtchenko

Anillus florentinus andreinii Jeannel, 1937
Unsexed
Italy, Toscana, Giuncarico 8.12.96 leg Degiovanni A. (cAA)
Copyright © Alexander Anichtchenko

Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
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Rhegmatobius strictus (Baudi di Selva, 1891)
Male adult
Sardegna, Gennargentu Mte., Spada Mt., 1000, 27.4.2003 A. Degiovanni leg. (cAA).
Copyright © Alexander Anichtchenko

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
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Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Diplocampa) fumigatum (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Sinechostictus (Sinechostictus) frederici (G. Muller, 1918)
Female adult
Spain, Jaen prov., Arroyo de la Piedra del Aguila; 3-4.05.2014, 37.683885, -3.485062, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Nothonepha) tetrapholeon Maddison, 2014 Paratype
Male adult
Argentina: Neuquén: Arroyo Queñi at Lago Queñi, 830m, 40.1575°S, 71.721°W, / 10–11.ii.2007. DRM 07.035. D.R. Maddison, S.A.Roig. Scale bar 1 mm. Image from Maddison D.R., 2014.
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Bembidion (Semicampa) chaudoirii Chaudoir, 1850b: 179
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult

Copyright © Kirill Makarov

Philipis cf. lawrencei Baehr, 2007
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Peryphophila) problematicum Schmidt, 2018 Holotype
Male adult
“NEPAL, Ghatte / Kirola – Khola. Camp / 3500m. Pasang 12.5.88” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) ahrensi Schmidt, 2018 Paratype
Male adult
“Uttar Ganga Ebene / bei Dhorpathan / 3000 m, 6.-8.3.”, “C-NEPAL-Himalaya / Dhaulagiri-Mts. / leg. Ahrens 1994” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) cordicolle Jacquelin da Val, 1852
Male adult
Turkey, Kemer distr., Mts.Taurus, Ulupinar riv.,36*27'01N 30*26'14E 21.9.2014, I. Enuschenko leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Pseudophilochthus) gemmulipenne (Wollaston, 1877)
Male adult

Copyright © David Maddison

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Platidiolus vandykei Kurnakov, 1960
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Patrobus cinctus Motschulsky, 1860
Female adult
Jewish Aut. Reg., Obluchie Distr., 3-5 km upper Radde Vill. on Lagar Riv., forest 24.07.2004 I.Melnik leg.
L=9.5 mm
Copyright © Kirill Makarov

Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
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Diplous (Diplous) sibiricus atratus Habu, 1951
Male adult

Copyright © Kirill Makarov

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Barytachys) tripunctatus (Say, 1830)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Tachys (Paratachys) micros (Fischer von Waldheim, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Setitachys macrops Baehr, 2016 Paratype
Female adult
Qld. Bentinck Is. Rokoti, 16 June 1963. P. Aitken, N. B. Tindale
Copyright © Martin Baehr

Tachyxysta howdenorum Boyd & Erwin, 2016 Holotype
Male adult
México, Chiapas, El Aguacero.
Copyright © Terry Erwin

Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Nothoderis rufotestacea Hayward, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Tachys) corax Leconte, 1852
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyura (Tachyphanes) patruelis Baehr, 2016 Holotype
Male adult
Turkey, Pr. Siirt Botan Cayi-Tal 500 m, 23.4.1983 Leg. W. Schacht (Zoologische Staatssammlung München).
Copyright © Martin Baehr

Tachyta (Eurytachyta) laticollis Baehr, 2016 Holotype
Female adult
Borneo, Sabah, Kinabalu NP, HQ vic., 1550 m, 22.–25.V.2005, R. GRIMM (CBM).
Copyright © Martin Baehr

Costitachys inusitatus Erwin, 1974 Holotype
Male adult
Santarem, Para, Brazil, IV.27.1963 FG Werner (MCZ)
Copyright © - - -

Pseudaphaenops jacobsoni Pliginskiy, 1912: 505
Male adult

Copyright © Kirill Makarov

Cimmeritodes (Xiangcimmerites) zhongfangensis Deuve & Tian, 2016 Holotype
Female adult
Chine, Hunan, Huaihua Shi, Zhongfang Xian, Huangyan Xiang, Fujia Cun, grotte Jinji Dong, 890 m, 27°28’20’’N - 110°4’7’’E (Tian Mingyi, Liu Weixin, Wang Xinhui, Tang Mingruo, 1.VII.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -

Trechepaphiopsis monochaeta Deuve & Kavanaugh, 2016 Holotype
Male adult
China, Yunnan, Gongshan County, Qiqi Trail at No. 12 Bridge Camp area, 16.3 airkm W of Gongshan, N27.71503°/E98.50244°, 2775 m. Image from Deuve et al, 2016
Copyright © - - -

Mexaphaenops elegans Barr, 1967
Female adult
Sierra Gorda, Pinal de Amoles, 2700m, Ojo de Agua - Mexico
Copyright © Vittorino Monzini

Cimmeritodes (Zhecimmerites) zhejiangensis Deuve & Tian, 2015 Holotype
Male adult
Chine, Zhejiang occidental, Quzhou City, Changshan Xian, Xinjiao Xiang, Dongling Cun, grotte Niujiao Dong, 546 mètres, 29°09’44,86’’N - 118°40’26,18’’E, Sunbin Huang & Xinhui Wang leg., 4.X.2014, in coll. South China Agricultural University, à Canton (Chine). Image from Deuve & Tian, 2015
Copyright © Mingyi Tian

Zhijinaphaenops jingliae Deuve & Tian, 2015
Male adult
Guizhou: Guiyang, Xifeng, Xishan, Shenli, Mafen Dong cave (息烽县西山镇马粪洞), 27°04'N, 106°37'E, 1157 m, 2020–II–29, leg. Guangyuan Cheng
Copyright © Mingyi Tian

Duvalius (Neoduvalius) lohaji Queinnec & Ollivier, 2021 Holotype
Male adult
Mracna Pecina, Livno, Prolog Planina, Bosnia and Herzegovina
Copyright © - - -

Himalotrechus humeratus Belousov, Kabak & Schmidt, 2019 Holotype
Male adult
India, Arunachal Pradesh, N of Bomdila, NW Chander, 10–15 km NW of, 27º25´ N 92º22´ E, ca 3200 m, 18 VI.2008 / Reuter (coll. J. Schmidt).
Copyright © - - -

Taniatrechus xeniae Belousov & Koval, 2023 Holotype
Female adult
“W Caucasus, Abkhazia, Bzyb Mt. Range, 3.4 km NEE of Mt. Tshiptshira, 2,220 m, Prostornaya Cave (= L-07 Cave), at a depth of 50 m, traps, 31.VIII.2010 – 11.VIII.2011. X.N. Gorbunova, T.V. Petrova leg.” (ZIN).
Copyright © - - -

Perileptus (Perileptus) melas Jeannel, 1926
Male adult
Philippines, Mindanao, Ugwad Falls (Kapuka), Caraga, Davao Or., 260m, 7.440359, 126.434027, 29–30.IV.2023, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
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Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
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Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
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Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -