Harpalus (Harpalus) flavescens (Piller & Mitterpacher, 1783)
Male adult
CZ, S Moravia, Bzenec; 11mm
Copyright © Aleš Sedláček

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Male adult
Image from Kataev, 2023
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Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest, on the sands, 17.V-25.VI.2024, R.Panin leg., L=13 mm
Copyright © Ruslan Panin

Harpalus (Harpalus) pumilus Sturm, 1818: 77
Male+Female adult
France, forêt de Fontainebleau (77300), 15/7/1985, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) pusillus (Motschulsky, 1850)
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) pumilus Sturm, 1818: 77
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) michaili Kataev, 1990 Paratype
Unsexed
Image courtesy of Biological Museum, Lund University
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Harpalus (Harpalus) masoreoides Bates, 1878
Female adult
Kirgizia, 12km from Beshkek, Alamedin riv., 7.7.1990 Anichtchenko A leg. (cAA) det. Kataev; 4.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) acupalpoides Reitter, 1900
Male adult
Mongolia, 2000m, 40km SW Arvayheer, 15-16.7.2006 Anichtchenko A. leg. & det. (cAA) 4.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) longipalmatus Mordkovitsh, 1969
Male adult
Russia, Chita reg., Daurian NP., Zun-Torey lake, 1.6.2002 Anichtchenko A. leg. (cAA) 4.3 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) clarkei Kataev & Schmidt, 2020
Male adult
Ethiopia, Mt. Kaka. Image from Kataev et al, 2020
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Harpalus (Harpalus) minutissimus Facchini, 2015 Holotype
Male adult

Copyright © Sergio Facchini

Harpalus (Harpalus) sinuaticollis Facchini, 2015 Holotype
Male adult

Copyright © Sergio Facchini

Harpalus (Harpalus) impressus Roth, 1851 Lectotype
Male adult
Abyssinia. Image from Kataev, 2021
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Harpalus (Harpalus) merkli Kataev, 2021 Holotype
Male adult
Abyssinia Kovács”, “Tschertscher [Chercher] 1911.X ( TMB).
Copyright © - - -

Harpalus (Harpalus) pseudoasemus Kataev, 2021 Paratype
Male adult
Kenya Naivasha , Rift Valley 1.900 m ”, “ Muséum de Paris Mission de l’Omo C. Arambourg P.-A. Chappuis & R. Jeannel 1932–33”, “ Harpalus asemus m. P. Basilewsky det.” ( MNHN) .
Copyright © - - -

Harpalus (Harpalus) asemus Basilewsky, 1946
Male adult
left - Yemen; right - Tanzania. Scale bars = 1.0 mm. Image from Kataev, 2021
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Harpalus (Harpalus) fuscoaeneus Dejean, 1829
Male adult
S Africa. Image from Kataev, 2021
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Harpalus (Harpalus) australasiae Dejean, 1829 Holotype
Female adult

Copyright © - - -

Harpalus (Afroharpalus) poussereaui Facchini & Giachino, 2011 Holotype
Male adult
La Réunion 974, Coll : J. Poussereau, Les Makes, Ft de Bon Accueil, Cadavre tangue [Tenrec ecaudatus (Schreber, 1778)], 10.02.2009" (MNHN). 6.7 mm; Image from Facchini & Giachino, 2011
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Harpalus (Afroharpalus) natalensis Boheman, 1848
Male adult
Image from Kataev, 2023
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Harpalus (Afroharpalus) nyassicus Basilewsky, 1946
Male adult
S Malawi, Mt. Mulanje, Lichenya Plateau (7.0 mm)
Copyright © Sergio Facchini

Harpalus (Afroharpalus) malawicus Facchini, 2023 Holotype
Male adult
Coll. Mus. Tervuren, Malawi North. Reg., Nyika Plateau, XII.1981 - R. Jocqué, Lake Kaulime, 2200 m, 6/19.XII.1981 (MRAC).
Copyright © Sergio Facchini

Harpalus (Afroharpalus) defector Peringuey, 1896
Male adult
Mesklip, S of Springbok (7.0 mm)
Copyright © Sergio Facchini

Harpalus (Afroharpalus) namibicus Facchini, 2023 Holotype
Male adult
S Namibia 5.1.2012, Hardap distr., 7 Km W Mariental, Koha GH, 1250 m, Murzin (CF).
Copyright © Sergio Facchini

Harpalus (Harpalus) honestus (Dufischmid, 1812)
Male adult
length: 7,4-10,6 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2007
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Harpalus (Harpalus) neglectus Audinet-Serville, 1821
Female adult
France, Ile d’Oléron (17) 12/6/2006, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) honestus (Dufischmid, 1812)
Male adult
CZ, NE Moravia, Moravskoslezské Beskydy - Smrk; 9mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) attenuatus Stephens, 1828a: 152
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) bellieri Reiche, 1861a: 201
Male+Female adult
France 20122 Corse Quenza plateau du Cuccione 08-06-2007
Copyright © Pascal Deschamps

Harpalus (Harpalus) nevadensis K. Daniel & J. Daniel, 1898
Male adult
Spain, Sierra Nevada, 5km N Capileira, 1800-2000m, 25.4.2003, Anichtchenko A. leg & det. (cAA) 9 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) sulphuripes goudotii Dejean, 1829
Male adult
Algeria, Mostaganem, 30.12.1972, V. Orlov leg. (cAA) 7mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) nevadensis K. Daniel & J. Daniel, 1898d: 65
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) dissitus Antoine, 1931
Male adult
Morocco, Grand Atlas Mts., pass near Oukaimeden, 2800-2900m, 18-20.6.2010 Anichtchenko A. leg. (cAA) 9mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) rufipalpis wagneri Schauberger, 1926
Male adult
Spain, Sierra Nevada, rio Trevelez, 1200m, 6km abajo Trevelez, 7.4.2004, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) decipiens decipiens
Male+Female adult
Espagne Alto de Morredero sierra de Teleno 21-05-2001 et Cabeza de Manzaneda 24-05-2001
Copyright © Pascal Deschamps

Harpalus (Harpalus) bellieri Reiche, 1861
Male adult
Italy, Sardinia, Fonni (NU), Mt. Bruncu Spina m 1800, 20.V.2005
Copyright © Gianni Allegro

Harpalus (Harpalus) decipiens decipiens
Male adult
Portugal, Serra Estrela, 5.VI.1993
Copyright © Gianni Allegro

Harpalus (Amblystus) wohlberedti Emden & Schauberger, 1932
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) spadiceus Dejean, 1829
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Harpalus (Harpalus) somnulentus Dejean, 1829
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Harpalus) latus (Linne, 1758)
Male adult
length: 10,2 mm
Russia., Sverdlovskaya area, the vicinities of Ekaterinburg city, Verkhnemakarovo vill., floodplain meadow
6.07.2005
Zinovyev E. leg.
Copyright © Evgeniy Zinoviev

Harpalus (Harpalus) progrediens Schauberger, 1922
Male adult
length: 8,4 mm
Russia., Sverdlovskaya area, Beloyarskyi distr., Chernousovo vill.
2.09.2007
Vlasov V. leg.
Copyright © Evgeniy Zinoviev

Harpalus (Harpalus) xanthopus winkleri Schauberger, 1923
Male adult
Ивановская обл.,Савинский р-н, Клязьминский заказник
Russia, Ivanovo prov., Savinsk distr.
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Harpalus (Harpalus) marginellus Gyllenhal, 1827: 432
Male adult
Moravia, Bílé Karpaty, 10 mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) torridoides Reitter, 1900a: 81
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) latus (Linne, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) xanthopus winkleri Schauberger, 1923
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) ussuricus Mlynar, 1979: 97
Male adult
Ю Приморье, Хасанский р-н, 7.5 км Ю п.Барабаш, устье руч. Олений 43°07'01"N 131°29'15"E 27.V.2015 leg. К.Макаров L=8.7 mm
Copyright © Kirill Makarov

Harpalus (Harpalus) solitaris Dejean, 1829a: 337
Female adult
W-Ukraine, Transcarpathian reg., Chornagora Massif, 8 km NE Bohdan vill., h=800m, 9-29.VI..2014, Panin R. leg. L=10.5 mm
Copyright © Ruslan Panin

Harpalus (Harpalus) rufiscapus Gebler, 1833
Male adult
Russia, Irkutsk reg., Alarskii distr., vic. Alar vill. 26-28.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) herbivagus Say, 1823
Male adult
USA (IL), Tazewell Co., Morton, Ulrich N.R. 10.8.1990 (cAA) 10 mm
Copyright © Alexander Anichtchenko

Harpalus (Anamblystus) torridoides Reitter, 1900
Male adult
Image from Kataev, 2023
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Harpalus (Anamblystus) luteicornis (Dufischmid, 1812)
Female adult
Latvia, Ilukste, 6.2010 M. Janovska leg (DUBC). det. Anichtchenko A. 2023
Copyright © Alexander Anichtchenko

Harpalus (Anophonus) pterostichus (Reitter, 1900)
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 10mm; det. I. Kabak, 2021
Copyright © Alexander Anichtchenko

Harpalus (Anophonus) cyanopterus (Tschitscherine, 1897)
Male adult
Image from Kataev, 2023
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Harpalus (Harpalus) ingenuus Tschitscherine, 1898a: 179
Male adult

Copyright © Kirill Makarov

Harpalus (Aristoharpalus) ingenuus Tschitscherine, 1898
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) dispar Dejean, 1829
Male adult
France, L’Albaron, 13123 Pascal 9/10/2001, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) dispar splendens (Gebler, 1829)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) petri Tschitscherine, 1902e: 596
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) kabakianus Kataev, 1988: 769
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) pygmaeus Dejean, 1829
Female adult
Greece bor. occ., 15 km W of Metsovo, Petra, 9.6.2008, leg. Orszulik
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) microthorax (Motschulsky, 1849)
Male adult
Spain, Toledo, lagunas de Quero, 28.7.2008, Anichtchenko A. leg & det. (cAA) 10 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) punctatostriatus Dejean, 1829
Female adult
Spain, Teruel, Montes Universales, pto del Portillo, 1780m, 31.5.2006, Anichtchenko A. leg & det. (cAA) 10mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) punctatostriatus Dejean, 1829
Male adult
Italy, Sardinia, Arborea (OR), 9.VI.1978
Copyright © Gianni Allegro

Harpalus (Harpalus) nigrans A. Morawitz, 1862
Female adult
Russia, Irkutsk vic., Mamony vill., Irkut riv., 11.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) sinuatus Tschitscherine, 1893
Male adult
Russia, 14 from Irkutsk by Aleksandrovskii road, 9.7.1994, Anichtchenko A. leg. & det. 6.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Asioharpalus) nigrans A. Morawitz, 1862
Male adult
Image from Kataev, 2023
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Harpalus (Harpalus) opacipennis (Haldemann, 1843)
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Harpalus (Harpalus) lederi Tschitscherne, 1899
Female adult
Buryatia, Hamar-Daban Mts., Taltsy riv., 1700m 19-25.5.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) cautus Dejean, 1829
Male adult
USA Washington, Thurston Co., 19.8.2005 (cAA) 10.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Bactroharpalus) obnixus Casey, 1924
Male adult
Image from Kataev, 2023
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Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
NW Podolian Upland, W UKRAINE,<br /> Lviv Reg., Zolochiv distr., near Gologirki vill., 30.04-19.05.07 Panin R.leg.<br /> L= 15mm
Copyright © Ruslan Panin

Harpalus (Harpalus) caspius (Steven, 1806)
Male adult
W UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa vill., Panin R. leg. 1.V-16.VI.2007
male, 13mm
Copyright © Ruslan Panin

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
Moravia, Kobylí; 11 mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) dimidiatus (P. Rossi, 1790)
Male+Female adult
France 21410 Pont de Pany Sainte-Marie-sur-Ouche 30-06-1984 et 18130 Dun sur Auron 18-05-2012
Copyright © Pascal Deschamps

Harpalus (Baryharpalus) caspius (Steven, 1806)
Male adult
Image from Kataev, 2023
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Harpalus (Harpalus) foveiger Tschitscherine, 1895
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) autumnalis (Duftschmid, 181)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) reflexus anadoluensis Kataev, 1993
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Brachyharpalus) kazanensis Jedlicka, 1958
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Brachypangus) antonowi (Tschitscherine, 1898)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Calathoderus) potanini Tschitscherine, 1906
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) chalcentus Bates, 1873a: 263
Male adult
Collection ground Minobu cho, Yamanashi Pref. 35617
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) chalcentus Bates, 1873a: 263
Endophallus
Collection ground Minobu cho, Yamanashi Pref. 35617
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Male adult
NW Caucas, Krasnodar reg., SE Goryachii Kluch, леса между пос. Асфальтовая Гора и Садовый (нежилой) 21.V.2012 leg. М. Смирнов
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Female adult
SK, S Slovakia, Veľký Krtíš; 12mm
Copyright © Aleš Sedláček

Harpalus (Caloharpalus) chalcentus Bates, 1873
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) chrysopus abasinus Rost, 1892
Male adult
Abkhasia, Bzybskii mt. rng., 1,8 km NE Napra
Абхазия, хр. Бзыбский, 1,8 км NE г. Напра, 2240 м, 43°19' N / 40°33' E, 18.VI.2004, А. Гусаков leg.
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Harpalus (Harpalus) chrysopus chrysopus
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Caucasoharpalus) aeneipennis (Faldermann, 1836)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Cephalophonus) cephalotes Fairmaire & Laboulbene, 1854
Male adult
SE Bulgaria, Primorsko; 16mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) kryzhanovskii Kataev, 1988: 772
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) kryzhanovskii Kataev, 1988: 772
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Male adult
Portugal, Algarve Portimao, 25/9/1983, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Cryptophonus) melancholicus Dejean, 1829
Male adult
Italy, Patro di Moncalvo (Asti).
Copyright © Gianni Allegro

Harpalus (Cryptophonus) fulvus Dejean, 1829
Male adult
Italy, Sardinia, Chia (CA), Stagno Campana, 29.VIII.1994
Copyright © Gianni Allegro

Harpalus (Cryptophonus) fulvus Dejean, 1829
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) pulvinatus lubricus Reitter, 1900
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) udege Lafer, 1989: 189
Male adult
Russia, Baikal lake, Listvianka vill., 2.8.1992, Anichtchenko A. leg. & det. 6.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) vittatus Gebler, 1833
Male adult
Tuva, Tes-Hem13.6.1948 (cAA) 6.5 mm; det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Harpalus (Diaharpalus) vittatus Gebler, 1833
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) atratus Latreille, 1804a: 360
Male adult
Germany, Bavaria, Eußenheim, F. Boetzl leg. & det. 19.07.14
Copyright © Fabian Bötzl

Harpalus (Drymoharpalus) atratus Latreille, 1804
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) ebeninus Heyden, 1870
Male adult
Spain, Picos de Europa, Macizo central, Llagu Bajero, 1890m, 6-7.8.2003, Anichtchenko A. leg & det. (cAA) 11mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) ebeninus Heyden, 1870
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Epiharpalus) punctipennis Mulsant, 1852
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Epiharpalus) ebeninus Heyden, 1870
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Eremoharpalus) medvedevi Kataev, 2006
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) lewisii Leconte, 1865
Male adult
Canada, Québec Berthierville Berthier 20-05-1978
Copyright © Pascal Deschamps

Harpalus (Euharpalops) fraternus Leconte, 1852
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) aequicollis Motschulsky, 1844
Female adult
Russia, 14 km from Irkutsk by Aleksandrovskii road, 9.7.1994, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) heyrovskyi Jedlicka, 1928
Male adult
Russia, Altaiskiy reg., Zmeinogorsk, 11.6.1984 (cAA) 10.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) compressus Motschulsky, 1844: 213
Male adult
SW Ukraine, Odessa reg., Ivanovka distr., Gudevichevo vill., steppe, 13.VI-11.VII.2022, A.Gontarenko leg. L=12 mm
Copyright © Ruslan Panin

Harpalus (Glanodes) obliquus G.Horn, 1880
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Haloharpalus) salinulus Reitter, 1900 Syntypes
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) brevicornis Germar, 1824
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Male adult
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Endophallus
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) froelichii Sturm, 1818: 117
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) hirtipes (Panzer, 1796)
Female adult
13.5 mm; Russia, Volgograd, Kamyshensky, Golubaja river valley, 150 m, leg. E.Alexeev, 15.-19.IV.2000; det. Wrase, 2016. Photo: U.Schmidt, 2017
Copyright © Udo Schmidt

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) brevis Motschiulsky, 1844
Male adult
Russia, Irkutsk reg., Zalari distr., Tungui vill. 20-24.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) macronotus Tschitscherine, 1893
Male adult
Russia, Irkutsk reg., Zalari distr., Tungui vill. 20-24.6.1997, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) zabroides Dejean, 1829
Male adult
E Siberia, Irkutsk, downtown, 20.6.2001, Anichtchenko A. leg & det. (cAA) 15 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) tichonis Jakobson, 1907
Male adult
Siberia, Chita reg., 30 km SE Hilok vill., 28.5.2002 Anichtchenko A. leg & det. (cAA) 11.5mm
Copyright © Alexander Anichtchenko

Harpalus (Haploharpalus) macronotus Tschitscherine, 1893a: 372
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) fuscipalpis Sturm, 1818
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Harpalus) viridanus viridanus (as fuscipalpoides)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) fuscicornis Menetries, 1832: 134
Female adult
Germany, Iphofen, 2018, leg. & det. Boetzl
Copyright © Fabian Bötzl

Harpalus (Harpalobius) viridanus Motschulsky, 1844
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) italus Schaum, 1860c: 583
Male adult

Copyright © Gianni Allegro

Harpalus (Harpalus) circumpunctatus Chaudoir, 1846a: 171
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) stevenii Dejean, 1829a: 242
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) terrestris (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) hospes hospes
Male adult
Slovakia, Horné Strháre; 12 mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) italus Schaum, 1860c: 583
Male adult
Italy, Piedmont, Rolasco (AL), 25.II. 2014
Copyright © Gianni Allegro

Harpalus (Harpalophonus) stevenii Dejean, 1829a: 242
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) affinis Schrank, 1781: 212 (Carabus)
Larva
Larva of 3rd instar
L = 17,4 mm
Moscow Reg., Istra Distr., env. Novorakovo 3.V.2008 leg. K.Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) affinis (Schrank, 1781)
Male adult
length: 9-12 mm
Helvetia, Graubünden, Ilanz
leg.det. U.Schmidt, 2004
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Harpalus (Harpalus) rubripes (Dufischmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) rubripes (Dufischmid, 1812)
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) amputatus obtusus (Gebler, 1833)
Male adult
Russia, Buryatia, Dzhida distr., Belozersk vic., 4-9.7.1997, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) michailovi Kataev, 1987 Paratype
Unsexed
Image courtesy of Biological Museum, Lund University
Copyright © - - -

Harpalus (Harpalus) semipunctatus Dejean, 1829
Male adult
Spain, Guadalajara, pto de Maranchon, 28.4.2006, 1300m, Anichtchenko A. leg. & det. (cAA) 11mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) semipunctatus Dejean, 1829
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) erosus Mannerheim, 1825
Male adult
E Siberia, lake Baikal, Listvyanka vill. 2.8.1992, Anichtchenko A. leg & det. (cAA) 11.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) tjanschanicus Semenov, 1889
Male+Female adult
Kazakhstan Bayankol vallée près de Narynkol du 27-05 au 01-06-1993
Copyright © Pascal Deschamps

Harpalus (Harpalus) kabakovi Kataev, 1987
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Hemipangus) badakschanus Jedlicka, 1956
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) metallinus Menetries, 1836: 149
Female adult
det. B. Kataev, 2013
Copyright © Kirill Makarov

Harpalus (Heteroharpalus) metallinus Menetries, 1836
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) tarsalis (as hokkaidensis) Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) tarsalis Mannenheim, 1825: 28 Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) tarsalis Mannenheim, 1825: 28 Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) tarsalis (as hokkaidensis Habu et Baba, 1963)
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39250
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Female adult
Russia
Copyright © Kirill Makarov

Harpalus (Harpalus) tardus (Panzer, 1796)
Male+Female adult
France, forêt de Fontainebleau (77300), 18/4/1987, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) modestus Dejean, 1829
Female adult
Russia, 13 km from Irkutsk by Aleksandrovskii road, 20.8.1992, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) albanicus Reitter, 1900
Male adult
Spain, Granada, Castell del Ferro, 5.4.2002 Anichtchenko A. leg. (cAA) 8.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
Italy, Piedmont, Sambuco (CN), m 1400, 4.VI.2006
Copyright © Gianni Allegro

Harpalus (Homaloharpalus) tangutorum Kataev, 1993
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) laevipes (as ainus) Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Hyloharpalus) tibeticus Andrewes, 1930
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) lumbaris Mannerheim, 1825: 27
Male adult

Copyright © Kirill Makarov

Harpalus (Hypsinephus) salinus Dejean, 1829
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Hypsoharpalus) arnoldii Kataev, 1988
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Idioharpalus) numidicus Bedel, 1893
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) serripes (Quensel, 1806)
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) flavicornis Dejean, 1829a: 366
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) politus Dejean, 1829a: 370
Male adult
Germany, Bavaria, Reiterswiesen, F. Boetzl leg. 21.05.14, M. Fritze det. 2014
Copyright © Fabian Bötzl

Harpalus (Harpalus) pseudoserripes Reitter, 1900
Female adult
Iran, Khorasan, Torbat-E-Heydariyeh, pass 5 km N Zharf 23.5.2008, Anichtchenko A. leg (cAA) 10 mm; det. Kataev B.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) flavicornis Dejean, 1829
Male adult
Italy, Piedmont, Ottiglio (Cave di Moleto), 24.V.2009
Copyright © Gianni Allegro

Harpalus (Baeticoharpalus) lopezi Serrano & Lencina, 2008
Male adult
Spain, Jaen prov., Sierra Magina, 2000m, 37.728701, -3.460772 3-4.05.2014, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Licinoderus) franzi Mateu, 1954
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Loboharpalus) rubefactus Bates, 1873
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Harpalus (Loboharpalus) platynotus Bates, 1873
Male adult
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Loboharpalus) platynotus Bates, 1873
Endophallus
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Loxophonus) agakhaniantzi (Mikhailov, 1972)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) laticeps Leconte, 1850
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Harpalus) providens Casey, 1914
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Harpalus (Harpalus) major (Motschulsky, 1850)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Macroharpalus) animosus Casey, 1924
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Megaharpalus) stoetznerianus Schauberger, 1932
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Pseudoophonus) caliginosus (Fabricius, 1775)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Megapangus) katiae F.Battoni, 1985
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Meroharpalus) fulvilabris Mannerheim, 1853
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) mitridati Pliginskiy, 1915
Male adult
Rostov Area, Kumyska Vill. 24.III.2006 leg. M.Nabozhenko
L=8.5
Copyright © Kirill Makarov

Harpalus (Mesoharpalus) kiritshenkoi Kataev, 1990
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Male adult
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Endophallus
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) smaragdinus (Duftschmid, 1812)
Male+Female adult
France, forêt de Saint Germain en Laye 78100, 30/8/85, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) cyclogonus cyclogonus (as sarmaticus)
Male adult
Russia, Volgograd reg., Palasovsk distr., N shore of Elton lake, 19-29.4.2007 Makarov K. leg & det. (cAA) 9 mm
Copyright © Alexander Anichtchenko

Harpalus (Nephoharpalus) pallidipennis A. Morawitz, 1862
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) anxius Duftschmid, 1812: 101 (Carabus)
Larva
Larva of 3rd instar
L = 16,2 mm
Imago from Volgograd Reg., env. of Elton Lake IV.2007 leg. K.Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) egorovi Lafer, 1989
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) amplicollis Menetries, 1848
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) calathoides Motschulsky, 1844: 200
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) anxius (Duftschmid, 1812)
Male+Female adult
France, Forêt de Fontainebleau 77300 18/6/88, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) servus (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) anxius (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) subcylindricus Dejean, 1829a: 374
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) amariformis Motschulsky, 1844
Female adult
Buryatia, Dzhida distr., Belozersk vic., 4-9.7.97, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) quasianxius Kataev, 1989 Paratype
Male adult
Image courtesy of Biological Museum, Lund University
Copyright © - - -

Harpalus (Harpalus) angustitarsis Reitter, 1887
Male adult
Spain, Segovia, Valsain, 3.5.2003 (cAA) det. Anichtchenko A. 7.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Opadius) cordifer Notman, 1919
Female adult
USA, Washington, Thurston Co., 19.8.2005 (cAA) 8.5mm
Copyright © Alexander Anichtchenko

Harpalus (Opadius) cordatus (Leconte, 1853)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Oreoharpalus) famelicus Tschitscherine, 1898
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) crates Bates, 1883: 239
Male adult
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883: 239
Endophallus
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883
Male adult
SE Maritime Prov., Lazovsky dist., Lazo env., 17-20.VI.2010, leg. М. Smirnov Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Harpalus (Harpalus) oblitus Dejean, 1829a: 273
Male+Female adult
France, Brétignolles sur mer 85470 17/5/85, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) oblitus Dejean, 1829a: 273
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) smyrnensis raddei Tschitscherine, 1897
Male+Female adult
Arménie Aragatsotn route du lac Kari 30-06-2014 Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) smyrnensis medicus Kataev, 1993
Female adult
Iran, East Azerbaijan prov., near Kandovan vill. 37.748092°N 46.286895°E 21-22.V.2017 leg.D.Kasatkin L=10.0
Copyright © Kirill Makarov

Harpalus (Harpalus) smyrnensis medicus Kataev, 1993
Male adult
Iran, East Azerbaijan prov., near Kandovan vill. 37.748092°N 46.286895°E 21-22.V.2017 leg.D.Kasatkin L=9.5 mm
Copyright © Kirill Makarov

Harpalus (Pharalus) indianus Csiki, 1932
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) optabilis Dejean, 1829
Male adult
Burjatia, Selenga riv., Deben vic., 15.6.2003 Anichtchenko A. leg & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Male adult
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Platyharpalus) ventralis Leconte, 1848
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Plectralidus) eraticus Say, 1823
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Plectralidus) retractus Leconte, 1863
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalus) distinguendus (Duftschmid, 1812)
Female adult
POLAND; CE 20 (Lower Vistula Valey)
Bruki Unislawskie
25.04.2009
leg. M. Traczyk
Photo of Mieczyslaw Stachowiak
Copyright © Mieczyslaw Stachowiak

Harpalus (Harpalus) distinguendus (Duftschmid, 1812)
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Harpalus (Harpalus) saxicola Dejean, 1829
Male adult
SE Slovakia, Slovenský kras; 7mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) angulatus scytha Tschitscherine, 1899
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) distinguendus distinguendus
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) contemptus Dejean, 1829
Male adult
Spain, Sierra Nevada, 5 km N Capileira, 1800-2000m, 25.4.2003, Anichtchenko A. leg. & det. (cAA) 10 mm
Copyright © Alexander Anichtchenko

Harpalus (Psammoharpalus) kozlovi Kataev, 1993
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus rufipes (Degeer, 1774)
Male adult
Russia, S.Ural
Copyright © Sunvas

Harpalus (Pseudoophonus) rufipes (Degeer, 1774)
Male adult
W-UKRAINE,
Lviv Region, Javorov distr., near Ivano-Frankovo 1-20.VI.07 Panin R. leg
long = 17mm
Copyright © Ruslan Panin

Harpalus (Pseudoophonus) pensylvanicus (Degeer, 1774)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Harpalus (Pseudoophonus) azumai Habu, 1968 Holotype
Male adult
Naha, Okinawa Is., OKINAWA. COL-129
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) azumai Habu, 1968 Holotype
Male adult
Naha, Okinawa Is., OKINAWA. COL-129
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) azumai Habu, 1968 Holotype
Male adult
Naha, Okinawa Is., OKINAWA. COL-129
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) erythropus Dejean, 1829
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Harpalus (Pseudoophonus) longicollis Leconte, 1848
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Harpalus (Pseudoophonus) pseudophonoides Schauberger, 1930
Male adult
Collection ground Tanagura cho, Fukushima pref. 37900
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) pseudophonoides Schauberger, 1930
Endophallus
Collection ground Tanagura cho, Fukushima pref. 37900
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Male adult
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) pastor niigatanus Schauberger, 1929
Male adult
Collection ground Himeji city, Hyogo Pref. 32778
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) pastor niigatanus Schauberger, 1929
Endophallus
Collection ground Himeji city, Hyogo Pref. 32778
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) ussuriensis vicarius Harold, 1878
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) ussuriensis vicarius Harold, 1878
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Male adult
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Endophallus
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Male adult
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) sinicus Hope, 1845
Male adult
Collection ground Hokkaido Pref. Kikonai～cho 34925
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) sinicus Hope, 1845
Endophallus
Collection ground Hokkaido Pref. Kikonai～cho 34925
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Male adult
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Endophallus
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) ussuriensis ussuriensis s.str.
Female adult

Copyright © Kirill Makarov

Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Female adult

Copyright © Kirill Makarov

Harpalus (Pseudoophonus) pastor pastor (tschiliensis Schauberger, 1829)
Male adult

Copyright © Kirill Makarov

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Kirill Makarov

Harpalus (Semiophonus) signaticornis (Duftschmid, 1812)
Male adult
France, 77860, Quincy-Voisins, 12-05-2018, leg. & det. P. Deschamps
Copyright © Pascal Deschamps

Harpalus (Sinoharpalus) hiekei Kataev & Wrase, 2010
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Smirnovia) kandaharensis Jedlicka, 1956
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Zangoharpalus) tinctulus luteicornoides (as ihai Habu) Holotype
Male adult
Hentona, Okinawa Is., OKINAWA, 28. IV. 1958, K. Iha. COL-136
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Zangoharpalus) tinctulus luteicornoides Breit, 1913 Holotype
Male adult
Hentona, Okinawa Is., OKINAWA, 28. IV. 1958, K. Iha. COL-136
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Zangoharpalus) tinctulus luteicornoides Breit, 1913 Holotype
Male adult
Hentona, Okinawa Is., OKINAWA, 28. IV. 1958, K. Iha. COL-136
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Zangoharpalus) tinctulus luteicornoides Breit, 1913 Holotype
Male adult
Hentona, Okinawa Is., OKINAWA, 28. IV. 1958, K. Iha. COL-136
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Zangoharpalus) tinctulus luteicornoides (as tsushimanus) Holotype
Male adult
Uchiyama, Tsushima Is., NAGASAKI, 26. V. 1957, K. Baba. COL-135
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Zangoharpalus) tinctulus luteicornoides (as tsushimanus) Holotype
Male adult
Uchiyama, Tsushima Is., NAGASAKI, 26. V. 1957, K. Baba. COL-135
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Zangoharpalus) tinctulus luteicornoides (as tsushimanus) Holotype
Male adult
Uchiyama, Tsushima Is., NAGASAKI, 26. V. 1957, K. Baba. COL-135
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Male adult
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Endophallus
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) praticola Bates, 1891
Male adult
Image from Kataev, 2023
Copyright © - - -