Bembidion (Ocydromus) atlanticum atlanticum Wollaston, 1854
Male adult
Uzbekistan, Chatkal mt. rng., 6 km NW Irtash, Iertash riv., 41.10.46N 70.17.28E, 1900m, 3.8.2021, Anichtchenko A. leg. (cAA). det. L. Toledano, 2026
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) basistriatum Fairmaire, 1893
Male adult
Ethiopia, Oromia, Mt Enkuolo, NE-slope, 3100-3200m, 07°24’41”N 39°22’26”E, 5-7.Xii.16, leg. J. Schmidt (CSChM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) corgenoma Maddison, 2020 Paratype
Male adult
USA: Oregon: Benton Co., Corvallis, Willamette River, 44.5491°N, 123.2449°W, 60 m
Copyright © David Maddison

Bembidion (Ocydromus) decorum caraffai Sainte-Claire Deville, 1906a: 15
Female adult
France Corse
Copyright © Pascal Deschamps

Bembidion (Ocydromus) decorum subconvexum K. Daniel & J. Daniel, 1902: 102
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) decorum decorum (Panzer, 1799b)
Male adult
CZECH REPUBLIC - Staré Hamry, 22.IX.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033. (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html). Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu. Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Label
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,7
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,7
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,8
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Urup Isl., Ukromnaya Bay, sea shore rocky debris, 20.viii.1996, leg. K.Eskov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5,2
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5.0
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,75 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Spermatheca
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,6 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,85 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,45
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=5.0 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=4,7 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,55 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Spermatheca
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,75
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Spermatheca
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,9 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Spermatheca
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,8
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,3 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,6 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=5 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov
Copyright © Kirill Makarov

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 (Label) Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) erosum (Motschulsky, 1850)
Male adult
USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051°N, 124.1005°W (V101453)
Copyright © David Maddison

Bembidion (Ocydromus) fontinale (Raffray, 1885)
Male adult
Ethiopia Amhara, Mt. Abuna yosef, n-slope, 3800-3950 m, 12°07’52”N 39°11’39”E, 4.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) fontinale (Raffray, 1885)
Male genitalia

Copyright © Joachim Schmidt

Bembidion (Peryphanes) grapii Gyllenhal, 1827
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov <br /> Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Ocydromus) hasurada Andrewes, 1924a: 77
Male adult
India, Uttarakhand, 10-12.iV.2012, 14km E Uttarkashi, Anichtchenko A. leg.; 5 mm. det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) hiekei Muller-Motzfeld, 1986b: 171 Paratype
Male adult

Copyright © Luca Toledano

Bembidion (Ocydromus) loebli Schmidt, 2014 Holotype
Male adult
Holotype male, with label data „INDIA Madras / Nilgiri 2100 m / Ootacamund - Pykara / 21.XI.72 / Besuchet Löbl Mussard“ in MHNG
Copyright © Qin Yuyao

Bembidion (Ocydromus) loebli Schmidt, 2014 Paratype
Male genitalia

Copyright © Qin Yuyao

Bembidion (Ocydromus) melanoceroides Toledano, Bonavita & Schmidt, 2021 Paratype
Male adult
Ethiopia, Oromia, Mt Enkuolo, N-slope, 3350-3600m, 07°22’48”N 39°21’07”E, 6-9.Xii.16, leg. J. Schmidt (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocydromus) melanocerum Chaudoir, 1876
Male adult
Ethiopia, Amhara, W-slope Mt. Choke, 2750-2900 m, 10°38’N 37°45’E, 1.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) modestum (Fabricius, 1801)
Male adult
CZECH REPUBLIC - Olomouc dist., Skalička, 11.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Colorado: Huerfano Co., Huerfano River at Badito, 37.7285°N 105.0167°W.
Scale bar is 1 mm.
Photo David R. Maddison & Alexander P. Swanson
Copyright © ---

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Arizona: Cochise Co., San Pedro R at Charleston, 31.6239°N, 110.1722°W (voucher V101461)
Copyright © David Maddison

Bembidium haplogonum Chaudoir, 1868 (NEOTYPE)
Male adult
USA: California: Lake Co., North Branch Cache Creek at hwy 20, 305 m 38.9881°N, 122.5400°W
Copyright © David Maddison

Bembidion (Ocydromus) primorjense Neri & Toledano, 2023 Holotype
Male adult
Holotype, ♂, “Russia N Primorskyi reg. / 70 km NW Plastun 18-26 VII / Podnebesnyi pass 600-800 m / A. Plutenko leg. 1996” [printed]” (CTVR)
Copyright © Qin Yuyao

Bembidion (Ocydromus) primorjense Neri & Toledano, 2023 Holotype Paratype
Male genitalia

Copyright © Qin Yuyao

Bembidion (Ocydromus) salweenum Schmidt, 2014 Holotype
Male adult
Holotype male, with label data “STIBET 26-29.IX.2010 / Kema SSE Naqu 4450 m / Salween River, banks, alpine / steppe, 31°16’34N 92°06’20E” in ZSM
Copyright © Qin Yuyao

Bembidion (Ocydromus) salweenum Schmidt, 2014 Paratype
Male genitalia

Copyright © Qin Yuyao

Bembidion (Ocydromus) sarpedon Casey, 1918
Male adult
USA: Colorado: Las Animas Co., Cokedale, Reilly Canyon, 37.1346°N, 104.6114°W (V101459)
Copyright © David Maddison

Bembidion (Ocydromus) saxatile Gyllenhal, 1827: 406
Female adult

Copyright © Kirill Makarov

Bembidion (Ocydromus) saxatile morsum Netolitzky, 1930
Male adult
Tajikistan, Khoja Obi Garm, 15.9.2011, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) saxatile himalotibeticum Schmidt, 2014 Holotype
Male adult
Holotype male, with label data “S TIBET 15.VII.2010 / YamTso lake shore near / Nagartse, near Samding / Monastery, 4420 m” in ZSM.
Copyright © Qin Yuyao

Bembidion (Ocydromus) saxatile himalotibeticum Schmidt, 2014 Paratype
Male genitalia

Copyright © Qin Yuyao

Bembidion (Ocydromus) saxatile flavipalpe Netolitzky, 1930
Male adult
Uzbekistan, Chatkal Mt. rng., Yangiobod, left conf of Dukentsay riv, 41.07.28N 70.06.21E 1300m 2.8.2021 Anichtchenko A. leg. (cAA). det. L. Toledano, 2026
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Ocydromus) sokolowskii Fassati, 1957b: 154 Holotype
Male adult
MHNG
Copyright © Luca Toledano

Bembidion (Ocydromus) terryerwini Neri & Toledano, 2021 Paratype
Male adult
N 29°21'14.7" E 57°20'31.7" / Iran, Prov. Kerman, Rāyen, Goroah, / Babtahoune, Bachufer-Schotterbank / 06.06.2014 2700 m / leg. Schnitter
Copyright © Luca Toledano

Bembidion (Ocydromus) transsylvanicum Bielz, 1852
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Marmarosh Mts., near Lug vill., bank of Kuziy (Kizi) stream, 400m, 26.IV.2014, Panin R. & Kanarsky Yu. leg. L= 5 mm
Copyright © Ruslan Panin

Bembidion (Peryphus) transversale Dejean, 1831
Male adult
Images from David R. Maddison & Alexander P. Swanson, 2010
Copyright © - - -

Bembidion (Ocydromus) transversale Dejean, 1831
Male adult
Canada: Alberta: Burbank, junction of Red Deer and Blindman Rivers, 52.3542°N, 113.7556°W (voucher V101454)
Copyright © David Maddison

Bembidion (Peryphanes) yukonum Fall, 1926: 131
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) zolotarewi Reitter, 1910b: 313
Male adult
Russia, NW Caucas, Krasnodar reg., Tuapse distr., Podchrebetovoe vicinity
(NW Кавказ, Краснодарский край, Туапсинский р-н, окр пос. Подхребeтовое 19.V.2012 leg. М. Смирнов)
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov