Bembidion (Actedium) pallidipenne (Illiger, 1802b)
Male adult
length: 4,1-4,7 mm, Niederlande, Maasvlakte, leg. A.Teunisse, 1989; det. G.Rößler, 1996, Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Andrewesa) patris Schmidt, 2010 Paratype
Male adult
NEPAL Annapurna Mts. / Madi Khola below Sikles / 1400 m, water power st. [station, 28°22N 84°08E] / 10.5.1996 leg. J. Schmidt. Image from Schmidt, 2010
Copyright © - - -

Bembidion (Antiperyphanes) mandibulare (Solier, 1849)
Male adult
Chile: Reg. X, Chiloé: Cucao, DRM voucher V100789. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Antiperyphanes) hirtipes (Jeannel, 1962)
Male adult
Argentina, Mendoza, Pampa Palauco. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Antiperyphanes) rufoplagiatum Germain, 1906
Male adult
Argentina, Neuquen, Arroyo Queni at Lago Queni. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Antiperyphanes) zanettii Toledano, 2008
Male adult
Ecuador: Napo: Rio Quijos W of Baeza. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Antiperyphanes) spinolai Solier, 1849
Male adult
Argentina: Chubut: Rio Azul at Lago Puelo. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Antiperyphus) philippii Germain, 1906
Male adult
Argentina: Neuquén: Rio Collón Curá ca 13 km S La Rinconada, DRM voucher V100787. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Apteromimus) platyderoides (Wollaston, 1877)
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Apteromimus) shepherdae Maddison, 2019
Female adult
St Helena
Copyright © David Maddison

Bembidion (Archaeophilochthus) christelae Ortuno & Arillo, 2010 Holotype
Male adult
Baltic Eocene amber
Copyright © Alexander Anichtchenko

Bembidion (Armatocillenus) sauteri (Jedlicka, 1958) Holotype
Female adult
Formosa, Sauter // Anping, 1909.X. // Typus [red] // Armatocillenus sauteri n.sp. [handwritten], det. ING. JEDLIČKA [pink] (HNHM).
Copyright © - - -

Bembidion (INCERTAE) bandotaro Morita, 1991
Female adult
Kuril Islands
photo Makarov K.
Copyright © Kirill Makarov

Bembidion (Asioperyphus) infuscatum Dejean, 1831
Female adult
length: 6,4 mm
West Siberia, KHMAO-Yugra, Khanty-Mansiysk distr., Ob river, Bolshoy Tchukhtinsiy island
4.08.2007.
Zinoviev E.leg.
Copyright © Evgeniy Zinoviev

Bembidion (Asioperyphus) postremum Say, 1830
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Asioperyphus) altestriatum Netolitzky, 1934a: 68
Male adult
Russia, SE Primorie, Lazo Nat. res., 10 km SW Sokol'chi vill.
(Приморский кр., Лазовский зап., 10 км SW пос. Сокольчи, окр. кордона Корпадь 43°15' N, 134°7' E, 23-27.06.10, leg. М.Смирнов)
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Asioperyphus) semilunium semilunium s.str.
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Asioperyphus) pamiricola pamiricola Lutshnik, 1930: 34
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Asioperyphus) bandotaro Morita, 1991a: 119
Female adult
Kuriles isl. det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Asioperyphus) xanthochiton Andrewes, 1922b: 176
Female adult
India, Uttarakhand, 14 km NW New Tehri, 950m, N30*28'25.3" E 078*20'56.2", 10.4.2012, A. Anichtchenko leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) ovale Motschulsky, 1844: 239
Male adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) lunatum Duftschmid, 1812: 211
Male adult
Latvia, Daugavpils distr., Silene, Ilgas, 18.5.1991, A. Barsevskis leg.
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) sajanum Shilenkov in Kryzhanovskij & al., 1995
Male adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) kazakhstanicum Kryzhanovskij, 1979 Paratype
Unsexed
Image courtesy of Biological Museum, Lund University
Copyright © - - -

Bembidion (Asioperyphus) notatum Andrewes, 1922b: 175
Female adult
India, Uttarakhand, 8 km NW Jaspur, lake shore, 22-23.4.2012, Anichtchenko A. leg. (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Australoemphanes) ateradustum Liebherr, 2008
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Bembidion) quadrimaculatum (Linne, 1761)
Male adult
length: 2,5-3,6 mm
Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1972
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Bembidion (Bembidion) quadrimaculatum oppositum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Bembidion) quadrimaculatum mandli Netolitzky, 1932
Female adult

Copyright © Kirill Makarov

Bembidion (Bembidion) humerale humerale Sturm, 1825: 176
Female adult
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha” NW part, 17.IV-10.VII.2018, Panin R. leg., L =2.95 mm
Copyright © Denis Khrapov

Bembidion (Bembidionetolitzkya) conforme Dejean, 1831
Male adult
length: 4,5-6 mm
Germania, Oberbayern, Oberstdorf
leg.det. U.Schmidt, 2005
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837
Female adult
Ukrainian Carpathians,
Ivano-Frankivsk reg.,Yaremcha distr.,Gorgany Mts. ,h=750 m. 22.VI.08 Panin R. leg.
7mm.
Copyright © Ruslan Panin

Bembidion (Bembidionetolitzkya) cyaneum Chaudoir, 1846a: 203
Male adult
N Кавказ, Карачаево-Черкесия, окр. г.Теберда 1.VII.1994 leg. А. Гусаков
N Caucas, Karachaevo-Cherkessia, Teberda vicinity
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)
Male adult
NW Кавказ, 20 км S г. Горячий Ключ, бер. горной речки 3.04.1996 leg. М. Смирнов
NW Caucas, 20 km S from Goriachii Kluch
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Bembidionetolitzkya) cyaneum Chaudoir, 1846
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) varicolor agurense Lutshnik, 1938
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) longipes K. Daniel, 1902a: 29
Male adult
France, 05380 Chateauroux les Alpes Le Rabioux, 02-VII-2014
Copyright © Pascal Deschamps

Bembidion (Bembidionetolitzkya) cimmerium Andrewes, 1922c: 238
Male adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A. leg. 5.5 mm. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) penninum Netolitzky, 1918a: 20
Male adult
Val Cairasca Sempione VI-1915 Mancini
Copyright © Jan Muilwijk

Bembidion (Bembidionetolitzkya) chaklaense Schmidt, 2018 Paratype
Male adult
“TIBET South Centr. 16.VII.07 / 50 km NNE Lhasa, SW of / Chak La pass 4100-4550m / Chakla river valley, lg. Schmidt”, “30°03’36.6N 91°16’51.7E / to / 30°06’10.1N 91°16’38.4E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) chaklaense Schmidt, 2018 Paratype
Male genitalia
“TIBET South Centr. 16.VII.07 / 50 km NNE Lhasa, SW of / Chak La pass 4100-4550m / Chakla river valley, lg. Schmidt”, “30°03’36.6N 91°16’51.7E / to / 30°06’10.1N 91°16’38.4E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) cymindulum Andrewes, 1930
Male genitalia
Tibet, Gyatso La S Lhatse, 5200-5300 m, 28°57’10”N 87°26’15”E, 21.VI.2011 leg. J. Schmidt
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lhai Schmidt, 2018 Paratype
Male genitalia
“TIBET South Centr. 17-20.VI.07 / Budha Vall. N of Yangpachem / ca. 30°10’56N 90°29’21E / 5000-5200 m, leg. J. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lhai Schmidt, 2018 Paratype
Male adult
“TIBET South Centr. 17-20.VI.07 / Budha Vall. N of Yangpachem / ca. 30°10’56N 90°29’21E / 5000-5200m, leg. J. Schmidt” (CSCHM).
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) pinnigerum Schmidt, 2018 Holotype
Male genitalia
“TIBET South Centr. 17/18.VII.07 / Reting Tsangpo Vall. E Reting / 4230-4800 m, Kiykiy place / ca. 30°24’15N 91°42’59E lg.J.S.” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) pinnigerum Schmidt, 2018 Holotype
Male adult
“TIBET South Centr. 17/18.VII.07 / Reting Tsangpo Vall. E Reting / 4230-4800 m, Kiykiy place / ca. 30°24’15N 91°42’59E lg.J.S.” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lunepabomaguvi lunepabomaguvi Paratype
Male genitalia
“INDIA Arunachal Pradesh / 10-15 km W Sela Pass / 34-3500 m, 24.VI.2008 / 27°30N 92°03E lg. Reuter” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lunepabomaguvi lunepabomaguvi Paratype
Male adult
“INDIA Arunachal Pradesh / 10-15 km W Sela Pass / 34-3500 m, 24.VI.2008 / 27°30N 92°03E lg. Reuter” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lunepabomaguvi bhutanicum Schmidt, 2018 Paratype
Male genitalia
“BT Wangdue Phodrang distr. / Phobjikha Vall., 2800 m / 28.IX.2017, leg. J. Schmidt / 27°24’40”N 90°12’12”E” (NBCB).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lunepabomaguvi bhutanicum Schmidt, 2018 Paratype
Male adult
“BT Wangdue Phodrang distr. / Phobjikha Vall., 2800 m / 28.IX.2017, leg. J. Schmidt / 27°24’40”N 90°12’12”E” (NBCB).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) namtso Schmidt, 2018 Paratype
Male genitalia
“TIBET (South Central) 12.VII.07 / S Namtso, Langma Valley / 5100-5150 m, leg. Schmidt / 30°37’39,1”N90°51’56,5”E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) namtso Schmidt, 2018 Paratype
Male adult
“TIBET (South Central) 12.VII.07 / S Namtso, Langma Valley / 5100-5150 m, leg. Schmidt / 30°37’39,1”N90°51’56,5”E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) retingense Schmidt, 2018 Paratype
Male genitalia
“CHINA Tibet 1.VII.2015 / side valley S Reting / 4400 m leg. J. Schmidt / 30°16’44”N 91°31’55”E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) retingense Schmidt, 2018 Paratype
Male adult
“CHINA Tibet 1.VII.2015 / side valley S Reting / 4400 m leg. J. Schmidt / 30°16’44”N 91°31’55”E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) smetanai smetanai Paratype
Male genitalia

Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) smetanai qingmaiense Schmidt, 2018 Paratype
Male genitalia
“China, S.W. Sichuan / Qingmai 3000 m / N 28°48’46.7”, E 099°52’09.2” / 3.7.2006, lgt. Janata, M.” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) smetanai qingmaiense Schmidt, 2018 Paratype
Male adult
“China, S.W. Sichuan / Qingmai 3000 m / N 28°48’46.7”, E 099°52’09.2” / 3.7.2006, lgt. Janata, M.” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) transhimalayanum Schmidt, 2018 Paratype
Female adult
“S TIBET, ca. 40km SW of / Yangpachem, 13.VII.2010 / Pijin moraine 5000-5500 m / 29°45’36N 90°17’54E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) wrzecionkoi Toledano & Schmidt, 2008
Male genitalia
specimen from the type locality.
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) wrzecionkoi Toledano & Schmidt, 2008
Male adult
south central Tibet, NE of Gyantse, Yung La pass and above, 4850-5400 m, 28°57’14”N 89°50’04”E, 17.VII.2010
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) yamtso Schmidt, 2018 Holotype
Male genitalia
“S TIBET 14/15.VI.2011 / E-slope Karo La pass / 4900-5300 m, leg. Schmidt / 28°55’N 90°18’E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) yamtso Schmidt, 2018 Holotype
Male adult
“S TIBET 14/15.VI.2011 / E-slope Karo La pass / 4900-5300 m, leg. Schmidt / 28°55’N 90°18’E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) zugmayeri Schmidt, 2018 Holotype
Male genitalia
“Sumdschiling / Ebene [plateau] W Tibet”, “Süssw. Tümp. [fresh water pond] / VII/06. Zugm.” (ex coll. W. Marggi, Thun, in CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) zugmayeri Schmidt, 2018 Holotype
Male adult
“Sumdschiling / Ebene [plateau] W Tibet”, “Süssw. Tümp. [fresh water pond] / VII/06. Zugm.” (ex coll. W. Marggi, Thun, in CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) bactrianum K. Daniel, 1902
Male adult
Tadzhikistan, Varzob, Khodja obi Garm, 2000m 21-24.3.2015 (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)
Female adult

Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)
Male adult

Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) conforme Dejean, 1831
Male adult
W Ukraine, NE Carpathians, Tiachiv distr., h~400 m, bank of stream Mala Ugolka, 23.VI.2016, Panin R. leg.
Copyright © Ruslan Panin

Bembidion (Bembidionetolitzkya) varicolor varicolor
Male adult
CZECH REPUBLIC - Bystřice nad Olší, 15.III.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Blepharoplataphus) hastii C.R. Sahlberg, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Blepharoplataphus) hastii C.R. Sahlberg, 1827
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Chilioperyphus) orregoi Germain, 1906
Male adult
Argentina: Chubut: Rio Azul at Lago Puelo. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Chlorodium) splendidum Sturm, 1825: 145
Female adult

Copyright © Kirill Makarov

Bembidion (Chlorodium) pygmaeum ab. bilunulatum Bielz, 1852
Male adult
N Moravia, Týn nad Bečvou; 2,5mm. det. J. Coulon, 2015
Copyright © Aleš Sedláček

Bembidion (Chlorodium) pygmaeum (Fabricius, 1792)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Chlorodium) splendidum luteipes (Motschulsky, 1844)
Male adult

Copyright © Kirill Makarov

Bembidion (Chlorodium) leucolenum Bates, 1873a: 275
Female adult
Kunashir Island, lower reaches of Krivonozhka Stream, 20-22.VI.2013, Yu. & L. Sundukov leg.
Copyright © Kirill Makarov

Bembidion (Chlorodium) leucolenum Bates, 1873a: 275
Male adult
Kunashir Island, watershed of Golovnina and Khlebnikova rivers west of Dubovoe, 28.IX.2015, Yu. Sundukov
Copyright © Kirill Makarov

Bembidion (Chlorodium) difforme (Motschulsky, 1844)
Female adult
Russia, Burjatia, Muhor-Shibirsk distr., 5km N Nikolsk vill., salt marsh, 27.8.1998, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Cillenus (Cillenus) lateralis Samouelle, 1819
Male adult
Netherlands, Bruinisse, leg. Krift, 1976; det. U.Schmidt, 1982
Photo: U.Schmidt, 2008
Copyright © Udo Schmidt

Bembidion (Cillenus) palosverdes Kavanaugh and Erwin, 1992
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Cillenus) lateralis Samouelle, 1819
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Cillenus (Desarmatocillenus) sumaoi (Morita, 1981)
Male adult
Chiba, Honshu, Japan<br /> (c) Field Museum of Natural History <br /> (Maruyama Munetoshi)
Copyright © Munetoshi Maruyama

Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Cillenus (Desarmatocillenus) kasaharai Habu, 1978 (Label) Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Cillenus (Desarmatocillenus) yokohamae (Bates, 1883)
Male adult

Copyright © Kirill Makarov

Bembidion (Desarmatocillenus) aestuarii (Habu & Ueno, 1954)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Desarmatocillenus) kasaharai Habu, 1978
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Desarmatocillenus) yakushimanum Sasakawa, 2007
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Desarmatocillenus) foochowense Lindroth, 1980
Male adult
TAIWAN: Lunglungchi, Fulung, Taipei Hsien, 31.vii.2001, Terada lgt.
Copyright © - - -

Bembidion (Diplocampa) transparens (Gebler, 1829)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Diplocampa) fumigatum (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Diplocampa) transparens (Gebler, 1829)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Diplocampa) transparens prostratum (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=3.6 mm
Copyright © Kirill Makarov

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Male genitalia
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Male genitalia
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.0 mm
Copyright © Kirill Makarov

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Female genitalia
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Spermatheca
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Ecuadion) ricei Maddison & Toledano, 2012 Paratype
Male adult
ECUADOR: Napo: Rio Chalpi Grande, 2800m, 0.3645°S, 78.0852°W, D.R. Maddison voucher V100622
Scale bar is 1 mm
image from (Maddison & Toledano, 2012)
Copyright © ---

Bembidion (Ecuadion) georgeballi Toledano, 2008
Male adult
ECUADOR: Pichincha: Quebrada Lozada, on road to Res. Yanacocha, 3460m, 0.1105°S, 78.5642°W, voucher V100658<br /> Scale bar is 1 mm.<br /> Image from Maddison & Toledano, 2012
Copyright © - - -

Bembidion (Ecuadion) andersoni Toledano, 2008
Male adult
ECUADOR: Pichincha: Reserva Yanacocha, start Andean Snipe Trail, DRM voucher V100655. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Ecuadion) chimborazonum (Bates, 1891)
Male adult
ECUADOR: Pichincha: Paso de la Virgen, DRM voucher V100793. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) sanctaemarthae Darlington, 1934
Male adult
ECUADOR: Napo: Rio Chalpi Grande. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) ricei Maddison & Toledano, 2012
Male adult
ECUADOR: Napo: Rio Chalpi Grande. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) cotopaxi Moret & Toledano, 2002
Male adult
ECUADOR: Pichincha: km 17 on route 28 W of Papallacta. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) walterrossii Toledano, 2008
Male adult
ECUADOR: Napo: Vinillos, 4.1 km S Cosanga. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) cotopaxi Moret & Toledano, 2002 Paratype
Female adult
Ecuador, Cotopaxi, Laguna Quilotoa, 3500m, 22.7.2001 P. Moret leg.
Copyright © Alexander Anichtchenko

Bembidion (Emphanes) tenellum Erichson, 1837
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Emphanes) latiplaga flavibase De Monte, 1956a: 179
Male adult
Morocco, Souss-Massa Draa, Zagora, OuedDraa riv., 1.2.2013. det. M. Toribio, 2017. 2.5 mm
Copyright © Alexander Anichtchenko

Bembidion (Emphanes) bulgani Jedlicka, 1968
Female adult
Russia: Primorskii krai: 20 km NWW Spassk-Dal'ny, Khanka Lake, Khankaysky Nat. Res., kordon Vostochnyi, 3-8.VI 2008, A. Napolov leg.
Copyright © Kirill Makarov

Bembidion (Emphanes) tenellum tenellum Erichson, 1837
Male adult
C Russia, Ivanovo ar., Pestyaki distr., Demidovo vill. env., Lukh riv. 4-5.VII.2014 leg. M. Smirnov ; Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Emphanes) latiplaga latiplaga Chaudoir, 1850
Female adult
NW-Ukraine, Volhynia, Turiysk env., 17-26.VI.2016, to light, Voytko P. leg./ Panin R. det., L=3.1 mm
Copyright © Denis Khrapov

Bembidion (Endosomatium) megalops (Wollaston, 1877)
Female adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Euperyphus) combustum combustum Menetries, 1832: 138
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=7.0 mm
Copyright © Kirill Makarov

Bembidion (Euperyphus) combustum combustum Menetries, 1832: 138
Female genitalia
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Euperyphus) combustum combustum Menetries, 1832: 138
Spermatheca

Copyright © Kirill Makarov

Bembidion (Euperyphus) giganteum reinigi Netolitzky, 1934 Syntypes
Unsexed
Maz, West-Pamir (15.—19. VIII. 1928), 3580 m
Copyright © Jan.Muilwijk

Bembidion (Euperyphus) giganteum giganteum J.R. Sahlberg, 1900
Female adult
S Kazakhstan, N slope Talas Mt.rng., Dzhabagly riv., 1100m 28.7.2004, A.Matalin leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) morawitzi Csiki, 1928
Female adult
China, Liaoning, Kuandian co., Banlaing, Ba-che-chuan (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Euperyphus) scapulare lomnickii Netolitzky, 1916: 260
Female adult
W Ukraine, NE Carpathians, Lviv reg., near Rozgirche vill., bank of Stryi river, 11.V.2014, Khrapov D. leg./det.Panin R., L= 5.4 mm
Copyright © Denis Khrapov

Bembidion (Euperyphus) scapulare lomnickii Netolitzky, 1916: 260
Male adult
W-Ukraine, Transcarpathian region, Vynogradiv distr., bank of Tisza river near Mala Kopanya vill., 5-7.VI.2018, Panin.R leg., L=5,8 mm
Copyright © Denis Khrapov

Bembidion (Euperyphus) eques Sturm, 1825
Female adult
Italy, Moggio Udinese, Torrente Aupa, 6.9.2023, 13.2007° E, 46.4084° N, J. Gebert leg.
Copyright © Jörg Gebert

Bembidion (Euperyphus) fluviatile fluviatile Dejean, 1831: 113
Male adult
CZECH REPUBLIC - Polanka nad Odrou, 6.IV.2017, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Eupetedromus) graciliforme Hayward, 1897
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Eupetedromus) sibiricum Dejean, 1831a: 66
Female adult
S Primorie, Ussurijski Reserve, Mirinov Klyuch 43°38.644' N 132°26.693' E 25.VIII.2007 leg. K.Makarov, A.Zaitzev
L =4.7
Copyright © Kirill Makarov

Bembidion (Eupetedromus) dentellum (Thunberg, 1787)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Eupetedromus) ruthenum Tschitscherine, 1895c: 298
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Eupetedromus) immaturum Lindroth, 1954 Paratype
Male adult
W Newfoundland, Deer Lake, 30.V.51, Lindroth. 4254:2 (MZLU)
Copyright © Lund University

Bembidion (Eurytrachelus) pogonoides Bates, 1883
Female adult
Far East, Lazovsky Nat.Res., shore of Japan Sea, cordon Petrova, Petrovski spring 10.06.1995 leg. Yu.Sundukov
female
L=6,2 mm
Copyright © Maksim Smirnov

Bembidion (Eurytrachelus) nitidum (Kirby, 1837)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Eurytrachelus) laticolle (Dufischmid, 1812)
Female adult
Orenburg Reg., Ilek Distr., Ilek env., valley of Ural Riv., beach 28.VII-8VIII.2005 S.Vastschenko leg.
L=6.2 mm
Copyright © Kirill Makarov

Bembidion (Eurytrachelus) vitiosum Gemminger & Harold, 1868
Female adult
Russia, Irkutsk reg., Nizhneudinsk distr., Uda riv., Bolshoy Uk waterfall, 22.7.1898, A.Anichtchenko leg. (DUBC).
Copyright © Alexander Anichtchenko

Bembidion (Furcacampa) versicolor (Leconte, 1848)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Furcacampa) impotens Casey, 1918
Female adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Furcacampa) affine Say, 1823
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Geocosmius) elescarabajo Maddison, 2023
Male adult

Copyright © David Maddison

Bembidion (Geocosmius) franiae Erwin, 1982
Male adult

Copyright © David Maddison

Bembidion (Gnatholymnaeum) spurcum Blackburn, 1881
Male adult
Hawaii. Image from J.K. Liebherr, 2008
Copyright © - - -

Bembidion (Hirmoplataphus) nigrum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Hirmoplataphus) hirmocaelum Chaudoir, 1850
Female adult
Siberia, Irkutsk, Irkut riv., 3.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Hirmoplataphus) pliculatum Bates, 1883
Male genitalia
S Sakhalin isl., Susuya riv. leg & det. Sundukov, 2023
Copyright © Kirill Makarov

Bembidion (Hirmoplataphus) pliculatum Bates, 1883
Male adult
S Sakhalin isl., Susuya riv. leg & det. Sundukov
Copyright © Kirill Makarov

Bembidion (Hirmoplataphus) hirmocaelum Chaudoir, 1850
Male genitalia
Primorski reg., Kedrovaya riv. det. Sundukov, 2023
Copyright © Kirill Makarov

Bembidion (Hirmoplataphus) hirmocaelum Chaudoir, 1850
Male adult
Primorski reg., Kedrovaya riv. det. Sundukov, 2023
Copyright © Kirill Makarov

Bembidion (Hydrium) laevigatum Say, 1830
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Bembidion (Hydrium) laevigatum Say, 1830
Male adult
USA: Utah: Grand Co., Moab, Colorado River, 1190 m, 38.5739°N, 109.5769°W
Copyright © David Maddison

Bembidion (Hydrium) mimbres Maddison, 2020 Paratype
Male adult
USA: New Mexico: Grant Co., Gila River, Billings Vista, 1320 m, 32.8163°N, 108.6032°W, 11.viii.2005. DRM 05.043. D.R. & J.H.A. Maddison
Copyright © David Maddison

Bembidion (Incertae sedis) equatoriale Vandyke, 1953 Holotype
Unsexed

Copyright © - - -

Bembidion (Jammuphanes) sinuosum Schmidt & Marggi, 2014 Paratype
Male adult
Pakistan, Kaghan valley, Shogran, 3050m, 19.6.2001. Imagr from Schmidt & Marggi, 2014
Copyright © - - -

Bembidion (Josefia) praeceptor Schmidt, 2018 Holotype
Male genitalia
“Tibet (South Central) 26.VI.07 / Gyama vall. ca. 60km E Lhasa / 4230-4620 m leg. J. Schmidt / ca. 29°41’48N 91°38’29E”, “south ascent of / Chungenpo Shong side vall” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Josefia) praeceptor Schmidt, 2018 Paratype
Male adult
“Tibet (South Central) 26.VI.07 / Gyama vall. ca. 60km E Lhasa / 4230-4620 m leg. J. Schmidt / ca. 29°41’48N 91°38’29E”, “south ascent of / Chungenpo Shong side vall” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Leuchydrium) tigrinum Leconte, 1879
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Kunashiri Isl., mouth of Medny Spring 44°16'31"N 145°56'29"E 27.VI.2008 leg. K.Makarov
L=4.3
Copyright © Kirill Makarov

Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Liocosmius) darlingtonielum Cooper & Maddison, 2014
Male adult

Copyright © David Maddison

Bembidion (Liocosmius) cachagua Maddison, 2023 Holotype
Male adult
CAS: “U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln.”, “C.A.S. Accession 29-V-48 Cachagua Creek”, “David R. Maddison DNA3842 DNA Voucher”
Copyright © David Maddison

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Thompson Island, 7.5.2008, Massachusetts
Images J. Rawlins
Copyright © ---

Bembidion (Lymnaeum) laticeps (Leconte, 1858)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Plataphus) altaicum Gebler, 1833
Male adult
Primorie reg., Lazo distr., near Lazo vill. Lazovka riv. 20.06.04 leg. M.& L. Smirnov photo from www.zin.ru
Copyright © Maksim Smirnov

Bembidion (Metallina) properans (Stephens, 1828b): 26 (Tachypus)
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
length: 2,7-4,1 mm
Germania, Bayern, Oberfranken, Neuenmarkt
leg.det. U.Schmidt, 1972
Photo: U.Schmidt, 2009
Copyright © U.Schmidt

Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Metallina) elevatum (Motschulsky, 1844)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Metallina) properans (Stephens, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Neja) nigricorne Gyllenhal, 1827: 402
Female adult
Czech Republic, Bohemia occ., Kraslice, Tisovec hill, 750-755 m a.s.l., 24.iv.1999, lgt. et det. J. Hejkal
Copyright ©

Bembidion (Neobembidion) constricticolle Hayward, 1897
Male adult

Copyright © David Maddison

Bembidion (Neoemphanes) shimoyamai Habu, 1978
Male adult
Far East, khanka Lake env., Bol.Usachi Riv. near Novokachalinsk 5.VII.2004 R.Filimonov leg.
L=3.5 mm
Copyright © Kirill Makarov

Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Neoemphanes) shimoyamai Habu, 1978 (Label) Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nepha) menetriesii (Kolenati, 1845)
Female adult
Russia, Orenburg reg., Sol'-Iletskii distr., 10 km W Troitsk vill. 6.05.09
leg. M. & L. Smirnov's
Female
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov

Bembidion (Nepha) genei illigeri Netolitzky, 1914
Male adult
length: 4,0-4,9 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2001
Foto: U.Schmidtz, 2010
Copyright © U.Schmidt

Bembidion (Nepha) genei illigeri Netolitzky, 1914: 54
Female adult
length: 4,6 mm
South Urals, Chelyabinsk area, Asha distr., the vicinities of Sim town, Sim river valley
14.05.2001.
Rudoiskatel' P.leg.
From the collection of A.Ivanov (Ekaterinburg)
Copyright © Evgeniy Zinoviev

Bembidion (Nepha) tetrasemum Chaudoir, 1846a: 207
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Nepha) caucasicum (Motschulsky, 1844)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Nepha) schmidti subcallosum Wollaston, 1864: 71
Male adult
3,6 mm; Spain, Canary Islands, Tenerife, Teno, Erjos, 1000 m, leg. J.Schoenfeld, 11.VII.2010; det. Lompe, 2012. Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Bembidion (Nepha) hauserianum Netolitzky, 1918
Female adult
Iran, 20 km SW Yasuj, 5-6.V.2007, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.1 mm
Copyright © Kirill Makarov

Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Male genitalia
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.2 mm
Copyright © Kirill Makarov

Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Female genitalia
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Spermatheca
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Nepha) glabrum (Motschulsky, 1850)
Male adult
Новосибирская обл., 9 км З г. Карасук, оз Кротовое. Image from Dudko, Bespalov, Zinovyev & Lyubechanskii, 2018
Copyright © - - -

Bembidion (Nepha) callosum subconnexum De Monte, 1953
Male adult
Spain, Gredos Mts., 2km E from La Mira, 1800m, 4-7.7.2007, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Nepha) schmidti alluaudi (Antoine, 1925)
Male adult
Morocco, Grand Atlas, 2200m Oukaimedene env., 16-17..6.2010, Anichtchenko A. leg. (DUBC). det. M. Toribio
Copyright © Alexander Anichtchenko

Bembidion (Nepha) ibericum Piochard de la Brulerie, 1868
Male adult
Spain, Sierra de Guadarrama, Laguna de los Cleveles, 6.VIII.2024, 2000m, Anichtchenko A. leg & det. (cAA).
Copyright © Alexander Anichtchenko

Bembidion (Nepha) genei trinacriae (Bonavita & Vigna-Taglianti, 2010)
Female adult
S Spain, Sierra de las Nieves, Los Quejigales 6-8.4.2010, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014
Habitat
northern Kunashir, 2.5 km NW of Cape Nelyudimyi
Copyright © Kirill Makarov

Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014 Holotype
Male adult
Northern Kunashir, 2.5 km NW of Cape Nelyudimyi, N44°29.433', E146°11.783', 4.VIII.2013, leg. K. Makarov & Y. Sundukov” and “HOLOTYPE Bembidion (Nipponobembidion) ruruy Makarov & Sundukov 2014 [printed on red paper]
Copyright © Kirill Makarov

Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014
Endophallus
Male genital apparatus of Bembidion (?Nipponobembidion) ruruy sp. n., holotype. Scale bars 0.25 mm. A aedeagus, right side B aedeagus, left side C right paramere D left paramere E sternite 9, ventral view F sternite 8 G tergite 8.
Copyright © Kirill Makarov

Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014 Paratype
Female adult

Copyright © Kirill Makarov

Bembidion (Notaphemphanes) ephippium (Marsham, 1802)
Male adult
S Hungary, Kiskunsági Nemzeti Park; 3,2mm
Copyright © Aleš Sedláček

Bembidion (Notaphemphanes) ephippium (Marsham, 1802)
Female adult

Copyright © Kirill Makarov

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult
4,4 mm
Namibia, Etosha National Park, Okaukuejo, 1400 mб leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012
Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Bembidion (Notaphocampa) foveolatum Dejean, 1831
Female adult
Bembidion sobrinum Boheman, 1848 det. M.Baehr, 2014; 4,2 mm. Namibia, Otjiwarongo, Kalkfeld, 30 km East, Mt. Etjo Safari Lodge, 1500 m, leg. U.Schmidt, 15.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult

Copyright © Kirill Makarov

Bembidion (Notaphus) patruele Dejean, 1831
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Notaphus) semipunctatum (Danovan, 1806)
Female adult

Copyright © Kirill Makarov

Bembidion (Notaphus) obliquum Sturm 1825: 160
Male adult
Russia, Ивановская обл., Пестяковский р-н, окр. дер. Пурежка, бер. р. Лух 24-25.VI.2011 М. Смирнов. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Notaphus) contractum Say, 1823
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Notaphus) indistinctum Dejean, 1831
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Notaphus) flohri Bates, 1878
Male adult

Copyright © David Maddison

Bembidion (Notaphus) brownorum Maddison, Sproul & Will, 2023 Holotype
Male adult
EMEC: 39.13841/−122.34621 USA: California: Colusa Co. Antelope Valley, Freshwater Creek uv light pan trap 133 m. 1.vii.2021 K.Will
Copyright © David Maddison

Bembidion (Notaphus) mormon Hayward, 1897
Male adult

Copyright © David Maddison

Bembidion (Notaphus) callens Casey, 1918 Paralectotypes
Female adult
Tuçson [Pima County], Arizona.
Copyright © David Maddison

Bembidion (Notaphus) obtusangulum Leconte, 1863
Male adult

Copyright © David Maddison

Bembidion (Notaphus) varium (Olivier, 1795)
Female adult
CZECH REPUBLIC - Olomouc dist., Skalička, 8.IV.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Notholopha) scitulum (Erichson, 1834)
Male adult
CHILE: Reg. VII: Los Niches E of Curicó, DRM voucher V100790; Image from Maddison D.R., 2014. Scale bar 1 mm.
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Bembidion (Notholopha) sexfoveatum Germain, 1906
Male adult
CHILE: Reg. IX: 16.3 km E Malalcahuello, Cuesta Las Raices, DRM voucher V100598.. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Nothonepha) eburneonigrum Germain, 1906
Male adult
Argentina: Neuquén: Rio Neuquén at Chos Malal, DRM voucher V100785. Image from Maddison D.R., 2014.
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Bembidion (Nothonepha) tetrapholeon Maddison, 2014 Paratype
Male adult
Argentina: Neuquén: Arroyo Queñi at Lago Queñi, 830m, 40.1575°S, 71.721°W, / 10–11.ii.2007. DRM 07.035. D.R. Maddison, S.A.Roig. Scale bar 1 mm. Image from Maddison D.R., 2014.
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Bembidion (Nothonepha) tetrapholeon Maddison, 2014 Paratype
Female adult
Argentina: Chubut: Rio Azul at Lago Puelo. Scale bar 1 mm. Image from Maddison D.R., 2014.
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Bembidion (Nothonepha) lonae Jensen-Haarup, 1910
Male adult
Argentina: Mendoza: Salinas del Diamante. Scale bar 1 mm. Image from Maddison D.R., 2014.
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Bembidion (Nothonepha) engelhardti engelhardti
Male adult
Argentina: Neuquén: Rio Salado at route 40. Scale bar 1 mm. Image from Maddison D.R., 2014.
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Bembidion (Nothonepha) germainianum (Toledano, 2002)
Male adult
Argentina: Neuquén: Rio Salado at route 40. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Nothonepha) tucumanum (Jeannel, 1962)
Male adult
Argentina: Mendoza: Salinas del Diamante. Scale bar 1 mm. Image from Maddison D.R., 2014.
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Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Colorado: Huerfano Co., Huerfano River at Badito, 37.7285°N 105.0167°W.
Scale bar is 1 mm.
Photo David R. Maddison & Alexander P. Swanson
Copyright © ---

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 (Label) Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033. (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html). Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu. Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Label
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) zolotarewi Reitter, 1910b: 313
Male adult
Russia, NW Caucas, Krasnodar reg., Tuapse distr., Podchrebetovoe vicinity
(NW Кавказ, Краснодарский край, Туапсинский р-н, окр пос. Подхребeтовое 19.V.2012 leg. М. Смирнов)
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Peryphanes) grapii Gyllenhal, 1827
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov <br /> Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Peryphanes) yukonum Fall, 1926: 131
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) transsylvanicum Bielz, 1852
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Marmarosh Mts., near Lug vill., bank of Kuziy (Kizi) stream, 400m, 26.IV.2014, Panin R. & Kanarsky Yu. leg. L= 5 mm
Copyright © Ruslan Panin

Bembidion (Ocydromus) decorum caraffai Sainte-Claire Deville, 1906a: 15
Female adult
France Corse
Copyright © Pascal Deschamps

Bembidion (Ocydromus) decorum subconvexum K. Daniel & J. Daniel, 1902: 102
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) saxatile Gyllenhal, 1827: 406
Female adult

Copyright © Kirill Makarov

Bembidion (Ocydromus) hasurada Andrewes, 1924a: 77
Male adult
India, Uttarakhand, 10-12.iV.2012, 14km E Uttarkashi, Anichtchenko A. leg.; 5 mm. det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) transversale Dejean, 1831
Male adult
Images from David R. Maddison & Alexander P. Swanson, 2010
Copyright © - - -

Bembidion (Ocydromus) saxatile morsum Netolitzky, 1930
Male adult
Tajikistan, Khoja Obi Garm, 15.9.2011, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) corgenoma Maddison, 2020 Paratype
Male adult
USA: Oregon: Benton Co., Corvallis, Willamette River, 44.5491°N, 123.2449°W, 60 m
Copyright © David Maddison

Bembidion (Ocydromus) transversale Dejean, 1831
Male adult
Canada: Alberta: Burbank, junction of Red Deer and Blindman Rivers, 52.3542°N, 113.7556°W (voucher V101454)
Copyright © David Maddison

Bembidion (Ocydromus) erosum (Motschulsky, 1850)
Male adult
USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051°N, 124.1005°W (V101453)
Copyright © David Maddison

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Arizona: Cochise Co., San Pedro R at Charleston, 31.6239°N, 110.1722°W (voucher V101461)
Copyright © David Maddison

Bembidium haplogonum Chaudoir, 1868 (NEOTYPE)
Male adult
USA: California: Lake Co., North Branch Cache Creek at hwy 20, 305 m 38.9881°N, 122.5400°W
Copyright © David Maddison

Bembidion (Ocydromus) sarpedon Casey, 1918
Male adult
USA: Colorado: Las Animas Co., Cokedale, Reilly Canyon, 37.1346°N, 104.6114°W (V101459)
Copyright © David Maddison

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,7
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,7
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,8
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Urup Isl., Ukromnaya Bay, sea shore rocky debris, 20.viii.1996, leg. K.Eskov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5,2
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5.0
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,75 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Spermatheca
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,6 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,85 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,45
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=5.0 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=4,7 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,55 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Spermatheca
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,75
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Spermatheca
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,9 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Spermatheca
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,8
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,3 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,6 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=5 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male genitalia
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov
Copyright © Kirill Makarov

Bembidion (Ocydromus) fontinale (Raffray, 1885)
Male adult
Ethiopia Amhara, Mt. Abuna yosef, n-slope, 3800-3950 m, 12°07’52”N 39°11’39”E, 4.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) fontinale (Raffray, 1885)
Male genitalia

Copyright © Joachim Schmidt

Bembidion (Ocydromus) basistriatum Fairmaire, 1893
Male adult
Ethiopia, Oromia, Mt Enkuolo, NE-slope, 3100-3200m, 07°24’41”N 39°22’26”E, 5-7.Xii.16, leg. J. Schmidt (CSChM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) melanoceroides Toledano, Bonavita & Schmidt, 2021 Paratype
Male adult
Ethiopia, Oromia, Mt Enkuolo, N-slope, 3350-3600m, 07°22’48”N 39°21’07”E, 6-9.Xii.16, leg. J. Schmidt (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocydromus) melanocerum Chaudoir, 1876
Male adult
Ethiopia, Amhara, W-slope Mt. Choke, 2750-2900 m, 10°38’N 37°45’E, 1.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) terryerwini Neri & Toledano, 2021 Paratype
Male adult
N 29°21'14.7" E 57°20'31.7" / Iran, Prov. Kerman, Rāyen, Goroah, / Babtahoune, Bachufer-Schotterbank / 06.06.2014 2700 m / leg. Schnitter
Copyright © Luca Toledano

Bembidion (Ocydromus) hiekei Muller-Motzfeld, 1986b: 171 Paratype
Male adult

Copyright © Luca Toledano

Bembidion (Ocydromus) sokolowskii Fassati, 1957b: 154 Holotype
Male adult
MHNG
Copyright © Luca Toledano

Bembidion (Ocydromus) decorum decorum (Panzer, 1799b)
Male adult
CZECH REPUBLIC - Staré Hamry, 22.IX.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Ocydromus) modestum (Fabricius, 1801)
Male adult
CZECH REPUBLIC - Olomouc dist., Skalička, 11.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.7 mm
Copyright © Kirill Makarov

Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Female adult

Copyright © Kirill Makarov

Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.9 mm
Copyright © Kirill Makarov

Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Endophallus

Copyright © Kirill Makarov

Bembidion (Ocyturanes) praeustum Dejean, 1831
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Ocyturanes) christophi Schmidt, 2018 Paratype
Male genitalia
“S TIBET 15.VII.2010 / Tibetan Himalaya / above Kampa La 48-4850 m” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) christophi Schmidt, 2018 Paratype
Male adult
“S TIBET 15.VII.2010 / Tibetan Himalaya / above Kampa La 48-4850 m” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) gracilipenne Schmidt, 2018 Paratype
Male genitalia
“S TIBET, N Lhasa Valley / above Chupsang Monast. / 19.IX.2010, 4900m / 29°44’01N 91°07’47E ” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) gracilipenne Schmidt, 2018 Paratype
Male adult
“S TIBET, N Lhasa Valley / above Chupsang Monast. / 19.IX.2010, 4900m / 29°44’01N 91°07’47E ” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) lhatseense Schmidt, 2018 Holotype
Male genitalia
“S TIBET 17.VI.2011 / above pass Tsuo La / 47-4940 m, leg. Schmidt / 29°03’37N 87°58’52E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) lhatseense Schmidt, 2018 Paratype
Male adult
“S TIBET 17.VI.2011 / above pass Tsuo La / 47-4940 m, leg. Schmidt / 29°03’37N 87°58’52E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) eucheres Netolitzky, 1943
Male adult
Tajikistan, Khoja Obi, Garm, 15.9.2011, Barsevskis A. leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) samai Neri & Toledano, 2017 Paratype
Male adult
Afghanistan, Vt. N Salang 1450 m, Khenjan, bord torrent, 11.8.1975, leg. Ledoux (DUBC)
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) muilwijki Neri & Toledano, 2017 Paratype
Male adult
S Iran, Sisakht Dena, 2500-3000 m, 13-14.6.1973
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) dyscheres Netolitzky, 1943
Female adult
N Iran, C Elburz, Val. Darband Sar, 2500-3000m 18.7.1970 (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) montislunae Neri & Toledano, 2022 Holotype
Male adult
Oman, Dhofer province / Jabal al Qamar / 20 km N Dhalqut / N 16.71092° E 53.15350° / (1100 m.n.m.) 28.8.2012 / lgt. P. Kučera (CTVR). Image from Paolo Neri & Luca Toledano, 2022
Copyright © - - -

Bembidion (Ocyturanes) ivanloebli Neri & Toledano, 2018 Holotype
Male adult
“Iran, Fars, Sepidan, 21-9-2005, Muilwijk leg.” (CTVR)
Copyright © Luca Toledano

Bembidion (Ocyturanes) waziristanum Andrewes, 1932c: 863
Male adult
Pakistan, W. Himalai mts., 20 km SW Chilez (CTVR)
Copyright © Luca Toledano

Bembidion (Ocyturanes) lobanovi Mikhailov, 1984b: 297
Female adult
N Tajikistan, Fanskie Mts. Khurdak (CTVR). Scale: 1 mm
Copyright © Luca Toledano

Bembidion (Ocyturanes) signatipenne Jacquelin du Val, 1852
Male adult
Imago: NW Turkey, Edremit, Kaz Dağ, Ay i Deresi (PN); Aedeagus specimen from Crete, Rethimno, Plakias, river Megalopotamos (PN).
Copyright © Luca Toledano

Bembidion (Ocyturanes) viduum Netolitzky, 1910a: 215 Holotype
Male adult
Kilikisch. Taurus (NHMW). Scale: 1 mm.
Copyright © Luca Toledano

Bembidion (Ocyturanes) kurdistanicum Netolitzky, 1920
Male adult
Imago: Iran, Mazandaran Prov., Karaj-Chalus road (MNB); Aedeagus: Holotype, Persia Septentr., Kerim (NHMW).
Copyright © Luca Toledano

Bembidion (Ocyturanes) heinzi Korge, 1971: 16 Paratype
Male adult
Anatolia bor., Köse Flußufer (MNB). Aedeagus of Holotype, Anatolia bor., Köse Flußufer (MNB).
Copyright © Luca Toledano

Bembidion (Ocyturanes) antennarium (Morvan, 1972) Holotype
Male adult
Iran, Zagros, Kurang (PM). Scale 1mm
Copyright © Luca Toledano

Bembidion (Ocyturanes) mirzayani (Morvan, 1973) Paratype
Female adult
Iran, Zagros Mts., Sisakht (PM). Scale: 1 mm.
Copyright © Luca Toledano

Bembidion (Omoperyphus) kempi Andrewes, 1922b: 178
Male adult
India, Uttarakhand, 13-15.4.2012, Uttarkashi distr., left tributary of Bhagirathi riv., 2150m, N30*57'43" E78*41'54.75", Anichtchenko A. leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Omoperyphus) hypocrita Dejean, 1831
Male adult
Spain, Jaen prov., Arroyo de la piedra del Aguila, 37.683885, -3.485062, 3-4.5.2014, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Omotaphus) alsium Coquerel, 1866
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Omotaphus) gorilla Bonavita, Toledano & Vigna Taglianti, 2016
Male adult

Copyright © David Maddison

Bembidion (Omotaphus) scotti Netolitzky, 1931
Male adult

Copyright © David Maddison

Bembidion (Omotaphus) mellissii Wollaston, 1869
Male adult

Copyright © David Maddison

Bembidion (Omotaphus) mixtum Schaum, 1863b: 90 Lectotype
Male adult
43101 [handwritten] // mixtum Sch. [handwritten] / dt. Netolitzky [printed] // Hist. -Coll. (Coleoptera) / Nr. 43101 (=55133) / Bembidion mixtum Klug / Aegypt. Ehrenberg / Zool. Mus. Berlin [blue, printed] // SYNTYPE / Bembidium mixtum / Schaum, 1863 / labelled by MFNB 2015 [red, printed] (MNB).
Copyright © Luca Toledano

Bembidion (Omotaphus) aethiopicum Raffray, 1885
Male adult

Copyright © Luca Toledano

Bembidion (Omotaphus) mixtum Schaum, 1863b: 90
Male adult

Copyright © Paolo Bonavita

Bembidion (Omotaphus) kivuanum (Basilewsky, 1951) Holotype
Male adult
Imago - Holotype: “HOLOTYPUS [red, printed] // édéage en preparation microscop. [hand�written] // I.R.S.A.C. – MUS. Congo, Uvira: Bassin Ulindi / 2850m. 29-IX-1950 / N. Leleup [printed] / (Bord d’un marais) [handwritten] // RMCA ENT / 000019230 “ (MRAC); Aedeagus - paratype.
Copyright © Paolo Bonavita

Bembidion (Omotaphus) kilimanum Alluaud, 1908 Holotype
Male adult
“AFRIQUE ORle ALLEMANDE / KILIMANDJARO / (ZONE DES PRAIRIES) / CH. ALLUAUD I-IV 1904 [white, printed] // TYPE [red, printed] // MUSEUM PARIS / Coll. Ch. ALLUAUD [blue, printed] // Bembid. / (Testediolum / kilimanum / n.sp. Alluaud / [white, handwritten]” (MNHN).
Copyright © Paolo Bonavita

Bembidion (Omotaphus) demeyeri Toledano & Bonavita, 2016 Holotype
Male adult
“Uganda Occidental / Province D’UNYORO / Rég. de l’Albert-Nyanza / CH. ALLUAUD II 1909 [white, printed] // MUSEUM PARIS / Coll. Ch. ALLUAUD [blue, printed] // Bembid. (Notaphus) / Bakeri / type. Alluaud / Alluaud détérm [white, handwritten, with a blue strip on top]” (MNHN)
Copyright © Paolo Bonavita

Bembidion (Omotaphus) gorilla Bonavita, Toledano & Vigna Taglianti, 2016 Holotype
Male adult
? Holotype. “RWERU: 2800/2900M, 20 - VI – 1964, Tamisage, Bambousaise // Coll. Mus. Tervuren, Zaire: P. N. Virunga, Volcan Mikeno, R. P. Celis VI-64” (MRAC)
Copyright © Paolo Bonavita

Bembidion (Omotaphus) mellissii Wollaston, 1869
Male adult
Image from Bonavita et al. 2016, with modifications
Copyright © Paolo Bonavita

Bembidion (Omotaphus) madagascariense Chaudoir, 1876
Male adult

Copyright © Paolo Bonavita

Bembidion (Omotaphus) picturatum Fairmaire, 1898
Male adult

Copyright © Paolo Bonavita

Bembidion (Omotaphus) scotti Netolitzky, 1931
Male adult

Copyright © Paolo Bonavita

Bembidion (Omotaphus) uhligi Toledano & Maddison, 2016 Holotype
Male adult
Holotype? RSA-Western Cape, NE of Botrivier env., 34°13’S, 19°12’E, 24.II.2014 R. Kmeco leg.
Copyright © Paolo Bonavita

Bembidion (Omotaphus) tropicum Chaudoir, 1876
Male adult

Copyright © Paolo Bonavita

Bembidion (Pamirium) zlotini Mikhailov, 1996: 71
Female adult
det. I.Kabak et B.Kataev SE Kazakhstan, Zailiysky Alatau mts. Kok-Zhaylau. N 43°08'15" / E 77°00'08". h=2270 m. 24-26.06.2013. S.Kolov leg. Image of S. Kolov, from www.zin.ru
Copyright © - - -

Bembidion (Pamirium) platypterum Solsky, 1874: 120
Female adult
Kazakhstan, Alma-Ata, 2.06.1990. det. Anichtchenko A. 2016
Copyright © Alexander Anichtchenko

Bembidion (Paraprincidium) karinae Schmidt, 2009 Holotype
Female adult
NEPAL, Prov. Koshi, Tumlingtar, Arun-Ufer [Arun river bank], 300 mNN, 20.X.2002 leg. J. Weipert (Naturkundemuseum Erfurt). Image from Schmidt, 2009
Copyright © - - -

Bembidion (Peryphanes) lacunarium (Zimmermann, 1869)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphanes) geberti Marggi, 2011 Holotype
Male adult
Türkei, Provinz Nevşehir, NE Avanos Kızılırmak, temporärer Flusslauf, 24.5.2001, leg. J. Gebert [38°46.070 N, 34°53.755 E], coll.
Naturhistorisches Museum Bern (NMBE).
Foto: Ortwin Bleich
Copyright © ---

Bembidion (Peryphanes) adygorum Belousov & Sokolov, 1996
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Peryphanes) olegleonidovici Fassati, 1990: 169
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) milleri Jacquelin du Val, 1852
Male adult
4,5 mm; Germany, Rheinland-Pfalz, Eifel, Gerolstein, leg. J.Scheuern, 30.IV.1983; det. F.Koehler, 1994; Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Bembidion (Peryphanes) dalmatinum dalmatinum Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) adygorum Belousov & Sokolov, 1996: 14
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) phryganobium Belousov et Sokolov, 1996:24
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=6.5 mm
Copyright © Kirill Makarov

Bembidion (Peryphanes) phryganobium Belousov et Sokolov, 1996:24
Female genitalia
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Peryphanes) phryganobium Belousov et Sokolov, 1996:24
Spermatheca
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=5.7 mm
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Male genitalia
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Female adult
Iran, Lorestan prov., Dorud distr., near Cham Chit vill. 13.V.2017 leg. D.Kasatkin L=5.6 mm
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Female genitalia
Iran, Lorestan prov., Dorud distr., near Cham Chit vill. 13.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Spermatheca
Iran, Lorestan prov., Dorud distr., near Cham Chit vill. 13.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov

Bembidion (Peryphanes) lirykense Reitter, 1908
Male adult
Iran, 40km S Ardabil, Goli, 20.5.2005, Anichtchenko A. leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphanes) fraxator Menetries, 1832
Male adult
Russia, Karachaevo-Cherkessia, urochishe Dzhalpak, 17-19.5.2009, A. Anichtchenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphanes) sanatum Bates, 1883: 274 Holotype
Male adult
“Type [round, bordered in red] // Japan / G. Lewis / 1910 – 310 [printed] // B. / sanatum / Bates [handwritten]” (NHMUK).
Copyright © Luca Toledano

Bembidion (Peryphanes) sanatum sanatum
Male adult
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum sanatum
Male genitalia
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum sanatum
Female adult
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum sanatum
Female genitalia
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum iwanai Makarov & Sundukov, 2021 Holotype
Male adult
Kunashir Island, valley of Kislaya River, below hot springs, 18.VI.2011, leg. A. Matalin
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum iwanai Makarov & Sundukov, 2021 Holotype
Male genitalia
Kunashir Island, valley of Kislaya River, below hot springs, 18.VI.2011, leg. A. Matalin (HT)
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum iwanai Makarov & Sundukov, 2021 Paratype
Female adult
Kunashir Island, valley of Kislaya River, below hot springs, 18.VI.2011, leg. A. Matalin
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum iwanai Makarov & Sundukov, 2021 Paratype
Female genitalia
Kunashir Island, valley of Kislaya River, below hot springs, 18.VI.2011, leg. A. Matalin
Copyright © Kirill Makarov

Bembidion (Peryphanes) milleri carpathicum J. Muller, 1918b: 88
Male adult
W Ukraine, W Podolian Upland, Rogatyn Opillia, Zaluzhzhya vill, 49°24'33"N 24°35'36"E, clay quarry, 30-31.VIII.2018, Panin R leg., L=4.6mm
Copyright © Denis Khrapov

Bembidion (Peryphanes) cf brunnicorne Dejean, 1831
Male adult
W.Ukraine, Transcarpathian, Polonynian Beskids, Yavirnyk Rng, h=600 m, near Ruskyi Mochar vill.,15.V.2021, V. Khrapova leg./R. Panin det., L=4.7 mm
Copyright © Denis Khrapov

Bembidion (Peryphanes) deletum deletum Audinet-Serville, 1821: 77
Male adult
CZECH REPUBLIC - Beskydy Mts., Kněhyně, 27.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Peryphanes) stephensii Crotch, 1866
Female adult
CZECH REPUBLIC - Moravian-Silesian reg., Opava dist., Závada, 25.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Testediolum) armeniacum kokandicum Solsky, 1874: 120
Female adult

Copyright © Kirill Makarov

Bembidion (Testediolum) gagates gagates Andrewes, 1924a: 72
Male adult
India, Uttarakhand, 13-15.4.2012, Uttarkashi distr., left tributary of Bhagirathi riv., 2300m, N30*57'41.57" E78*41'54.75", Anichtchenko A. leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Peryphidium) validum salangense Neri & Toledano, 2021 Paratype
Male adult
“Afghanistan / Lac du Salang 3700 m / 4.8.74 Leg. Ledoux”
Copyright © Luca Toledano

Bembidion (Peryphiolus) caligatum Jeanne & Muller-Motzfeld, 1982
Male adult
Spain, Guadarrama, Cotos, 25.8.2001, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Bembidion (Peryphiolus) monticola monticola Sturm, 1825: 135
Male adult
CZECH REPUBLIC - Frýdek-Místek dist., Staré Hamry, 22.IX.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Peryphodes) ephippigerum (Leconte, 1852)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Peryphophila) problematicum Schmidt, 2018 Holotype
Male genitalia
“NEPAL, Ghatte / Kirola – Khola. Camp / 3500m. Pasang 12.5.88” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphophila) problematicum Schmidt, 2018 Holotype
Male adult
“NEPAL, Ghatte / Kirola – Khola. Camp / 3500m. Pasang 12.5.88” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphophila) pseudoclarum Schmidt, 2018
Male genitalia
“NEPAL Manaslu Mts. / Marsyangdi Khola vall. / Bhachok Khola 1000m / 13.4.1999 leg. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphophila) pseudoclarum Schmidt, 2018 Holotype
Male adult
“NEPAL Manaslu Mts. / Marsyangdi Khola vall. / Bhachok Khola 1000m / 13.4.1999 leg. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) tetracolum Say, 1823: 89
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphus) petrosum Gebler, 1833: 275
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Label
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) ehikoense Habu, 1984 (Label) Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Peryphus) tetracolum Say, 1823: 89
Male adult
Germania, Bayern, Oberfranken, Kulmbach, Melkendorf
leg.det. U.Schmidt, 1975
Photo: U.Schmidt, 2011
Copyright © U.Schmidt

Bembidion (Peryphus) captivorum Netolitzky, 1943
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) subcostatum subcostatum (Motschulsky, 1850)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) bruxellense Wesmael, 1835
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) obscurellum obscurellum (Motschulsky, 1845)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) captivorum Netolitzky, 1943a: 29
Male adult
Россия, SE Приморский кр., Лазовский зап., ~12 км S пос. Старая Каменка, окр. кордона Америка, 43°15'N / 134°01'E, 10.07.04, leg. М. & Л. Смирновы. image from www.zin.ru
Copyright © Maksim Smirnov

Bembidion (Peryphus) tetracolum tetracolum Say, 1823: 89
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) bracculatum Bates, 1889b: 212
Female adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A.; 5.5 mm, det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion () eutherum Andrewes, 1923b: 96
Male adult
India, Uttarakhand, 15 km SW New Tehri, 869m, 18-20.4.2012, Anichtchenko A. leg.; det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) jedlickai jedlickai Fassati, 1945: 22
Female adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) bualei bualei Jacquelin du Val, 1852
Male adult
Italy, Lombardia, Brescla, Sabbio Chiese, 20.5.2011, E. Ruzzier leg.
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) hummleri hummleri J. Muller, 1918
Male adult
Morocco, Quarzazate prov., Quwd Drag N 30*30'26" W006*06'01", near Tin-Zoulin vill., 27.6.2010, A. Anichtchenko leg. det. M. Toribio, 2017
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) irregulare Netolitzky, 1934b: 90
Male adult
Shyock valley betw. Karghalik & Saser Brangsa, 4150-4500 m, 24.VI-11.VIII.1930
Copyright © Jan Muilwijk

Bembidion (Peryphus) sejunctum Casey, 1918
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Peryphus) ahrensi Schmidt, 2018 Holotype
Male genitalia
“Uttar Ganga Ebene / bei Dhorpathan / 3000 m, 6.-8.3.”, “C-NEPAL-Himalaya / Dhaulagiri-Mts. / leg. Ahrens 1994” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) ahrensi Schmidt, 2018 Paratype
Male adult
“Uttar Ganga Ebene / bei Dhorpathan / 3000 m, 6.-8.3.”, “C-NEPAL-Himalaya / Dhaulagiri-Mts. / leg. Ahrens 1994” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) aparupa Schmidt, 2018 Paratype
Male genitalia
“W-NEPAL Baglung Lekh / W Pokhara, 10-15km NW / Baglung, 2350-2550m / leg. Schmidt 10-12.5.04 / 28°18‘36”N 83°31‘56“E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) aparupa Schmidt, 2018 Paratype
Male adult
“W-NEPAL Baglung Lekh / W Pokhara, 10-15km NW / Baglung, 2350-2550m / leg. Schmidt 10-12.5.04 / 28°18‘36”N 83°31‘56“E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) dhaulagiricum Schmidt, 2018 Holotype
Male genitalia
“NEPAL W Dhaulagiri / below Dhule 26-2700m / 7.IX.12 leg. J. Schmidt / 28°41‘07N 82°56‘31E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) dhaulagiricum Schmidt, 2018 Holotype
Male adult
“NEPAL W Dhaulagiri / below Dhule 26-2700m / 7.IX.12 leg. J. Schmidt / 28°41‘07N 82°56‘31E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) glabripenne Schmidt, 2018 Holotype
Male genitalia
“NEPAL, Prov. Karnali / 29°04,33‘N 82°42,41’E / Bachtal bei Pahada / 3100-3400 m NN / 2.VI.1997 / leg.: J. Weipert” (NME).
Copyright © Joachim Schmidt

Bembidion (Peryphus) glabripenne Schmidt, 2018 Paratype
Female adult
“Nepal, District Karnali, / Nähe Pohada 02.06.1997 / 3010-3400 mNN, leg. Grill, / Flußufer / HF, 29°04’N, 82°42’E” (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Peryphus) sillemi Netolitzky, 1935b: 92 Holotype
Unsexed

Copyright © Jan Muilwijk

Bembidion (Peryphus) cordicolle Jacquelin da Val, 1852
Male adult
Turkey, Kemer distr., Mts.Taurus, Ulupinar riv.,36*27'01N 30*26'14E 21.9.2014, I. Enuschenko leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) persicum Menetries, 1832
Male adult
Tajikistan, Khoja Obi, Garm, 15.9.2011, Barsevskis A. leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) abbreviatum pulpani Fassati, 1955
Male adult
Tadzhikistan, Ayni distr., Sughd prov., Iskanderkul lake, 2300m 5.7.2012, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) baleense Toledano, Bonavita & Schmidt, 2021 Holotype
Male adult
Ethiopia, Oromia, Bale Mts., Sebsebe Washia Forest, Salgen Vall. 3130 m, 07°02’08”n 39°36’06”e, 4.ii.2019, leg. R.Emmerich, J. Schmidt, Yeshitla M. (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) baleense Toledano, Bonavita & Schmidt, 2021
Male genitalia

Copyright © Joachim Schmidt

Bembidion (Peryphus) damota Toledano, Bonavita & Schmidt, 2021 Holotype
Male adult
Ethiopia, Sodo, Mt. Damot, 2380 m, 06°53’26”N 37°46’51”E, 3.Xii.2017, leg. D. Hauth, J. Schmidt, Yeshitla M.” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) damota Toledano, Bonavita & Schmidt, 2021
Male genitalia

Copyright © Joachim Schmidt

Bembidion (Peryphus) scottustulatum Netolitzky, 1937
Male adult
Ethiopia, Amhara, n-slope Mt. Choke, 3800-3950 m, 10°43’16”N 37°51’15”E, 26.ii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSChM)
Copyright © Joachim Schmidt

Bembidion (Peryphus) scottustulatum Netolitzky, 1937
Male genitalia

Copyright © Joachim Schmidt

Bembidion (Peryphus) psuchrum Andrewes, 1922c: 241
Male adult
Inde, Himachal Pradesh, Mahari, 3500 m (CTVR)
Copyright © Luca Toledano

Bembidion (Peryphus) tricuspis Neri & Toledano, 2018 Paratype
Male adult
“SW Iran, Hoseiniyeh / 28 km NNW Andimeshk / 12-13. 4. 1977, 360 m // Loc. no. 286 / Exped Nat Mus / Praha” (NMPC).
Copyright © Luca Toledano

Bembidion (Peryphus) subcostatum vau Netolitzky, 1913: 213
Male adult
CZECH REPUBLIC - Olomouc reg., Přerov dist., Skalička, 8.IV.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Philochthus) pallidiveste Carret, 1905
Male adult
Volgogrda Area, Pallasovka Distr., near Elton Vill. 4.V.2009 leg. K.Makarov
L = 3.8
Copyright © Kirill Makarov

Bembidion (Philochthus) guttula (Fabricius, 1792)
Male adult
length: 2,8-3,8 mm
Germania, Bayern, Oberfranken, Rudolfstein
leg.det. U.Schmidt, 2001
Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Philochthus) biguttatum (Fabricius, 1779)
Male adult

Copyright © Kirill Makarov

Bembidion (Philochthus) guttula (Fabricius, 1792)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) inoptatum Schaum, 1857a: 149
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) mannerheimii C.R. Sahlberg, 1827
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) cf. neresheimeri J. Muller, 1929
Female adult
Ивановская обл., г. Иваново, парк, 1905 г., 29.05.2014, М. Смирнов leg.; det. K.Makarov. Image from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Philochthus) lunulatum (Geoffroy, 1785)
Male adult
Turkey, Marmaris, Hisarönü, leg. U.Schmidt, 18.V.1997; det. Marggi, 2014; Photo: U.Schmidt, 2014. 3,8 mm
Copyright © Udo Schmidt

Bembidion (Philochthus) biguttatum (Fabricius, 1779)
Male adult
Russia, Ivanovo, "Kharinka" city park, 30.V.2015, M. Smirnov leg.
Copyright © Maksim Smirnov

Bembidion (Philochthus) ellipticocurtum Netolitzky, 1935c: 167
Female adult
Lebanon, Jbail prov., Laqlouq, 8.5.2000. det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Philochthus) neresheimeri J. Muller, 1929: 101
Male adult
W Ukraine, Lviv city, Sykhiv, waste-ground, 49°47'29"N 24°3'18"E, 7.IV.2018, Panin R. leg., L=3.1mm
Copyright © Denis Khrapov

Bembidion (Phyla) obtusum Audinet-Serville, 1821: 83
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Phyla) phoeniceum Huber & Marggi, 1997: 778
Female adult
Turkey, Kirish distr., Camuyuva riv. 28.9.2014, I. Enischenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Phyla) incommodum Netolitzky, 1926
Male adult
Iran, 40 km S Ardabil, Goli, 20.5.2005, Anichtchenko A. leg. (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Plataphus) simplex Hayward, 1897
Male adult
Great Smoky Mountains National Park. photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Plataphus) gebleri (Gebler, 1833)
Male adult
Far East, Lazovskij Nat. Res., Formozova spring 43°00'06"N 134°05'13"E 10.VII.2005 K.Makarov leg.
L=4.5 mm
Copyright © Kirill Makarov

Bembidion (Plataphus) prasinum (Duftschmid, 1812)
Female adult

Copyright © Kirill Makarov

Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 (Label) Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) fujiyamai Habu, 1958 (Label) Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Alexander Anichtchenko

Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) tetraporum Bates, 1883: 270
Female adult
Kuriles isl., Kunashir
Copyright © Kirill Makarov

Bembidion (Plataphus) gebleri persuasum Netolitzky, 1938a: 38
Male adult
SE Приморский кр., Лазовский зап., побережье Японского моря, бухта Тачингоуза, 27.06.04 leg. М.& Л.Смирновы
Russia, SE Primorie, Lazo Nat. res., seashore, Tachingouz bay
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Plataphus) infuscatipenne Netolitzky, 1938
Male adult
Russia, SE Primorie, Lazo Nat. res., ~10 км SW Sokolchi vill., Korpad' vicinity 43°15'N / 134°7'E, 23-27.06.10, leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Plataphus) lucillum lucillun Bates, 1883
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Plataphodes) tetraporum Bates, 1883: 270
Endophallus
Aedeagus of Bembidion (Plataphodes) tetraporum Bates, 1883. Scale bars 0.25 mm. A right side B left side.
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Male adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Endophallus
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Endophallus
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion (Plataphus) coelestinum (Motschulsky, 1844)
Female adult
N Mongolia, Tuul-Gol R., Engin Aimak Tev, 14.8.2013, I. Enuschenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Plataphus) difficile (Motschulsky, 1844)
Male adult
Russia, Tuva rep.,Todzhynsk distr., E Sayan mts., Arzhkhan-Khem, 2200 m, 21.6.2017, I. Enuschenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Plataphus) pseudolucillum Netolitzky, 1938a: 37
Male adult
Kibune, Kyoto (NHMW)
Copyright © Luca Toledano

Bembidion (INCERTAE) amurense Motschulsky, 1859
Male adult
photo Makarov K.
Copyright © Kirill Makarov

Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) umeyai Habu, 1959 (Label) Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) gotoense Habu, 1973 (Label) Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) phaedrum Andrewes, 1923b: 97
Female adult
India, Uttarakhand, 15 km SW New Tehri, 869 m, N30*15'874" E 078*21'587", 18-20.4.2012, A. Anichtchenko leg.; 5 mm; det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Politophanes) inventor Schmidt, 2018 Holotype
Male genitalia
“E. NEPAL: KOSI / Val. Induwa Kola / 2000 m, 18.IV.84 / Löbl - Smetana” (MHNG).
Copyright © Joachim Schmidt

Bembidion (Politophanes) inventor Schmidt, 2018 Holotype
Male adult
“E. NEPAL: KOSI / Val. Induwa Kola / 2000 m, 18.IV.84 / Löbl - Smetana” (MHNG).
Copyright © Joachim Schmidt

Bembidion (Princidium) compactum Andrewes, 1922c: 240
Female adult
India, Uttarakhand, 10 km NNE of Ramnagar, gorge of left trib. of Kosi River, 400-450 m,
29°29' N, 79°09,5' E, R. Dudko, I. Lyubechanskii, 19-20.04.2012
det. R. Dudko 2012
Copyright © Roman Dudko

Bembidion (Princidium) punctulatum punctulatum s.str.
Male adult
Germany, Bavaria, Ascholding, Isar, leg. det. U.Schmidt, 3.V.1975 Photo: U.Schmidt, 2012
Copyright © Udo Schmidt

Bembidion (Princidium) punctulatum Drapiez, 1820
Male adult
Pechoro-Ilychskii N.P. (Печеро-Илычский зап)
Copyright © Kirill Makarov

Bembidion (Pseudophilochthus) nubigena (Wollaston, 1877)
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Pseudophilochthus) evanescens (Wollaston, 1877)
Female adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) gemmulipenne (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) fossor (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) trechoides (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) sublimbatum (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) dicksoniae (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) grayanus (Wollaston, 1877)
Male adult

Copyright © David Maddison

Sakagutia umi Sasakawa, 2007
Male adult
Far East, Lazo Distr., 4 km VE Preobrazhenie Vill., Ezhovaja Bay 42°55'59N 26.VII.2005 133°59'29E K.Makarov leg.<br /> L=5.5 mm
Copyright © Kirill Makarov

Sakagutia umi Sasakawa, 2007
Male adult
SE Приморье, Лазовский р-н., побережье Японского моря, 3 км NE пос. Преображение бухта Ежовая (Тасовая), литораль, 12.07.2010, leg. М. Смирнов. Image from www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Sakagutia umi Sasakawa, 2007
Habitat

Copyright © Dooyoung Kim

Bembidion (Sakagutia) umi Sasakawa, 2007
Male adult
S. Korea, Gyeongsanbuk-do, Pohang-si, collected by digging the pebbles near the estuary
Copyright © Dooyoung Kim

Sakagutia umi Sasakawa, 2007
Male genitalia

Copyright © Dooyoung Kim

Bembidion (Semicampa) schueppelii Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Semicampa) chaudoirii Chaudoir, 1850b: 179
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Sloanephila) tahitiense Liebherr & Maddison, 2013 Holotype
Male adult
(MNHN): French Polynesia: Tahiti Nui / Mt. Mauru lava tube / el. 705 m 18-IX-2006 lot 01 / 17˚38.017´S 149˚21.284´W / pyr. fog moss/liverworts / along stream J.K. Liebherr // HOLOTYPE / Bembidion / tahitiense / Liebherr & / Maddison 2012 (black-bordered red label).
Copyright © David Maddison

Bembidion (Sloanephila) jacksoniense Guerin Meneville, 1830
Female adult
Wiluna W.A. / ix 25 1931 //Australia / Harvard Exp. / Darlington (MCZ). Scale bars 0.5 mm.
Copyright © David Maddison

Bembidion (Taiwanobembidion) aliense Habu, 1973
Male adult

Copyright © David Maddison

Bembidion (Talanes) subfasciatum Chaudoir, 1850b: 187
Female adult
Krasnodar Reg., Taman Peninsula, Tuzlo Lake 6.V.2009
2.7 mm
Copyright © Kirill Makarov

Bembidion (Talanes) aspericolle (Germar, 1829)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Terminophanes) pulcherrimum (Motschulsky, 1850)
Male adult
Caucasus, Elbrus, Itkol (CTVR). Image from Toledano L. & Gueorguiev B., 2015
Copyright © - - -

Bembidion (Terminophanes) kaschmirense Netolitzky, 1920
Female adult
India, Uttarakhand, 5 km NE Govind Ghat, road to Gangaria, 9-10.6.2011, Anichtchenko A. leg. det. J. Schmidt 2015 8 mm
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) mckinleyi scandicum Lindroth, 1943
Female adult
E Siberia, Lake Baikal, Malyi Kochergat vill., Goloustnaya riv., 16.VII.1991, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) reibnitzi Schmidt, 2018 Holotype
Male genitalia
“S TIBET Kyi Chu river / betw. Lhasa and airport / ca. 3650 m, 22.IX.2010 / leg. J. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Terminophanes) reibnitzi Schmidt, 2018 Paratype
Male adult
“S TIBET Kyi Chu river / betw. Lhasa and airport / ca. 3650 m, 22.IX.2010 / leg. J. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Terminophanes) terminale kirgisorum Netolitzky, 1934
Male adult
Tajikistan, Ayni distr., Sughd prov., Sarytag, 9km S lake Iskanderkul, 2380m, 6.7.2012, A. Barsevskis leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) consummatum Bates, 1873
Male adult
Russia, E Siberia, Irkutsk reg., Bolshaya Rechka vill, 3.8.1990, Anichtchenko A. leg. & det
Copyright © Alexander Anichtchenko

Bembidion (Testediolum) julianum De Monte, 1943b: 53
Male genitalia

Copyright © Augusto Degiovanni

Bembidion (Testediolum) julianum De Monte, 1943
Male adult
Italy
Copyright © Augusto Degiovanni

Bembidion (Testedium) bipunctatum (Linne, 1761)
Male adult
length: 3,6-4,8 mm
Austria, Schladminger Tauern, Mariapfarr
leg.det. U.Schmidt, 1984
Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Testedium) laetum Brulle, 1836: pl.2
Female adult
Morocco, 65 km N Essaouria, 5 km N Jamaa Shoim, 23-25.III.2011, A. Gusakov. Фотография с сайта zin.ru\Animalia\Coleoptera.
Copyright © Maksim Smirnov

Bembidion (Trepanedoris) concretum Casey, 1918
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Trepanedoris) atripes (Motschulsky, 1844)
Male adult

Copyright © Kirill Makarov

Bembidion (Trepanedoris) faraticolle (Jeanne, 1995) Holotype
Female adult
Image courtesy Biological Museum, Lund University
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Bembidion (Trepanedoris) doris (Panzer, 1797)
Male adult

Copyright © Kirill Makarov

Bembidion (Trepanes) articulatum (Panzer, 1796)
Female adult
length: 3,0-3,9 mm
Germania, Bayern, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1992
Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Trepanes) octomaculatum (Goeze, 1777)
Male adult

Copyright © Kirill Makarov

Bembidion (Trepanes) articulatum (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Trepanes) octomaculatum (Goeze, 1777)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Trichoplataphus) inaense Habu, 1956 (Label) Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Trichoplataphus) ozarkense Maddison & Hildebrandt, 2011 Paratype
Male adult
USA: Missouri: Carter Co., Current River at Van Buren, voucher DRMV100311
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Bembidion (Trichoplataphus) rolandi Fall, 1922
Male adult
USA: Virginia: Rockbridge Co., Maury River at Glasgow, voucher DRMV100193
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Bembidion (Trichoplataphus) parconaturaviva Toledano, 2010: 394
Male adult
SE Maritime Prov., Lazovsky dist., ~10 кm SW Sokoltchi vill., 23-27.VI.2010 leg.М. Smirnov. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов. det. K. Makarov 2016
Copyright © Maksim Smirnov

Bembidion (Trichoplataphus) kara Andrewes, 1921d: 250
Female adult
India, Uttarakhand, 15 km SW New Tehri, 869 m, N30*15'874" E 078*21'587", 18-20.4.2012, A. Anichtchenko leg.; 6 mm; det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Trichoplataphus) deplanatum A. Morawitz, 1862a: 262
Male adult
E Siberia, Irkutsk, Irkut riv., 3.VI.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Trichoplataphus) oxyglymma Bates, 1883
Male adult
Japan, Honshu Pref., Gifu, Tohuyama Dam, Ibigawa river, 16.VII.1992, leg. B. Kuznetsov. det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Zeactedium) giachinoi Toledano, 2005
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Zecillenus karikari Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zecillenus) alacre (Broun, 1921)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zecillenus) albescens (Bates 1878)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zecillenus) tepaki Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zecillenus) tillyardi (Brookes, 1927)
Female adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zecillenus) waimarama Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zecillenus) puponga Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zecillenus) chalmeri (Broun, 1886)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zemetallina) wanakense Lindroth, 1976
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
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Bembidion (Zemetallina) chalceipes Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
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Bembidion (Zemetallina) bullerense Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
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Bembidion (Zemetallina) hokitikense Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
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Bembidion (Zeperyphodes) nesophilum Broun, 1886
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zeperyphodes) callipeplum Bates, 1878
Female adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zeplataphus) tairuense Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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