Carabidae

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Coptodera (Coptodera) relucens Bates, 1869
Male adult
Peru, Junin prov., Satipo env., 800m, 13-14.III.2005, leg. A. Petrov. det. Anichtchenko A. 2015
Copyright © Kirill Makarov

Coptodera (Coptodera) megalops Bates, 1869
Male adult
Peru. det. A. Anichtchenko, 2015
Copyright © Kirill Makarov

Coptodera (Coptodera) stockwelli Shpeley & Ball, 1993
Male adult
Panama, Darien Nat. Park, near Rancho Frio, 4-15.I.2017, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Coptodera (Coptodera) lineata (Bates, 1883) Syntypes
Female adult
Stenoglossa lineata Bates, 1883. "Almolonga, Mexico, Hoege"
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Coptodera (Coptodera) sahlbergi Chaudoir, 1869
Female adult
La Plata, Buenos Aires Province, XI-2014. det. A. Cicchino
Copyright © Armando Cicchino

Coptodera (Coptodera) relucens Bates, 1869
Male adult
Peru, Junin dep. Satipo Rio Venado, 1122m.
Copyright © Augusto Degiovanni
Coptodera sensu stricto taxon page (8 photos)
Coptodera incertae sedis taxon page
Coptodera (Coptoderina) madara Habu, 1957 Holotype
Female adult
Kurokawa, NIIGATA, 03. VIII. 1938, K. Baba Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-172 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Coptodera (Coptoderina) eluta Andrewes, 1923 (madara Habu, 1957 ) Holotype
Female adult
Kurokawa, NIIGATA, 03. VIII. 1938, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-172
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Coptodera (Coptoderina) madara Habu, 1957 Holotype
Female adult
Kurokawa, NIIGATA, 03. VIII. 1938, K. Baba Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-172 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Coptodera (Coptoderina) madara Habu, 1957 Holotype
Label
Kurokawa, NIIGATA, 03. VIII. 1938, K. Baba Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-172 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Coptodera (Coptoderina) subapicalis Putzeys, 1877a: 84
Male adult
Collection ground Shonai machi, OoitaPref. 34095 (C) Working Group for Biological Indicator Ground Beetles Database, Japan Prohibited secondary use of all data
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Coptodera (Coptoderina) subapicalis Putzeys, 1877a: 84
Endophallus
Collection ground Shonai machi, OoitaPref. 34095 (C) Working Group for Biological Indicator Ground Beetles Database, Japan Prohibited secondary use of all data
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Coptodera (Coptoderina) flexuosa Schmidt-Gobel, 1846
Female adult
N INDIA, Uttarakhand, 18-20.04.2012, 15 km SW New Tehri, h = 869 m, N30°15.874' E078°21.587', Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) cyanella Bates, 1869
Male adult
Indonesia E., West Papua, S Bird's Neck, Kaimana 47 km E Triton Bay, Kamaka (former Warika) vill. 03*46'43S 134*10'18E, 10.09.2010, M. Kalins leg. 6 mm det. M. Baehr, 2013
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) fasciolata (W.J.Macleay, 1887)
Male adult
Australia. L 3.3 mm det. M. Baehr. Image from Baehr M., 2014. A new species of the genus Coptodera Dejean, 1825 from Tanimbar Islands
Copyright © Martin Baehr

Coptodera (Coptoderina) bastai Baehr, 2014 Paratype
Male adult
INDONESIA, Tanimbar is., Yamdema isl., 20 km NE of Saumlaki, 150 m. 1.-30.i.2007, M. Obořil lgt. Image from Baehr M., 2014. A new species of the genus Coptodera Dejean, 1825 from Tanimbar Islands. 3.4 mm
Copyright © Martin Baehr

Coptodera (Coptoderina) interrupta Schmidt-Gobel, 1846: 53
Male adult
8,3 mm Thailand, Khao Lak, Khao Lak Beach, at light leg. U.Schmidt, 29.IV.2105; det. M.Baehr, 2015; U.Schmidt, 2015
Copyright © Udo Schmidt

Coptodera (Coptoderina) triloba s.str.
Male adult
Ghana, Atewa Hills, 6*14N 0*33W, 24.3.2010 (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) chaudoiri Andrewes, 1919
Male adult
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) chaudoiri Andrewes, 1919
Endophallus
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) chaudoiri Andrewes, 1919
Male genitalia
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) eluta Andrewes, 1923
Unsexed
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) japonica Bates, 1883
Male adult
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) japonica Bates, 1883
Endophallus
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) japonica Bates, 1883
Male genitalia
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) maculata Dupuis, 1912
Male adult
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) maculata Dupuis, 1912
Male genitalia
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) maculata Dupuis, 1912
Endophallus
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) marginata Dupuis, 1912
Endophallus
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) marginata Dupuis, 1912
Male genitalia
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) occulta Hunting & Yang, 2019 Holotype
Male adult
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) occulta Hunting & Yang, 2019
Male genitalia
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) proksi Jedlicka, 1963
Male adult
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) proksi Jedlicka, 1963
Male genitalia
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) taiwana (Nakane, 1956)
Male adult
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) taiwana (Nakane, 1956)
Male genitalia
Taiwan. Image from Hunting & Yang, 2019
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Coptodera (Coptoderina) triloba minima (Hansen, 1968)
Female adult
det. Anichtchenko A. 2025
Copyright © Igor Solodovnikov

Coptodera (Coptoderina) crockerensis Kirschenhofer, 2014 Holotype
Female adult
Malaysia - Sabah prov. Banjaran Crocker Mts., Gunung Alab peak 30.IV. 27.V.1996, 1650-1800 m, M. Štrba & R. Hergovits leg. (CHZ).
Copyright © Erich Kirschenhofer

Coptodera (Coptoderina) papuella Darlington, 1968 Holotype
Male adult
Image courtesy MCZ
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Coptodera (Coptoderina) congoensis Burgeon, 1937
Male adult
Cameroon 2-5.2.2023, 35km S Kribi (cAA). 6.6 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) eluta Andrewes, 1923a: 30
Female adult
Wao, Lanao del Sur, Mindanao, August 2018 (DUBC)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) eluta Andrewes, 1923
Male adult
N Viet-Nam, 160 km NNW Hanoi, env. Na Hang 26.V.-14.VI.1996 leg. A. Napolov (MSPU).
Copyright © Kirill Makarov

Coptodera (Coptoderina) eluta Andrewes, 1923a: 30
Female adult
Andaman Is. 19.15–38”, “NHMUK 010791022 (BMNH) 6 mm
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) eluta Andrewes, 1923a: 30
Male adult
E Luzon, Quirino, Tapsoy, Sierra Madre, VIII.2014” (DUBC) 6.5 mm
Copyright © Alexander Anichtchenko

Coptodera sapaensis Kirschenhofer, 1994 Holotype
Female adult
Image courtesy Harald Schillhammer (The Vienna Natural History Museum, Austria)
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Coptodera (Coptoderina) eluta Andrewes, 1923a: 30
Male genitalia

Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) luzonensis Jedlicka, 1934 Holotype
Male adult
8.5mm. Image courtesy Jiří Hájek (National Museum of Natural History, Praha, Czech Republic)
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Coptodera (Coptoderina) luzonensis Jedlicka, 1934
Male adult
Sierra Madre, Dismongal, Madela, Quirino, Eastern Luzon, January 2016 (DUBC)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) luzonensis Jedlicka, 1934
Male adult

Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) luzonensis Jedlicka, 1934
Male adult

Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa Schmidt-Gobel, 1846: 55
Male adult
variability of Coptodera flexuosa
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) proksi Jedlicka, 1963d: 341
Male genitalia
Formosa
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) proksi Jedlicka, 1963d: 341
Male adult
“Sulsharyo, Near Mt. Ari, Formosa, 16.V.38”, “NHMUK 010790942” (BMNH).
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) subapicaloides Jedlicka, 1956: 218 Paratype
Male adult
“Kuatun, Fukien / China meridian. / Klapperich”, “Cotype [red label]”, “subapicaloi- / des sp.n. / det. Ing. Jedlička [pink label]”, “Mus. Nat. Prague / 65205 / Inv. [orange label”, “ex coll. A. Jedlička / National Museum of / Prague, Czech Republic” (NMPC).
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) shimianensis Kirschenhofer, 2012 Holotype
Male adult
“China: W-Sichuan / Ya’an Pref., Shimian Co. / Xiaoxiang Ling, side-valley ab. /Nanya Cun nr. Caluo, 11km S / Shimian, 1250m, 7.VII.1999 / leg. A. Pütz”, “Holotypus / Coptodera / (Coptoderina)/ shimianensis sp.n. / des. Kirschenhofer 2012 [red label]”, “Coll. Wrase / Berlin”, “Coll. Wrase / Gusow-Platkow /Germ. [green label]” (cDW).
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) shimianensis Kirschenhofer, 2012
Female adult
MYANMAR: “19809”, “Doherty”, “Birmah, RubyM”, “Fry Coll. 1905.100”, “NHMUK 010791101” (BMNH).
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) shimianensis Kirschenhofer, 2012
Male adult
China, Prov. Yunnan, Nujiang Autonom. Ref., NW Gaoligong Shan, Lushui county, Pianma env., 20.V.2012, leg. Hu (cDW)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) shimianensis Kirschenhofer, 2012
Male genitalia
China, Prov. Yunnan, Nujiang Autonom. Ref., NW Gaoligong Shan, Lushui county, Pianma env., 20.V.2012, leg. Hu (cDW)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa cyanella Bates, 1869
Male adult
“Tjamba, Zuid Celebes, Doherty VIII. IX.”, “NHMUK 010791016 QR code” (BMNH)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa cyanella Bates, 1869
Female adult
“Crocker Range, Sabah, v.1988, Khoo leg.” (cRS)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa cyanella Bates, 1869
Male adult
“Salwatty”, “Bowring, 63.47”, “NHMUK 010791168 QR code” (BMNH)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa cyanella Bates, 1869
Female adult
“Ins. Ternate [Maluku], Doherty VIII”, “Coptodera impicta Chaud., Compared with type H.E.A”, “ŅHMUK 010791315” (BMNH)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa cyanella Bates, 1869
Female adult
“Ins. Ternate [Maluku], Doherty VIII”, “Coptodera impicta Chaud., Compared with type H.E.A”, “ŅHMUK 010791315” (BMNH)
Copyright © Alexander Anichtchenko

Coptodera viridana Jedlička, 1935 Holotype
Male adult
“Type [round label]”, “Philippine Is, / Coll. Bottcher. / B.M. 1929-201”, “Coptodera / viridana / sp.n. / Det. Ing. Jedlička”, “NHMUK 010791080 QR code” (BMNH).
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa subapicalis Putzeys, 1877
Male adult
“Higo”, “Japan, Lewis col” (BMNH)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa philippinensis Anichtchenko, 2025 Paratype
Male adult
“Philippines, E Visayas, Samar, Marabut, xi.2015” (DUBC) 5.8mm
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) proksi Jedlicka, 1963d: 341
Male adult

Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa taiwana (Nakane, 1956)
Male adult
“Taiwan, Taichung City, Beitun, Dakeng No 4 Hiking Trail, 24.175510N, 120.790522E, 525m, 28.II.2018, B.H. Ho lgt. At night” (cAA). 4.5 mm
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa taiwana (Nakane, 1956)
Female adult
“Taiwan, Taichung City, Beitun, Dakeng No 4 Hiking Trail, 24.175510N, 120.790522E, 525m, 28.II.2018, B.H. Ho lgt. At night” (cAA). 5.5 mm
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa cyanella Bates, 1869
Unsexed
“Wallace”, “Moluccas, Jilolo [Jailolo]”, “NHMUK 010791343” (BMNH)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa cyanella Bates, 1869
Unsexed
“Ceram”, “NHMUK 010791202” (BMNH).
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa taiwana (Nakane, 1956)
Male adult

Copyright © Alexander Anichtchenko

Coptodera viridana Jedlička, 1935 Holotype
Male adult

Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa Schmidt-Gobel, 1846: 55
Male genitalia
23, C. viridana, syn. (holotype) = C. flexuosa cyanella Bates, 1869; 24, C. flexuosa cyanellaBates, 1869 (Salawati isl.); 25, C. flexuosa cyanella Bates, 1869 (Sulawesi isl.); 26, C. flexuosa philippinensis (Samar isl.); 27, C. flexuosa subapicalis; 28, C. flexuosa taiwana; 29–30, C. flexuosa s. str. (from the same locality, India)
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa flexuosa
Female adult
“C. Sulawesi, Palolo, 10.2013” (cAA). 6mm
Copyright © Alexander Anichtchenko

Coptodera (Coptoderina) flexuosa flexuosa
Male adult
“N Viet-Nam, 160 km NNW Hanoi, env. Na Hang 26.V.–14.VI.1996 leg. A. Napolov” (MSPU)
Copyright © Kirill Makarov

Coptodera (Coptoderina) flexuosa flexuosa
Female adult
“Roxas, Palawan, Philippines, Nov. 2018” (DUBC) 5.3 mm
Copyright © Alexander Anichtchenko
Coptoderina taxon page (92 photos)
Coptodera (Coptoderinella) congoana Burgeon, 1937
Female adult
Gabon E., Ogooué-Ivindo Prov., Makokou 7.7 km SW Ivindo NP, Ipassa forest research station, 0°30’52N, 12°48’21E, 18-19.VI.2016, 520m, edge of secondary lowland rainforest, MV light, leg. D. Telnov
Copyright © Alexander Anichtchenko

Coptodera (Coptoderinella) congoana Burgeon, 1937
Male adult
Gabon E., Ogooué-Ivindo Prov., Makokou 7.7 km SW Ivindo NP, Ipassa forest research station, 0°30’52N, 12°48’21E, 18-19.VI.2016, 520m, edge of secondary lowland rainforest, MV light, leg. D. Telnov
Copyright © Alexander Anichtchenko

Coptodera (Coptoderinella) martinui Anichtchenko, 2025 Holotype
Male adult
Cameroon 2.–5.2. / 35 km S Kribi 2023 / Martinŭ Ivo lgt.
Copyright © Alexander Anichtchenko

Coptodera (Coptoderinella) martinui Anichtchenko, 2025 Paratype
Male adult
Cameroon 2.–5.2. / 35 km S Kribi 2023 / Martinŭ Ivo lgt.
Copyright © Alexander Anichtchenko

Comparison
Male adult
3, Pronotum of Coptodera (Coptoderinella) congoana Burgeon, 1937 (Gabon, Ivindo NP); 4, Pronotum of Coptodera (Coptoderinella) martinui sp. nov.; 5, Aedeagus of C. (C.) congoana (a—dorsal, b—lateral view; specimen from Gabon); 6, Aedeagus of C. (C.) martinui sp. nov., holotype (a—dorsal, b—lateral, c—ventral view). Scale bar 1 mm
Copyright © Alexander Anichtchenko
Coptoderinella taxon page (5 photos)
Haplocrepis taxon page