Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Zapotecanillus montanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 52miles N of Oaxaca, Ber.202, sink litter, 17 May 1971 S.B.Peck collector (CMNH)
Scale bar = 0.5mm
Copyright © Igor Sokolov

Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro

Austranillus macleayi (Lea, 1906) Syntypes
Female adult

Copyright © Museums Victoria

Pelodiaetus nunni Sokolov, 2019 Paratype
Male adult
NZ, Canterbury, Ahuriri Scenic Reserve. 1.27 mm
Copyright © Igor Sokolov

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Anillus florentinus florentinus Dieck, 1869
Unsexed
Emilia Romagna, Sasseleone (BO), 26.3.2002 350m, A.Degiovanni leg. (cAA)
Copyright © Alexander Anichtchenko

Rhegmatobius strictus (Baudi di Selva, 1891)
Male adult
Sardegna, Gennargentu Mte., Spada Mt., 1000, 27.4.2003 A. Degiovanni leg. (cAA).
Copyright © Alexander Anichtchenko

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Mioptachys flavicauda (Say, 1823)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphanes) lacunarium (Zimmermann, 1869)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Testediolum) armeniacum kokandicum Solsky, 1874: 120
Female adult

Copyright © Kirill Makarov

Bembidion (Asioperyphus) serorum Netolitzky, 1934
Male adult

Copyright © Kirill Makarov

Bembidion (Terminophanes) pulcherrimum (Motschulsky, 1850)
Male adult
Caucasus, Elbrus, Itkol (CTVR). Image from Toledano L. & Gueorguiev B., 2015
Copyright © - - -

Bembidion (Bracteon) argenteolum Ahrens, 1812
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Hydrium) mimbres Maddison, 2020 Paratype
Male adult
USA: New Mexico: Grant Co., Gila River, Billings Vista, 1320 m, 32.8163°N, 108.6032°W, 11.viii.2005. DRM 05.043. D.R. & J.H.A. Maddison
Copyright © David Maddison

Bembidion (Peryphiolus) caligatum Jeanne & Muller-Motzfeld, 1982
Male adult
Spain, Guadarrama, Cotos, 25.8.2001, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Ocys andreae (Jeannel, 1937)
Female adult
Spain, Andalusia, Sierra Nevada, pico Veleta, 2500-2800m, 29-30.9.2001 Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Odontium (Bracteon) conicolle Motschulsky, 1844: 273
Female adult
Burjatia, Barguzin depr., Alla riv., 8.8.1997, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Dimorphopatrobus ludmilae Casale et Sciaky, 1994: 47
Female adult
Tibet, Motuo, Hanmi, 8.2013. det. A. Anichtchenko, 2015 17 mm
Copyright © Alexander Anichtchenko

Platidiolus rufus Chaudoir, 1878
Male adult
Russia, Hamar-Daban Mt. rng., Komarinskiy Mt., upper flow of Slyudyanka riv. 15.7.2013, A. Shavrin leg. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
Copyright © - - -

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Deltomerus (Deltomerus) corax Peyerimhoff, 1922
Male adult
Morocco, Grand Atlas, 2200-2300m, N31.13.09.5 W07.49.29.7 16-17.6.2010 Anichtchenko A. leg. & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Elaphropus) latissimus (Motschulsky, 1851)
Female adult
Kunashiri Isl., Calder of Golovnina Volcano, S coast od Goryachee Lake 43°52'19"N 145°29'43"E 12.VIII.2008 leg.K.Makarov, A.Zaitzev
2.0 mm
Copyright © Kirill Makarov

Elaphropus (Tachyura) ocellatus (Bates, 1892)
Female adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko

Lymnastis galilaeus Piochard de la Brullerie, 1876
Male adult

Copyright © Gianni Allegro

Tachyura ascendens (Alluaud, 1917)
Female adult
TANZANIA, Kilimanjaro, Nkuansa, grassland,1660m, 3.7.2011, R. Felix det. as Elaphropus (Tachyura) exiguus (BOHEMAN 1848)
Copyright © Peter Schüle

Elaphropus (Ammotachys) marchantarius Boyd & Erwin, 2016 Paratype
Male adult
Venezuela, San Carlos de Rio Negro
Copyright © Terry Erwin

Tachys (Paratachys) arcanicola Blackburn, 1878 Paralectotypes
Female adult
Oahu: Honolulu (Bates collection, box 389). Image from Liebherr, 2021
Copyright © James K. Liebherr

Micratopus anaisae Lemaire, 2021 Holotype
Female adult
Saint-Félix, commune du Gosier, Guadeloupe, 16.210 N, 61.461 O, J. M. Lemaire leg. 15/04/2016. (MNHN). Image from Lemaire, 2021
Copyright © - - -

Tachyura (Tachyura) thoracicus (Kolenati, 1845)
Female adult
India, Uttarakhand, 14 km E Uttarkashi, 1500m, 10-12.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr as octostriata (Netolitzky, 1929)
Copyright © Alexander Anichtchenko

Liotachys antennatus Bates, 1871
Female adult
Bolivia: Santa Cruz Prov. Andrez Ibanez Potrerillo de Guenda, Terebinto. / 3-4 October 1994 / leg. P. Bettella / det. R. Davidson. Carnegie Museum of Natural History, Pittsburg, PA, USA
Copyright © Curt Harden

Shiqianaphaenops majusculus Ueno, 1999c: 47 Holotype
Male adult
Cave Feng Dong, Tangshan, Shiqian County, 27°29'10"N/108°15'23"E, 700 m in altitude, Mingyi Tian & Jingli Cheng leg.Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Rakantrechus (Paratrechiama) tofaceus Ueno, 1970b: 106
Male adult
Japan, Kumamoto Ken
Copyright © Vittorino Monzini

Speotrechus mayeti (Abeille de Perrin, 1875)
Male adult
France, mines de fer Privas
Copyright © Vittorino Monzini

Agonotrechus sinotroglophilus Deuve, 2000
Male adult
Hanwang Dong, Zhongbai, Zengjia, Chaotian, Guangyuan, Sichuan (四川省广元市朝天区曾家镇中柏村汉王洞), 32.577297°N, 106.106979°E, 1210 m, 2018-IV-06, leg. Li He (SCAU). Image from Tian & He, 2020
Copyright © - - -

Zhijinaphaenops liuae Deuve & Tian, 2015 Holotype
Male adult
Chine, Guizhou, Guiyang, Xifeng Xian, Shidong Zhen, grotte Hejia Dong, 1198 mètres, 27°2’30,89’’N - 106°31’40,38’’E, Weixin Liu, 13.VII.2013, in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2015
Copyright © Mingyi Tian

Aphaenops (Cerbaphaenops) michaeli Foures, 1954: 4
Male adult

Copyright © Vittorino Monzini

Guizhaphaenopsodes solidior Tian & X. Huang, 2021 Holotype
Male adult
cave Tangle Dong, suburb of Jishou, Xiangxi Tujia & Miao Autonomous Prefecture, west Hunan Province, 28°18’21”N /109°39’15”E, 270 m in altitude, 2018-I-15, leg. Xinglong Huang (SCAU).
Copyright © Mingyi Tian

Velebitaphaenops giganteus Casale & Jalzic, 2012
Female adult
Image from cavebiology.net
Copyright © Creative Commons

Chenotrechus parvulus Tian, Huang & Jia, 2023 Holotype
Male adult
the cave Xiniuyan, Guhang, Luxin, Wuxuan, Laibin, Central Guangxi Zhuang Autonomous Region, Southern China, 109.55°E, 23.56°N, 115 m, 2021-I-15, leg. Mingyi Tian, Weixin Liu & Yi Zhao, in SCAU
Copyright © Mingyi Tian

Aphaenops (Geotrechus) discontignyi Fairmaire, 1867
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -

Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -