Anillinus affabilis (Brues, 1902) Lectotype
Male adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm.
Copyright © Igor Sokolov

Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro

Illaphanus pteridophyticus Giachino, 2005 Holotype
Male adult

Copyright © Alexander Anichtchenko

Pelodiaetodes (Pelodiaetodes) constricticollis Sokolov, 2015 Paratype
Male adult
NZ, Northland, Radar Bush. Image from Sokolov, 2015
Copyright © - - -

Nesamblyops sp.
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Rhegmatobius quadricollis (Ehlers, 1883)
Unsexed
Majella, Abruzzo, Italy
Copyright © Vittorino Monzini

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Anillus florentinus florentinus Dieck, 1869
Unsexed
Emilia Romagna, Sasseleone (BO), 26.3.2002 350m, A.Degiovanni leg. (cAA)
Copyright © Alexander Anichtchenko

Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
Copyright © - - -

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Erwiniana punctisculptilis (Erwin, 1994)
Male adult
Ecuador, Yasuni
Photo of Sokolov I. M.
Copyright © Igor Sokolov

Bembidion (Eurytrachelus) laticolle (Dufischmid, 1812)
Female adult
Orenburg Reg., Ilek Distr., Ilek env., valley of Ural Riv., beach 28.VII-8VIII.2005 S.Vastschenko leg.
L=6.2 mm
Copyright © Kirill Makarov

Bembidion (Semicampa) chaudoirii Chaudoir, 1850b: 179
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Amerizus (Tiruka) gaoligongensis Gueorguiev, 2015 Paratype
Female adult
CHINA, Yunnan, Tengchong County, cave Wu Shi Shan 1-2-3, near vil. Kongshuhe, ca. 2580 m, N25.73111 E98.66459, 11.11.2013, B. Petrov leg.; Scale bar = 1.5 mm.
Copyright © Borislav Gueorguiev

Bembidion (Notaphus) obliquum Sturm 1825: 160
Male adult
Russia, Ивановская обл., Пестяковский р-н, окр. дер. Пурежка, бер. р. Лух 24-25.VI.2011 М. Смирнов. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Trichoplataphus) kara Andrewes, 1921d: 250
Female adult
India, Uttarakhand, 15 km SW New Tehri, 869 m, N30*15'874" E 078*21'587", 18-20.4.2012, A. Anichtchenko leg.; 6 mm; det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) fontinale (Raffray, 1885)
Male adult
Ethiopia Amhara, Mt. Abuna yosef, n-slope, 3800-3950 m, 12°07’52”N 39°11’39”E, 4.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) basistriatum Fairmaire, 1893
Male adult
Ethiopia, Oromia, Mt Enkuolo, NE-slope, 3100-3200m, 07°24’41”N 39°22’26”E, 5-7.Xii.16, leg. J. Schmidt (CSChM)
Copyright © Joachim Schmidt

Bembidion (Geocosmius) elescarabajo Maddison, 2023
Male adult

Copyright © David Maddison

Bembidion (Testedium) laetum Brulle, 1836
Male adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 3.9 mm
Copyright © Kirill Makarov

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Deltomerus (Deltomerus) tatricus L. Miller, 1859: 306
Male adult
N Slovakia; Vysoké Tatry Mts., 10mm
Copyright © Aleš Sedláček

Dimorphopatrobus ludmilae Casale et Sciaky, 1994: 47
Female adult
Tibet, Motuo, Hanmi, 8.2013. det. A. Anichtchenko, 2015 17 mm
Copyright © Alexander Anichtchenko

Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Patrobus septentrionis volgensis Zamotajlov & Isaev, 2006
Male adult

Copyright © Igor Solodovnikov

Deltomerodes ovicollis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Tachyura) ocellatus (Bates, 1892)
Female adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko

Pericompsus (Upocompsus) habitans (Sloane, 1896) Syntypes
Male adult

Copyright © Museums Victoria

Meotachys (Hylotachys) ballorum Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Tapurucuara
Copyright © Terry Erwin

Polyderidius sp.
Male adult
México, San Luis Potosí
Copyright © Terry Erwin

Tachys (Tachys) corax Leconte, 1852
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (incertae) mulwalensis Sloane, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Paratachyta) coracina (Putzeys, 1875)
Male adult
Malaysia, Langkawi, environment of Hotel Berjaya, leg. U.Schmidt, 7.-22.XI.2009; det. M.Baehr, 2010; Photo: U.Schmidt, 2021; 2.1mm
Copyright © Udo Schmidt

Elaphropus (Elaphropus) globulus (Dejean, 1831)
Male adult
Spain, Granada, rio Darro, barranco de Teatino, 20.V.2005, Anichtchenko A. leg & det. (cAA). 1.8 mm
Copyright © Alexander Anichtchenko

Micratopus anaisae Lemaire, 2021 Holotype
Female adult
Saint-Félix, commune du Gosier, Guadeloupe, 16.210 N, 61.461 O, J. M. Lemaire leg. 15/04/2016. (MNHN). Image from Lemaire, 2021
Copyright © - - -

Trechus (Epaphius) nigricornis Motschulsky, 1844
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Perileptus (Perileptus) imaicus Jeannel, 1923
Male adult
N India, Uttarakhand, 10-12.04.2012, Uttarkashi distr., N30*44,999' E078*34,412', 1446m, 14 km E Uttarkashi, Anichtchenko A. leg & det
Copyright © Alexander Anichtchenko

Amblystogenium pacificum (Putzeys, 1869)
Male adult
Ile de la Possesion, Crozet, T.A.A.F, II.1962 Lerri leg. (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Luraphaenops gaudini (Jeannel, 1952)
Male adult

Copyright © Vittorino Monzini

Junnanotrechus wrasei Belousov & Kabak, 2014 Holotype Paratype
Male+Female adult
China (N Yunnan), Dali Bai Nat. Aut. Pref., Diancang Shan, pass 43 km NW Dali, 3104 m, 25 ° 59 ´ 33.5 ´´ N / 99 ° 52 ´ 12.5 ´´ E, (pasture & shrubs, litter, moss and mushrooms sifted), 23.08. 2009, (M. Schülke) ( ZISP)
Copyright © - - -

Cimbrodytes heterognathus Piva, 2021 Holotype
Male adult
Photo Erminio Piva
Copyright © Vittorino Monzini

Trechus (Trechus) minitrechus Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Oromia, Mt Enkuolo NE slope, alt. 3200–3300 m, 07°23′45″ N, 39°22′27″ E, 8 Dec. 2016
Copyright © Joachim Schmidt

Duvalius (Neoduvalius) lohaji Queinnec & Ollivier, 2021 Holotype
Male adult
Mracna Pecina, Livno, Prolog Planina, Bosnia and Herzegovina
Copyright © - - -

Paratrechus obrieni Mateu, 1981
Male adult
Costa Rica, Cerro Buena Vista, 3400 m a.s.l., N 9.5548°, W 83.7552°, 15.IX.2023. Image from Donabauer, 2024
Copyright © - - -

Petraphaenops unguiculatus Delić, Kapla & Colla, 2020 Holotype
Male adult
SLOVENIA, Luče, Raduha, Devettisoča jama (Cave Registry N o. 9000), 46.41231°N, 14.75095°E, found at –450 m (pitfalltrap), collected by T. Delić, 14 June 2017. Image from Delić et al, 2019 (2020)
Copyright © - - -

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -