Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Anillinus langdoni Sokolov & Carlton, 2004
Male adult
USA-TN, Sevier Co, GSMNP, secondary site, 490 m, ex leaf et log litter, 35°43.157' N, 83°23.309' W, 19.04.2011, J.-S. Park det. I. Sokolov, 2013
Copyright © Roman Dudko

Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro

Typhlocharis laurentii Magrini, 2000 Paratype
Unsexed
España, Almeria, Sierra Almagrera ; carretera N332, 30.XI.98, leg. Fancello (MNHN, Antoine Mantilleri, 2016)
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Scotodipnus alpinus Baudi di Selve, 1871
Female adult
Piemonte, Valle Po, Crissolo (CN), 27.V.2000, 1250m, fagus, leg. F. Angelini
Copyright © Alexander Anichtchenko

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Anillus florentinus florentinus Dieck, 1869
Unsexed
Emilia Romagna, Sasseleone (BO), 26.3.2002 350m, A.Degiovanni leg. (cAA)
Copyright © Alexander Anichtchenko

Medusapyga hehalis LaBonte, 2023 Paratype
Female adult
WASHINGTON, Thurston Co., 11.3 km NW Littlerock, elev. ~400 m, D.J. Cox and J.R. LaBonte, 6 May 2007, 46.9600°N 123.13450°W. Image from LaBonte & Maddison, 2023
Copyright © - - -

Megastylulus pivai Giachino & Sciaky, 2003 Holotype
Male adult
leg. E. Piva - Isl. St.Lucia Caraibi
Copyright © Vittorino Monzini

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Furcacampa) versicolor (Leconte, 1848)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Actedium) pallidipenne (Illiger, 1802b)
Male adult
length: 4,1-4,7 mm, Niederlande, Maasvlakte, leg. A.Teunisse, 1989; det. G.Rößler, 1996, Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Bracteon) foveum Motschulsky, 1844: 271
Male adult
Khabarovsk Reg., bank of Amur Rin. in Khabarovsk city 48°28'33"N 135°02'39"E 7.VIII.2003 I.Melnik leg.
L=6.0 mm
Copyright © Kirill Makarov

Bembidion (Philochthus) biguttatum (Fabricius, 1779)
Male adult

Copyright © Kirill Makarov

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult
4,4 mm
Namibia, Etosha National Park, Okaukuejo, 1400 mб leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012
Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Bembidion (Antiperyphanes) hirtipes (Jeannel, 1962)
Male adult
Argentina, Mendoza, Pampa Palauco. Image from Maddison D.R., 2014.
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Bembidion (Jammuphanes) sinuosum Schmidt & Marggi, 2014 Paratype
Male adult
Pakistan, Kaghan valley, Shogran, 3050m, 19.6.2001. Imagr from Schmidt & Marggi, 2014
Copyright © - - -

Neolovricia ozimeci Lakota, Jazlic & Moravec, 2009 Holotype
Female adult
Spilja u Radinovcima, Dedici, Rastovac, Biokovo, 25.10.2006 Coleoptera (1) Leg: Roman Ozimec. Deposited in the Croatian Natural History Museum, Zagreb. Image from Lakota, Jazlic & Moravec, 2009
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Sinechostictus (Pseudolimnaeum) cf. pendleburyi (Andrewes, 1931)
Female adult
Philippines, N Luzon, Mt. Province, Mt. Polis 7.2014. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) marginipenne Solsky, 1874: 130
Male adult
Uzbekistan, Chatkal Mt. rng., Yangiobod, left conf of Dukentsay riv, 41.07.28N 70.06.21E 1300m 2.8.2021 Anichtchenko A. leg. (cAA). det. L. Toledano, 2026; 3.8 mm
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Platidiolus vandykei Kurnakov, 1960
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Deltomerus (Deltomerus) pseudoplatynus pseudoplatynus s.str.
Female adult
NW-Caucasus,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
long - 11mm.
Copyright © Ruslan Panin

Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Penetretus andalusicus (Reitter, 1897)
Male adult
Spain, Sierra Nevada, Lagunillos de la Virgen, 2850m, 18.5.2005, A. Anichtchenko leg. & det (cAA)
Copyright © Alexander Anichtchenko

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Tachys (Paratachys) micros (Fischer von Waldheim, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Stigmatachys uvea Boyd & Erwin, 2016 Holotype
Female adult
Perú, Loreto, Campamento San Jacinto
Copyright © Terry Erwin

Tachys (incertae) mulwalensis Sloane, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) terryli Liebherr, 2021 Paratype
Female adult
Kauai: Halelea F. R.: Namolokama Mtn., Waioli Str., litter, sift, ohia/ferns, 22°08.00'N, 159°29.85'W, 1340 m el., 21-v-2005, lot 1 Liebherr
Copyright © James K. Liebherr

Tachyta (Eurytachyta) laticollis Baehr, 2016 Holotype
Female adult
Borneo, Sabah, Kinabalu NP, HQ vic., 1550 m, 22.–25.V.2005, R. GRIMM (CBM).
Copyright © Martin Baehr

Tachyta (Paratachyta) quadrinotata Baehr, 2016 Holotype
Female adult
320 Ilam Distr., betw. Hodia Khola Valley to Sktia, 250–500 m, Shorea forest, dry, 7 Apr 1988, MARTENS & SCHAWALLER leg. / Nepal-Expeditionen JOCHEN MARTENS (SMNS).
Copyright © Martin Baehr

Aenigmatachys rubens Baehr, 2020 Holotype
Male adult
“Australia Black B . . . (unreadable) / G.C. Champion Coll. B.M. 1927-409” (NHM).
Copyright © Martin Baehr

Tachyura (Tachyura) thoracicus (Kolenati, 1845)
Female adult
India, Uttarakhand, 14 km E Uttarkashi, 1500m, 10-12.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr as octostriata (Netolitzky, 1929)
Copyright © Alexander Anichtchenko

Trechus (Microtrechus) tuckaleechee Barr, 1962
Male adult
Great Smoky Mountains National Park
Copyright © Igor Sokolov

Epaphiopsis (Epaphiama) jacobsoni Sokolov et Shilenkov, 1987: 10
Male adult
Altai Republic: Holzun mt.rng., Bannaya riv., 1300-1600 m,12-14.06.1999, R.Yu & A.Yu. Dudko leg. & det.
Copyright © Roman Dudko

Dianotrechus gueorguievi Tian, 2016 Holotype
Female adult
China, Yunnan Province, Ermu Vill., Kunming District, Dashi Dong (Big Rock Cave), 24°49'13"N/102°27'56"E, 1940 m in altitude, XI-8- 2011, B. Petrov leg.” (SCAU). Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Geotrechus orpheus Dieck, 1869
Male adult
France, Ariege, Aven de Santhe Catherine, VIII.1981. Leg. Sciaky (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Sporades (Perileptosporades) jaechi Liebherr, 2020 Holotype
Male adult
NEW CALEDONIA (NC 27) / N-Prov., Poum Com. / unnamed river / 9.XI.2016, leg. M.A. Jäch // ca. 7 km NE Poum / ca. 15 m a.s.l. / 20°12'18.3"S 164°5'27.7"/ almost dried out lowland river (NHMW).
Copyright © James K. Liebherr

Haixiaphaenops jinxiaohongae Tian, Cheng & Huang, 2021 Paratype
Male adult
Guizhou, Qingzhen, Anliu, Ximi, Changtu Dong cave (贵州省清镇市暗流镇长土洞), 26°51‘N, 106°21‘E, 1249 m, 2020–VI–27, leg. Chenggang Wang, Xiaohong Jing, Guangyuan Cheng, Yi Zhao & Mingyi Tian, SCAU.
Copyright © Mingyi Tian

Trechus (Trechus) mattisi Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Bale Mts, Harenna Forest, E Rira, alt. 3150 m, 06°44′51″ N, 39°46′40″ E, Feb. 2007
Copyright © Joachim Schmidt

Ushijimaella uenoi P. Moravac et Wrase, 1998: 213 Paratype
Female adult
China, Shaanxi, Qin Ling Shan, 107.56E, 35.45N, autoroute km 93 S of Zhouzhi, 108km SW Xian Mountain Forest, sifted, 1650m, 1-2.9.1995, leg. A. Putz (cAA)
Copyright © Alexander Anichtchenko

Aspidaphaenops qujingensis Deuve & Tian, 2020 Holotype
Male adult
Chine, Yunnan, Préfecture de Qujing, Luoping Xian, Machang, grotte Shuiyuan Dong, 1530 mètres, 24°49’33’’N 104°21’48’’E (Tian M. Y., Lui W. X., Wang X. H., Tang M. R. leg., 18.VI.2015), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve

Microblemus rieae Ueno, 2007 Holotype
Male adult
Image from Naito, 2018
Copyright © - - -

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
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Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
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Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

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