Pterostichus (Adelopterus) ambiguus (Fairmaire, 1858)
Female adult
France, 20224, Corse, Calacuccia forêt de Valdu Niellu 12-09-2000
Copyright © Pascal Deschamps

Pterostichus (Adelosia) macer (Marsham, 1802)
Male adult
Latvia, Kuldiga Co. 2019
Copyright © Alexander Anichtchenko

Pterostichus (Anilloferonia) malkini (Hatch, 1953)
Male adult
Image from Labonte, 2013
Copyright © - - -

Pterostichus (Anomostichus) anomostriatus Sciaky, 1995 Paratype
Male adult
China – pr. Sichuan, Emei mt. 2500 4-20.V.1989 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Aphaonus) compressus Rost, 1892
Male adult
Russia, Krasnodar reg., 6 km NE Soloniki vill.,
Sochi Nat.Park, 28-30.V.2009 Chibilov E. leg.
photo A. Anichtchenko
det.I.A.Solodovnikov
Copyright © Alexander Anichtchenko

Pterostichus (Aphaonus) cylindriformis Reitter, 1887e: 252
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Aphaonus) mariamae Zamotajlov, Solodovnikov & Fominykh, 2015 Holotype
Male adult
Abkhazia, Gulripsh Distr., Tsebelda Plateau near village Ablukhvara, left bank of River Kelasur, slopes of Mt. Bashlug, 650 m, 15.05.-19.07.2012, pitfall traps (D. Fominykh) (ZISP).
Copyright © Kirill Makarov

Pterostichus (Argutor) commutabilis (Motschulsky, 1866)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Argutor) leonisi Apfelbeck, 1904: 257
Male adult

Copyright © Igor Solodovnikov

Pterostichus (Argutor) leonisi Apfelbeck, 1904: 257
Female adult

Copyright © Igor Solodovnikov

Pterostichus (Asioplatysma) medvedevi Kryzhanovskij & Mikhailov, 1972: 319
Male adult

Copyright © Kirill Makarov

Pterostichus (Badistrinus) laticollis (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Badistrinus) haptoderoides haptoderoides
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult

Copyright © Kirill Makarov

Pterostichus (Bothriopterus) aeneocupreus (Fairmaire, 1887)
Male adult
Bhutan, near Gangcheyting, [Camp 2] 3689 m, 27° 24 14.0''N, 90° 14' 04.2''E, 9.oct. 2017
Copyright © Joachim Schmidt

Pterostichus (Calopterus) pilosus (Host, 1789)
Male adult

Ukrainian Carpathians,
Transcarpathian Reg., Rakhov Distr., Chornohora range h~1850m, 29.VI-28.VII.2007, Panin R. leg.
male,18 mm.

Copyright © Ruslan Panin

Pterostichus (Calopterus) selmanni prevostii (Dejean, 1828)
Male adult
France, Jura, Lajoux, 1190m, 20/05/2013, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Pterostichus (Calopterus) selmanni roubali Schauberger, 1927
Female adult
CZ; SW Bohemia, Šumava Mts.; 16mm
Copyright © Aleš Sedláček

Pterostichus (Calopterus) pilosus pilosus s.str.
Female adult
CZ; NE Moravia, Štramberk; 15mm
Copyright © Aleš Sedláček

Pterostichus (Calopterus) pilosus pilosus (Host, 1789)
Male adult
W Ukraine, NE Carpathians,, Chornagora Mt Range, h=2000m, alpin zone, 11.VI-1.IX.2015, Panin R. leg 16 mm
Copyright © Ruslan Panin

Pterostichus (Calopterus) selmanni selmanni (ab.niger Schilsky, 1888)
Female adult
Austria, Upper Austria, Pyhrnpass Umgebiet, h=1250 m (Pyhrn pass area) Wurzeralm, 6-18.VI.2024, M. Fuchs leg., L= 18.5 mm
Copyright © Ruslan Panin

Pterostichus (Cheporus) burmeisteri Heer, 1838
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m 9.VI-25.VII.2007 Panin R. leg.
male,15 mm.

Copyright © Ruslan Panin

Pterostichus (Cheporus) burmeisteri baldensis Schaum, 1862
Female adult
NE Italy, Veneto, Mts Lessini, Vallone del Malera h=1500 m., 28-30.V.2019, Gontarenko A. leg./det.Panin R. L=14 mm
Copyright © Ruslan Panin

Pterostichus balthasari Jedlička, 1937 Lectotype
Male adult
(NMPC) Image from Dorjderem S, Shi H, Liang H., 2020
Copyright © - - -

Pterostichus (Circinatus) bullatus Allegro & Sciaky, 2010
Male adult

Copyright © Gianni Allegro

Pterostichus (Circinatus) dentifer Allegro & Sciaky, 2010 Holotype
Male adult
China, C Sichuan, Dayi distr., / Chadiping env., 1500 m / 5–7.08.1996 / A. Zamotajlov, A. Miroshnikov & / D. Fedorenko. Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) bullatus Allegro & Sciaky, 2010
Female adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) beneshi Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Wolong 150 km NW Chengdu, 9.VII.1994 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Circinatus) tonkinensis Straneo, 1980
Male adult
Vietnam, 40 km W of Cao Bang, Phia Oac Mt., 22°36´50´´N 105°52´0´´E, h~1800 m, deciduous bamboo forest, 3–11.X.2018 (D. Fedorenko)
Copyright © Dmitri Fedorenko

Pterostichus (Circinatus) medogensis Shi & Wang, 2026 Paratype
Male adult
“CHINA, Xizang, Mêdog county, 62 km at road Bomê to Mêdog, mixed forest, 2794 m, N29.70937 E95.59230”, “2012.VII.27, by pitfall trap, LIANG Hongbin, SHI Hongliang, LIU Ye, WANG Yanan leg., Institute of Zoology, CAS”. Image from Wang et al, 2026
Copyright © - - -

Pterostichus (Cophosus) cylindricus (Herbst, 1784)
Female adult
CZ; S Moravia, Břeclav; 20mm
Copyright © Aleš Sedláček

Pterostichus (Cryobius) kaninensis Poppius, 1906
Male adult

Copyright © Kirill Makarov

Pterostichus (Cryobius) pinguedineus Eschscholtz, 1823
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Cryobius) middendorffi J.R. Sahlberg, 1875: 103
Female adult

Copyright © Kirill Makarov

Pterostichus (Cryobius) middendorffi J.R. Sahlberg, 1875
Male adult
Магаданская обл., верховья р. Ола, снежник c Salix reticulata h~1127 60°39'N 151°16'E 7-12.VIII.2011 leg. О.Макарова,А.Бабенко. det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) quadraticollis Chaudoir, 1846a: 140
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) pumilio (Dejean, 1828)
Male adult
Spain, Navarra, Sierra de Aralar, 9.5.2006, Altxueta, 1000m, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) aralarensis Espanol & Mateu, 1945
Female adult
Spain, Mts, Cantabricos, Soto de Sajambre, Canto Cabronero Mt., 1850m, 12.VIII.2008, Anichtchenko A. leg. & det. 7.5mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) homalonotus sojot Shilenkov, 2000 Paratype
Female adult
Бурятия, Восточный Саян, окр. влк. Кропоткина 2050м, 10-15.7.98, Aнищенко А. leg. (cAA) 7.8mm [Buryatia, W Sayan mts., volcan Kropotkin vic., Anichtchenko A. leg]
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) fulvescens (Motschulsky, 1844)
Male adult
Burjatia, W Sajan Mts., Tunkinskie Goltsy, Mondy vill. vic., 1500m, 1-7.7.2001, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) riparius asiaticus Poppius, 1906
Female adult
Far East, Khabarovsk reg., Verhnebureinskii distr., Bureinskii Nat. Res., kordon Strelka, 23-25.VII.2006, 51.38'37"E, 134.15'41"E, A. Barsevskis leg. (DUBC). 8.5 mm; det. Yu. Sundukov, 2025
Copyright © Alexander Anichtchenko

Pterostichus (Cylindrocharis) rostratus (Newman, 1838)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Eosteropus) aethiops (Panzer, 1796)
Female adult
W-Ukraine, Lviv reg., Zhydachiv distr., near Vybranivka vill., forest, under the bark, Panin R. leg. 12.X.2007 female,14mm
Copyright © Ruslan Panin

Pterostichus (Eosteropus) rufitarsis cordatus Letzner, 1842
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2000m., Panin R. leg., 29.VI-28.VII.2007 L=12 mm
Copyright © Ruslan Panin

Pterostichus (Eosteropus) discrepans A.Morawitz, 1862
Female adult

Copyright © Kirill Makarov

Pterostichus (Eosteropus) orientalis (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Eosteropus) tuberculiger tuberculiger
Male adult
S Primorie, Ussuriisk distr., 3km NE Kaimanovka vill. 22-25.8.2007, K. Makarov A.Zaitsev leg. (DUBC) 13mm
Copyright © Alexander Anichtchenko

Pterostichus (Eosteropus) virescens (Gebler, 1833)
Female adult
NE Altay Mts., Altaiskii res., S slope Korbu mt. rng., Kokshi, 650m, Teletskoe lake 4.7.2002. 11.5mm
Copyright © Alexander Anichtchenko

Pterostichus (Epinialoe) fumikoae Y.Kurosa, 2019 Holotype
Male adult
Kiyozasa-tôge, Fujieda-shi, 11.XI.2017, Y. Kurosa leg. (National Museum of Nature and Science, Tsukuba). Image from Kurosa, 2019
Copyright © - - -

Pterostichus (Ethira) pilifer (Bates, 1878)
Male adult
Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -

Pterostichus (Falsargutor) pseudopedius pseudopedius Reitter, 1887
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Feronidius) ? hungaricus (Dejean, 1828)
Male+Female adult
W. UKRAINE, Transcarpathian region, Mukachevo Distr., riverine forest near Znyatsevo vill., 16-31.VII.2008, Panin R. leg., female, 16 mm
Copyright © Ruslan Panin

Pterostichus (Feronidius) melas (Creutzer, 1799)
Male adult

W-Ukraine,
Lviv Region, Zolochiv distr., near Gologirky, in steppe, 16-29.V.2007 Panin R. leg.
long-17mm

Copyright © Ruslan Panin

Pterostichus (Feronidius) fornicatus (Kolenati, 1845)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Feronidius) hungaricus (Dejean, 1828)
Male adult
Romania, Mts Trascau, Rimetea env., Mt Piatra Secuiului h~1100 m, 27.V-11.VIII.2008, I.Retezár & B.Tallosi leg. L=15 mm
Copyright © Ruslan Panin

Pterostichus (Gutta) kongshuhensis Gueorguiev, 2015 Holotype
Male adult
CHINA, Yunnan Prov., Tengchong County, vil. Kongshuhe near cave Wu Shi Shan 1-2-3 N25.73111 E98.66459, ca. 2500-2600m, 11.11.2013, B. Petrov leg.
Copyright © Borislav Gueorguiev

Pterostichus (Haplomaseus) caucasicus (Menetries, 1832)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Huaius) hanwang Tian & He, 2020 Holotype
Male adult
cave Hanwang Dong, Zhongbai, Zengjia, Chaotian, Guangyuan, Sichuan (四川省广元市朝天区曾家镇中柏村汉王洞), 32°34′40.14″N, 106°6′25.32″E, 1210 m, IX-22-2018, leg. Li He (SCAU). Image from Tian & He, 2020
Copyright © Mingyi Tian

Pterostichus (Huaius) changgui Jiang & Zeng, 2025 Paratype
Male adult
CHINA, Chongqing City, Chengkou County [城口县], Miaoba Town [庙坝镇], Guannei Village [关内村], Laoying Cave [老鹰洞], N 31°55'39.51", E 108°30'22.22", alt. 700 m, 11-IV-2025, Yu-Yang Zhou & Fan-Hao Zeng leg. Image from Jiang & Zeng, 2025
Copyright © - - -

Pterostichus (Hypherpes) tristis (Dejean, 1828)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Hypherpes) lama (Menetries, 1843)
Female adult
USA, WA, Clark Po near Vancouver Lake forest, 2.VI.2012. 25 mm
Copyright © Alexander Anichtchenko

Pterostichus (Hypherpes) lama (Menetries, 1843)
Male adult
USA: Washington, 34 mm
Copyright © Igor Solodovnikov

Pterostichus (Hypherpes) lama (Menetries, 1843)
Female adult
USA: Washington
Copyright © Igor Solodovnikov

Pterostichus (Koreonialoe) microps Heyden, 1887
Male adult

Copyright © Kirill Makarov

Pterostichus (Koreonialoe) micropoides Yin, Shi & Liang, 2021 Holotype Paratype
Male+Female adult
Paratype from Hunchun city, Jilin
Copyright © - - -

Pterostichus (Koreonialoe) ishikawai Nemoto, 1988
Female adult
Sud Korea, Juwangsan National Park
Copyright © Vittorino Monzini

Pterostichus (Galapterus) galae Farkac & Plutenko, 1996
Male adult

Far East, S Sikhote-Alin, Snezhnaya Mt., source of Ussuri Riv., h = 1400 -1682 m. 1-6.07.2004 Yu.Sundukov leg.
male
L=13,0 mm

Copyright © Maksim Smirnov

Pterostichus (Lenapterus) punctatissimus (Randall, 1838)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Lenapterus) vermiculosus (Menetries, 1851)
Male adult

Copyright © Kirill Makarov

Pterostichus (Lenapterus) agonus G.Horn, 1880
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Lenapterus) subrugosus Straneo, 1955
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Leptoferonia) pumilus Casey, 1913
Female adult
Oregon, Portland

Copyright © Igor Sokolov

Pterostichus (Leptoferonia) hackerae Will, 2016 Holotype
Male adult

Copyright © Kipling W. Will

Pterostichus (Lianoe) dufourii boisgiraudii L. Dufour, 1843
Male adult
France, Pyrenee Ariege, Vall. du Biros, Eylie Cirque de la Plagne, 1100-1500m, 20.6.1993 (DUBC). det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Pterostichus (Lianoe) arribasi Ortuño, 1991
Male adult
Ezcaray, 16.8.97, Valdezcaray, La Rioja, Spain, M. Toribio leg. & det. (cAA). 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Licentius) cuii Tian & He, 2021 Holotype
Male adult
cave Tangren Dong, Beijiao Cun, Fozizhuang Xiang, Fangshan Qu, Beijing, 39° 47ʹ42ʺN 115°51ʹ55ʺE, 324 m, 20.XII.2020, leg. Qingwu Cui (SCAU).
Copyright © Mingyi Tian

Pterostichus (Lyropedius) lyroderus (Chaudoir, 1846)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Lyrothorax) caspius (Menetries, 1832)
Male adult
CZ, SE Moravia, Břeclav; 11mm
(introduced)
Copyright © Aleš Sedláček

Pterostichus (Incertae) zhejiangensis Kirschenhofer, 1997 Holotype
Male adult

Copyright © Kipling W. Will

Pterostichus (Melanius) castor Goulet & Bousquet, 1983
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Melanius) aterrimus aterrimus
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Melanius) aterrimus aterrimus (Herbst, 1784)
Male adult
W Ukraine, Lviv, Holosko, bank of Natural lake, 24.V-8.VII.2016, Panin R. leg. L=15 mm.
Copyright © Ruslan Panin

Pterostichus (Melanius) elongatus (Duftschmid, 1812)
Female adult
Marruecos, Gada de Debdou, 9.6.1993 (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) orion (Tschitscherine, 1901)
Male adult

NE Chita reg., Kodar Mt rng., Chara riv.
15 km N Leprindo, 2200m 5-15.7.2001 Brinev leg.
14 mm

Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) mirus (Tschitscherine, 1894)
Female adult
Russia, Siberia, SW Tuva, pass Barlyk, 2100m, 10.VII.2010
14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) interruptus (Dejean, 1828)
Male adult

Copyright © Kirill Makarov

Pterostichus (Metallophilus) rugosus (Gebler, 1823)
Male adult
Russia, Chita reg., Malye Kovali vill. vic., near Chichatka riv., 22-29.7.1999, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Micronialoe) chogyesanus Park; Kwon & Lafer, 1996
Male adult
male
Copyright © Dooyoung Kim

Pterostichus (Micronialoe) spinulosus Kim & Suh, 2024 Holotype
Male adult

Copyright © Dooyoung Kim

Pterostichus (Monoferonia) diligendus (Chaudoir, 1868)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Feroperis) vladivostokensis Lafer, 1979 : 13
Male adult

Copyright © Kirill Makarov

Pterostichus (Morphnosoma) petulans Jedlicka, 1938
Male adult
Russia, Primorskii krai, Shkotovskii distr., Anisimovka vill., 17-26.6.2008 (DUBC). 15 mm. det. A. Anichtchenko, 2021
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) jungens (Tschitscherine, 1893)
Male adult
Russia, S Primorie, Chernye Mts., valley of Poima riv., 30.7.1999, A. Anichtchenko leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) melanarius cardioderus (Chaudoir, 1846)
Female adult
Georgia, Racha distr., near Okureshi vill, Khvamli Mt h=1000-1100 m, 19-24,VI.2024, I.Uspenskyi leg./det. R Panin, L=22 mm
Copyright © Ruslan Panin

Pterostichus (Morphohaptoderus) schuelkei Sciaky & Wrase, 1997: 1102 Holotype
Female adult
body length = 10.8 mm, “CHINA (Shaanxi) / Qin Ling Shan / 108.47 E / 33.51 N / Mt. W pass / autoroute km 70,47km. S Xian 2350–2500 m / 7.–19.VII.1996 / Kleinfeld & Schutze”; “COLL.WRASE / BERLIN”; “HOLOTYPUS / Pterostichus / (Circinatus)/ schuelkei sp. n. / Sciaky & Wrase des. 1997” [red label]
Copyright © Hongliang Shi

Pterostichus (Morphohaptoderus) saueri Sciaky, 1994
Male adult
China, Sichuan, Ganzi pref, Daxue Shan 102.00E 30.03N 5km E Kangding, river valley 3000m 20-23.5.1997 leg A. Putz (cAA) 6.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Morphohaptoderus) glabellus Fedorenko, 2023 Paratype
Female adult
N[orthern] Vietnam, Lao Cai Prov[ince]./ Hoang Lien Son Mt.ridge,/ env. Fansipan [= Phan Xi Pang] Mt, Tram Ton/ h = 1950-2100 [m],/ 15-30./ leg. VII.2007
Copyright © Dmitri Fedorenko

Pterostichus (Morphohaptoderus) unisetosus Fedorenko, 2023 Holotype
Male adult
(ZMMU): N-Vietnam, 40 km WNW of/ Lao Cai, env. Y Ty, Bat Xat/ Nat[io]n.[al] Park, h=1600-1900 m/ 22°37´-37.6´N/ 103°37.3´-38.3´E, 16-21./ leg. D. Fedorenko X.2018
Copyright © Dmitri Fedorenko

Pterostichus (Morphohaptoderus) huashanus Sciaky, 1994
Female adult
7.5 mm (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Morphohaptoderus) quyuani Wu, Chen & Shi, 2025 Holotype Paratype
Male+Female adult
Hubei Province, Shennongjia, Hongping Town, Wenshui Village, 1700 m, pitfall trap, 2013.VIII.13, Hao Huang leg.” [in Chinese]. Image from Wu, Chen & Shi, 2025
Copyright © - - -

Pterostichus (Morphohaptoderus) dentellus Facchini & Sciaky, 2003
Male+Female adult
Hubei Province, Shennongjia. Image from Wu, Chen & Shi, 2025
Copyright © - - -

Pterostichus (Myosodus) rudestriatus (Reitter, 1883)
Female adult
S Russia, NW-Caucasus, Karachay-Cherkessia, Teberda, Alibek Mountain Pass h=2800 m, 23.VII.2010, Solodovnikov I.A leg. <br L=16 mm</p>
Copyright © Ruslan Panin

Pterostichus (Myosodus) lacunosus roubalianus (Lutshnik, 1928)
Female adult
S Russia, W Caucasus, <br /> Tuapse distr., Kashina (=Lysaya) Mt. h=1300m., 10.V.2006, Solodovnikov I.A. leg.<br /> Female, 18 mm
Copyright © Ruslan Panin

Pterostichus (Myosodus) schoenherri Faldermann, 1836
Male adult
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Rng, Zekari Pass env. h~2100-2200m, 26-30.VI.2006, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin

Pterostichus (Myosodus) zamotajlovi Belousov, 1991
Male adult
Russia, Karachaevo-Cherkessia, N slopes of Sophia Mt., upper of Sophia riv., h = 2700 m, 07-25.VII.2020 leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Pterostichus (Myosodus) aibgensis adelaidae Reitter, 1899
Male adult

Copyright © Igor Solodovnikov

Pterostichus (Myosodus) aibgensis adelaidae Reitter, 1899
Female adult

Copyright © Igor Solodovnikov

Pterostichus (Myosodus) sodalicius Heyden, 1885
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. & det. (cAA). 15mm
Copyright © Alexander Anichtchenko

Pterostichus (Myosodus) schoenherri Faldermann, 1836
Female adult
Georgia, Lesser Caucasus, Imereti prov., Meskheti Mt Rng, N slope, Zekari Pass h~2150 2220 m alpine meadows 8-9.VII.2021, E.Rujan leg./det. R. Panin
Copyright © Ruslan Panin

Pterostichus (Myosodus) lacunosus (Chaudoir, 1844)
Male adult
S Georgia, Samtskhe-Javakheti, Akhaltsikhe distr., Atskuri h=1200-1350 m, 1-7.VII.2013, E.Rutjan leg., L=19.5 mm
Copyright © Ruslan Panin

Pterostichus (Myosodus) starcki Heyden, 1885
Female adult
L=20 mm
Copyright © Ruslan Panin

Pterostichus (Neohaptoderus) catei Sciaky & Wrase, 1997 Paratype
Male adult

China,Shaanxi, Qin Ling Shan, 110.06E 34.27N Hua Shan Mt., N Valley, 1200-1400m 118 km E Xian, sifted 18-20.08.1995 A.Pütz leg.
10 mm
photo & coll. Anichtchenko A. 2009

Copyright © Alexander Anichtchenko

Pterostichus (Neohaptoderus) austrinus Fedorenko, 2023 Holotype
Male adult
(ZMMU), labelled: N-Vietnam, 40km W of/ Cao Bang, Phia Oac Mt./ 22°36´25N 105°52´08E/ h~1650-1700 m,/ deciduous forest, 3-11.X./ leg. D.Fedorenko 2018
Copyright © Dmitri Fedorenko

Pterostichus (Neohaptoderus) ignavus (Tschitscherine, 1897)
Male adult
(cAA). 9.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. Holotype
Male adult
Mt. Daisen, Tottori Pref., Japan, 14. VII. 1956, T. Fujimura
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-041
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) fujisanus Tanaka & Suga, 1972 Holotype
Male adult
Mt. Fuji (south side), 1400 m, Shizuoka Pref., Japan, 31. VII. 1971, K. Tanaka & S. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Oreophilus) variolatus variolatus Dejean, 1828
Male adult
S Slovenia, Ilirska Bistrica, Snežnik Mts,15-28.VI.2009, Manfred Egger leg.
L=19 mm

Copyright © Ruslan Panin

Pterostichus (Oreophilus) multipunctatus (Dejean, 1828)
Female adult

Copyright © Kirill Makarov

Pterostichus (Oreophilus) externepunctatus Dejean, 1828
Female adult
NW-Italy, Piemonte prov.,Cottian Alps, Mt Albergian, 11.VI.1979, S. Dacatra leg., L=15 mm
Copyright © Ruslan Panin

Pterostichus (Oreophilus) jurinei (Panzer, 1803)
Female adult

Copyright © Kirill Makarov

Pterostichus (Oreophilus) bicolor Aragona, 1830
Male adult
H. Penna, Senova, 1400m 18.6.1978 (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Oreophilus) impressus (Fairmaire & Laboulbene, 1854)
Male adult
France, Orvault, Loire-Atlantique, 16.12.1962 [label error] (DUBC) 17 mm
Copyright © Alexander Anichtchenko

Pterostichus (Oreoplatysma) cecchiniae Jakobson, 1907
Male adult

Russia, Krasnodar reg., 6 km NE Soloniki vill.,
28-30.V.2009 Anichtchenko A. leg.
male, long. 10 mm

Copyright © Alexander Anichtchenko

Pterostichus (Oreoplatysma) koenigi (Reitter, 1887)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Orientostichus) pulcher Sciaky & Allegro, 2013 Holotype Paratype
Male+Female adult
Label of Holotype male: China, S Sichuan, Ebian, Yizhi Yakou, Xiaoliang Shan-Si He, 2800-3100 m, 19.VII.2007, Cavazzuti P.F. leg. (IZAS). Scale bar 5 mm
Copyright © Gianni Allegro

Pterostichus (Orientostichus) perlutus Jedlicka, 1938a: 5
Male adult
W Sichuan, pass W Kangding, Zheduo mts, 4200m, 12.7.2013, V. Patrikeev leg. (cAA) 16 mm; det. H. Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Orientostichus) jialini Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023
Copyright © - - -

Pterostichus (Orientostichus) pemphis Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023
Copyright © - - -

Pterostichus (Orsonjohnsonus) johnsoni Ulke, 1889
Male adult

USA, Oregon, Clackamas County

Copyright © Igor Sokolov

Pterostichus (Paraferonia) lubricus LeConte, 1852
Male adult

Great Smoky Mountains National Park

Copyright © Igor Sokolov

Pterostichus (Paranialoe) biexcisus subparallelus Straneo, 1955
Male adult
Nishinoshima Is., Mita (Mt. Takuhi-yama). Image from Takashi Shimizu and Ryo Nakamura, 2023
Copyright © - - -

Pterostichus (Parapterostichus) grajus (Dejean, 1828)
Male adult
Italia, Oropa (Bi) Lago Mucrone, 14.6.2003 S. Dacatra leg. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) tomensis (Gebler, 1847)
Male adult

Kemerovo Reg., env. Osinniki, source of Krasnaya Kaltank Riv., Populus tremulae-fores, W slope , h = 450 м 28.V-6.VI.03 Yu.Sundukov leg.
L=18,0 mm

Copyright © Maksim Smirnov

Pterostichus (Petrophilus) turanensis Jedlicka, 1959
Male adult
Russia, E Sayan Mts., Tunkinskie Goltsy, Kyngarga riv., 2200 m, 18-23.VII.1995, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) montanus (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) tundrae (Tschitscherine, 1894)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) triseriatus (Gebler, 1847)
Male adult
Tyva, Kyzyl-Taiga mt. rng., 1500m, Ak-Sug riv. 21.6.2006, A. Menshikov leg. (cAA) det. A. Anichtchenko, 2021; 10.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) subaeneus (Chaudoir, 1850)
Male adult
Russia, E Siberia, Irkutsk reg., Nizhneudinsk distr., 9km SW Telezhnyi vill., 1040m, 28.7.1998, A. Anichtchenko leg & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) septentrionis (Chaudoir, 1868)
Male adult
E Siberia, Hamar-Daban mts., Osinovka riv., 2000m, 28-31.7.1995, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Phaenoraphis) sagittus Park, 2013 Holotype
Male adult
5-6,VI,2012, Mt. Jeombongsan, Jindong-ri,Girin-myeon, Inje-gun, Gawangwondo Prov. Coll.D.P.Lyu. Image from Park et al, 2013
Copyright © - - -

Pterostichus (Phonias) corrusculus Leconte, 1873
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Phonias) eobius (Tschitscherine, 1899)
Male adult

Copyright © Kirill Makarov

Pterostichus (Phonias) longinquus Bates, 1873
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) strenuus (Panzer, 1796)
Female adult
Russia, Moscow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) diligens (Sturm, 1824)
Male adult
Russia, Moskow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) jankowskyi (Tschitscherine, 1897)
Female adult
Russian Far East, Primorskii krai, Khasanskii district, 9 km SSE Barabash, middle course of Kedrovaya River N43°06'24" E131°32'17" h~120m asl 22-30.V.2015 leg. K.Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) burkhan O.Berlov & Anistschenko, 2005 Paratype
Male adult
Russia, Chita reg., 23km SW Borzia vill. Borzinskoe lake, 1.6.2002, Anichtchenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Platypterus) lineatopunctatus L.Miller, 1850
Male adult
Upper Austria, NE-Alps,Windischgarsten, Rettenbach, Mt. Hoher Nock, h=1960 m,15.VI.2012, Manfred Egger leg.

Copyright © Ruslan Panin

Pterostichus (Platypterus) panzeri (Panzer, 1803)
Female adult
Austria, 47.32N 14.30E, path to Sparafeld Mt, 18.6.2017, L. Minkina leg (cAA) 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Platysma) niger niger s.str.
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.2007 Panin R. leg.

Copyright © Ruslan Panin

Pterostichus (Platysma) eschscholtzii (Germar, 1824)
Male adult

Copyright © Kirill Makarov

Pterostichus (Platysma) cordaticollis Heyden, 1884
Male adult
S Kazakhstan, N slope Talas Mt. rng., Dzhabagly riv., 110m, 28.7.2004, A. Matalin leg.
Copyright © Alexander Anichtchenko

Pterostichus (Plectes) consors (Tschitscherine, 1893)
Female adult
NE Kazakhstan, S Alai Mts., Kalbinsky mt. rng., Ognevka vill., 16.6.2000 (DUBC). det. K. Makarov, 2021
Copyright © Alexander Anichtchenko

Pterostichus (Plectes) drescheri (Fischer, 1817)
Male adult
Russia, W Sayan Mts., Cheremushki vill. 3.8.1998 (DUBC) 18 mm; det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Pterostichus (Pledarus) gibbicollis (Motschulsky, 1844)
Female adult

Copyright © Kirill Makarov

Pterostichus (Pseudethira) ganesh J.Schmidt, 2005 Paratype
Male adult
Nepal, Ganesh Himal, W of Pansang pass, 3200-3500m, 18.6.2000 (cAA); 15mm
Copyright © Alexander Anichtchenko

Pterostichus (Pseudoferonina) lolo Bergdahl, 2011 Holotype
Male adult
deposited in CAS, labeled: “USA: IDAHO,<br /> Clearwater Co.; Cottonwood Creek @ USFS Rd 250; Orogrande Ck. Rd.; NF Clearwater<br /> R. watershed; elev. ~870 m (2860 ft); 12.5 MI. ESE Headquarters, 3.5 MI. SW<br /> of Bungalow; 46°33'10"N, 115°33'08"W; T38N, R7E, S34; 17 July 2009; #73-2009;<br /> J. C. Bergdahl, coll.”/ “HOLOTYPE Pterostichus lolo Bergdahl designated 2011” [red<br /> label].<br /> Photo of James C. Bergdahl & David H. Kavanaugh / ZooKeys 104: 77–96 (2011)
Copyright © - - -

Pterostichus (Pseudomaseus) luctuosus (Dejean, 1828)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Pseudorites) nicaeensis (A. Villa & G.B. Villa, 1835)
Female adult
Italy, Passo della Guardia - Imperia
Copyright © Gianni Allegro

Pterostichus (Pseudosteropus) illigeri (Panzer, 1803)
Female adult
Austria, UTM VN96, 47.30N 14.53E, path to Eisenerzer Reichenstein 21.6.2019, L. Minkina leg. (cAA) 10 mm
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) hagenbachii Sturm, 1824
Male adult
France, Jura, Lajoux, 1190m, 20/05/2013, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Pterostichus (Aphaon) rhilensis kourili Maran, 1933c: 74
Female adult
Bulgaria, Pirin mountains, Vihren, 08.07.2010, E. Svec lgt.; 12 mm, V. Zieris det. 2012, collection Boetzl
Copyright © Fabian Bötzl

Pterostichus (Pterostichus) fasciatopunctatus (Creutzer, 1799)
Female adult
Germany, Bavaria, Passau, Laufenbach dell, F. Bötzl leg. & det., 08.09.2014
Copyright © Fabian Bötzl

Pterostichus (Pterostichus) rutilans (Dejean, 1828)
Male adult
NW Italy, Piemonte,(VC), Oropa, Lago del Mucrone (Lake), h~1900m, 22.VII.1984, L. Diotti leg. L=13 mm
Copyright © Ruslan Panin

Pterostichus (Pterostichus) rufipes (Dejean, 1828)
Male adult
France, 30450, Concoules, 27-05-2017
Copyright © Pascal Deschamps

Pterostichus (Pterostichus) dux L.Schaufuss, 1862
Female adult
Spain, Mts Cantabricos, Macizo Central, Picu Urriellu, 2000m, 13.8.2008 (cAA) leg & det. Anichtchenko A.; 18 mm;
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) honnoratii (Dejean, 1828)
Male adult
France, Vanduse, Mt. Serein, 13.5.1983 (cAA). 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) bruckii Schaum, 1859
Female adult
Serbia, Zaskovci at Priot, 3.21'41N, 22.36'07E, 1620m, 20.04.2024, leg. L. Minkina (cAA). 12.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pterostichus (Qianius) lii Jia & Tian, 2021 Holotype
Male adult
cave Hei Dong, Longjiang Cun, Shenxi Zhen (Town), Honghuagang Qu (District), Zunyi Shi (City), Guizhou Province, 27°36ʹ46ʺN 106°58ʹ08ʺE, 822 m, 22.VIII.2020, leg. Yunchun Li and Yucheng Lin (SCAU).
Copyright © Mingyi Tian

Pterostichus (Rhagadus) solskyi (Chaudoir, 1878)
Female adult

Copyright © Kirill Makarov

Pterostichus (Sinoreophilus) scalptus Sciaky & Wrase, 1997
Male adult

Copyright © Gianni Allegro

Pterostichus (Sinoreophilus) lanista (Tschitscherine, 1898)
Male adult
C Sichuan, Jiyuding Shan, 30km N Wenchuan, 3550m, 2.7.2017, leg. V. Patrikeev (cAA). det. Hongliang Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Sinoreophilus) potanini Tschitscherine, 1889
Female adult
China, N Sichuan, Langmusi vill. 3700 5.8.2015, leg. V. Patrikeev (cAA) det. Hongliang Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Sinosteropus) horstweiperti Schmidt & Gueorguiev, 2013 Paratype
Male adult
NEPAL Arun Vall. to Makalu / Kalo Pokhari 4090m / 29.VI.2013 leg. S. Tamang / 27°41’35N 87°12’18E
Copyright © Borislav Gueorguiev

Pterostichus (Lianoe) masumotoi Tanaka; Morita & Suga, 1987 Holotype
Male adult
Mt. Fuji (south side), 1600 m, Shizuoka Pref., Japan, 31. VII-7. VIII. 1971, K. Tanaka & K. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Steropanus) cavifrons Fedorenko, 2018 Holotype
Male adult
Vietnam, Gia Lai Province, ~50 km N of An Khe, Kon Chu Rang Nat[ure]. Reserv[e]., 14°30–31'N/108°32'E, h=1000– 1040 m, pitfall traps, 24.V–2.VI.2016, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) glymmiger (Andrewes, 1937)
Male adult
Vietnam, Lao Cai Prov., Sa Pa Distr., Fan Si Pan [Phang Xi Pang] Mt., 1400–1500 m, 26.V.–6.VI.1999, N.L.Orlov [leg.], 22°18'56''N 103°49'35''E;
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) pseudoviolaceus Fedorenko, 2018 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2–3 km W of Ngoc Linh Mt., 15°05'N, 107°57'E, h=1600–2150 m, pitfall traps, 19.III– IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) securipenis Fedorenko, 2020 Holotype
Male adult
(ZMMU): Vietnam, Quang Nam Prov[ince]., Nam Gian Distr[ict]., Song Thanh Nat[io]n[al]. Park, 15°33´48´´N/ 107°23´22´´E, h=1050 m, 23.IV–11.V.2019, leg. D. Fedorenko.
Copyright © Dmitri Fedorenko

Pterostichus (Steropus) ebenus (Quensel, 1806)
Male adult
SC Spain, Jaen provincia, Linares, 30.IX.2001, M.A. Lopez Vergara leg.
L=21 mm
Copyright © Ruslan Panin

Pterostichus (Steropus) insidiatrix (Brulerie, 1872)
Male adult
Spain, Teruel, Montes Universales, Pto del Portillo 1780m 31.5.2006, Anichtchenko A. leg & det. (cAA) 19mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) ghilianii Putzeys, 1846
Male adult
Spain, Sierra de Guadarrama, pto Somosierra, 1500m 3.6.2006, Anichtchenko A. leg & det. (DUBC) 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) madidus (Fabricius, 1775)
Male adult
Belgium, Limburg, Forest reserve of Kolmont, 24.4.2016 (DUBC). det. Anichtchenko A., 2021. 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Male adult
Spain, Navarra, Bera de Bidasoa, Larrun Mt. 850m 9-15.9.2001, A. Anichtchenko leg. & det. (cAA) 19 mm
Copyright © Alexander Anichtchenko

Pterostichus (Tausternus) hoii Fedorenko, 2020 Paratype
Male adult

Copyright © Dmitri Fedorenko

Pterostichus (Troglorites) breuili mendizabali Jeannel, 1921
Male adult
Spain, Guipuzcoa, Albistur, (Tolosa) cueva Mendicute, M.Toribio leg.
10 mm
Copyright © Alexander Anichtchenko

Pterostichus (Troglorites) ochsi (Fagniez, 1921)
Female adult
Courmes, Alpes-Maritimes, France
Copyright © Jean Michel Lemaire

Pterostichus (Jedlickaia) sterbai Jedlicka, 1934 Paratype
Male adult
Pe Yen Tsin, Yunnan (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Tschitscherinea) krali Sciaky, 1997 Holotype
Male adult
China, Yunnan, Habashan mts., SE slopes, 4-5.VI.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Tubuliphallus) megacephalus Sciaky & Allegro, 2013 Holotype
Male adult
China, C Sichuan, Xiling Snow Mts., 2100-3100 m, 1-3.VIII.1996 (CSc)
Copyright © Gianni Allegro

Pterostichus (Vietoderus) laevibasis Fedorenko, 2023 Paratype
Male adult
N-Vietnam, 40km W of / CaoBang, Phia Oac Mt./ E-slope, h=1600-1800 m/ 22°36´27´´N105°52´0´´E/ 22.V-6.VI.2018/ leg. A.Abramov.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) abramovi Fedorenko, 2017 Holotype
Male adult
(ZMMU): ‘Vietnam, Dak Lak Prov [ince]., Chu Yang Sin Nat[io]n[al]. Park, 12º222403N 108º212113E, 1.5 km W [of ] Chu Pan Phan Mt, h=1650 m, 30.III–11.IV.2012.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) annamita chuyangsin Fedorenko, 2017 Holotype
Male adult
(ZMMU): ‘Vietnam, Dak Lak Prov.[ince], Chu Yang Sin Natn. Park, 12º232483N/ 108º202593E, Krong Kmar riv[er]., upper flow, h=1200–1650 m, 30.III–14.IV.2012, D. Fedorenko leg.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) kontumensis Fedorenko, 2017 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2-3 km W of Ngoc Linh Mt., 15º052N, 107º572E, h=1600–2150 m, pitfall traps, 19.III–9.IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) subirideus Fedorenko, 2023 Paratype
Male adult
Vietnam, Ha Tinh Prov[ince]./ Vu Quang nat[io]n[al]. Park,/ Kim Quang env., h~200 m/ 18°17´38´´N 105°22´06´´E/ 24.V-1.VI.2022/ leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Wraseiellus) crassiapex Shi & Sciaky, 2013 Paratype
Male adult
China – W Hubei, road Badong-Yesanguan, Tiechanghuang, 1300 m, 27.VI-11.VII.2003 (coll. R. Sciaky)
Copyright © Riccardo Sciaky