Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Male adult
W-Ukraine, Lviv Region, Zhydachiv distr., near Vybranivka, in forest, under a bark of a tree. Panin R. leg.16.X.2008 long 13 mm
Copyright © Ruslan Panin

Diachromus germanus (Linne, 1758)
Male adult
W-Ukraine,
Lviv Reg., Peremyshliany Distr., near Zalissia 28.V.07 Panin R. leg.
11mm .
photo Panin R.
Copyright © Ruslan Panin

Diachromus germanus (Linne, 1758)
Male adult
length: 7,4-9,5 mm
Helvetia, Graubünden, Ilanz
leg.det. U.Schmidt, 2004
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Gynandromorphus etruscus etruscus Quensel, 1806
Male adult
S Ukraine,
Krym Peninsula,Kizil-Koba,15.IV.2009, Shaporinskiy V. leg.
L=10.5mm
Copyright © Ruslan Panin

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Male adult
Collection ground Mukawa cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Diachromus germanus (Linne, 1758)
Male adult
NW Caucas, Krasnodar reg., SE Goriachii Kluch, 21.V.2012 leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Anisodactylus (Anisodactylus) binotatus (Fabricius, 1787)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Anisodactylus (Anisodactylus) nemorivagus (Duftschmid, 1812)
Male adult
Spain, Sierra de Guadarrama, 2 km E Navacerrada, 1900m, 1.9.2002, Anichtchenko A. leg & det. (cAA) 7.5mm
Copyright © Alexander Anichtchenko

Harpalus rufipes (Degeer, 1774)
Male adult
Russia, S.Ural
Copyright © Sunvas

Acinopus (Osimus) ammophilus Dejean, 1829
Male adult
S Ukraine, Krym Peninsula,
Shchelkino distr., near Mysovoe vill., 03.VIII.2008V. Shaporinskiy leg.
23mm
Copyright © Ruslan Panin

Harpalus (Harpalus) affinis (Schrank, 1781)
Male adult
length: 9-12 mm
Helvetia, Graubünden, Ilanz
leg.det. U.Schmidt, 2004
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Harpalus (Harpalus) honestus (Dufischmid, 1812)
Male adult
length: 7,4-10,6 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2007
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Harpalus (Pseudoophonus) rufipes (Degeer, 1774)
Male adult
W-UKRAINE,
Lviv Region, Javorov distr., near Ivano-Frankovo 1-20.VI.07 Panin R. leg
long = 17mm
Copyright © Ruslan Panin

Harpalus (Harpalus) distinguendus (Duftschmid, 1812)
Female adult
POLAND; CE 20 (Lower Vistula Valey)
Bruki Unislawskie
25.04.2009
leg. M. Traczyk
Photo of Mieczyslaw Stachowiak
Copyright © Mieczyslaw Stachowiak

Harpalus (Harpalus) latus (Linne, 1758)
Male adult
length: 10,2 mm
Russia., Sverdlovskaya area, the vicinities of Ekaterinburg city, Verkhnemakarovo vill., floodplain meadow
6.07.2005
Zinovyev E. leg.
Copyright © Evgeniy Zinoviev

Ophonus (Metophonus) puncticeps Stephens, 1828
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Male adult
length: 6,0-9,1 mm Germania, Bayern, Pegnitz<br /> leg.det. U.Schmidt, 1972<br /> Photo: U.Schmidt, 2008
Copyright © U.Schmidt

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Harpalus (Harpalus) distinguendus (Duftschmid, 1812)
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Harpalus (Harpalus) laevipes (as ainus) Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) serripes (Quensel, 1806)
Female adult

Copyright © Kirill Makarov

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Male adult
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) smaragdinus (Duftschmid, 1812)
Male+Female adult
France, forêt de Saint Germain en Laye 78100, 30/8/85, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) anxius (Duftschmid, 1812)
Male+Female adult
France, Forêt de Fontainebleau 77300 18/6/88, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) neglectus Audinet-Serville, 1821
Female adult
France, Ile d’Oléron (17) 12/6/2006, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) pumilus Sturm, 1818: 77
Male+Female adult
France, forêt de Fontainebleau (77300), 15/7/1985, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) tardus (Panzer, 1796)
Male+Female adult
France, forêt de Fontainebleau (77300), 18/4/1987, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Male adult
Portugal, Algarve Portimao, 25/9/1983, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Harpalus (Cephalophonus) cephalotes Fairmaire & Laboulbene, 1854
Male adult
SE Bulgaria, Primorsko; 16mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) honestus (Dufischmid, 1812)
Male adult
CZ, NE Moravia, Moravskoslezské Beskydy - Smrk; 9mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) ardosiacus (Lutshnik, 1922)
Female adult
CZ, S Moravia, Mikulov; 10,5mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) stictus Stephens, 1828
Male adult
CZ, SE Moravia, Uherský Brod; 12mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) flavescens (Piller & Mitterpacher, 1783)
Male adult
CZ, S Moravia, Bzenec; 11mm
Copyright © Aleš Sedláček

Acinopus (Osimus) ammophilus Dejean, 1829
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) froelichii Sturm, 1818: 117
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) servus (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) rubripes (Dufischmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) autumnalis (Duftschmid, 181)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Hesperophonus) azureus (Fabricius, 1775)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) anxius (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) attenuatus Stephens, 1828a: 152
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) cordatus (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) puncticeps Stephens, 1828
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) latus (Linne, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) atratus Latreille, 1804a: 360
Male adult
Germany, Bavaria, Eußenheim, F. Boetzl leg. & det. 19.07.14
Copyright © Fabian Bötzl

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) pumilus Sturm, 1818: 77
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) rubripes (Dufischmid, 1812)
Female adult

Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Male adult
det. A. Matalin
Copyright © Maksim Smirnov

Ophonus (Ophonus) stictus Stephens, 1828
Female adult
W Ukraine, NW Podolian Upland, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.V-15.VI.2017, Panin R. & Kanarsky Yu. leg. L=15 mm
Copyright © Ruslan Panin

Harpalus (Harpalus) solitaris Dejean, 1829a: 337
Female adult
W-Ukraine, Transcarpathian reg., Chornagora Massif, 8 km NE Bohdan vill., h=800m, 9-29.VI..2014, Panin R. leg. L=10.5 mm
Copyright © Ruslan Panin

Harpalus (Semiophonus) signaticornis (Duftschmid, 1812)
Male adult
France, 77860, Quincy-Voisins, 12-05-2018, leg. & det. P. Deschamps
Copyright © Pascal Deschamps

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) modestus Dejean, 1829
Female adult
Russia, 13 km from Irkutsk by Aleksandrovskii road, 20.8.1992, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Cryptophonus) melancholicus Dejean, 1829
Male adult
Italy, Patro di Moncalvo (Asti).
Copyright © Gianni Allegro

Ophonus (Metophonus) melletii (Heer, 1837)
Male adult
Slovakia, Sahy-Tesmak. 7.5 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) schaubergerianus (Puel, 1937)
Male adult
Mor. b. Stramberk. det Wrase. 9 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) parallelus (Dejean, 1829)
Male adult
Spain, Andalucia, Sierra de las Nieves, Los Quejigales, 6-8.4.2010 Anichtchenko A. leg & det. (DUBC) 7 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) rupicola (Sturm, 1818)
Male adult
Russia, NW Caucasus, Krasnodar reg., Taman env., 23.4.2000 (DUBC)
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) laticollis Mannerheim, 1825
Male adult
Russia, NW Caucasus, Novorossiisk distr., Markoth Mts., 23.5.2005 (DUBC) det. Anichtchenko A. 2021; 10.5mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) dimidiatus (P. Rossi, 1790)
Male+Female adult
France 21410 Pont de Pany Sainte-Marie-sur-Ouche 30-06-1984 et 18130 Dun sur Auron 18-05-2012
Copyright © Pascal Deschamps

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
Italy, Piedmont, Sambuco (CN), m 1400, 4.VI.2006
Copyright © Gianni Allegro

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Drymoharpalus) atratus Latreille, 1804
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Anamblystus) luteicornis (Dufischmid, 1812)
Female adult
Latvia, Ilukste, 6.2010 M. Janovska leg (DUBC). det. Anichtchenko A. 2023
Copyright © Alexander Anichtchenko

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest, on the sands, 17.V-25.VI.2024, R.Panin leg., L=13 mm
Copyright © Ruslan Panin

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Kirill Makarov

Harpalus (Amblystus) rufipalpis rufipalpis
Male adult
Latvia, Ilgas 10.6.92 (DUBC) 9.8 mm
Copyright © Alexander Anichtchenko

Harpalus (Actephilus) picipennis (Duftschmid, 1812)
Male adult
Latvia, Ilgas 18.6.96 (DUBC). 6.3 mm
Copyright © Alexander Anichtchenko

Harpalus (Actephilus) picipennis (Duftschmid, 1812)
Female adult
Latvia, Ilgas 18.6.96 (DUBC) 6.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Cryptophonus) litigiosus Dejean, 1829
Male adult
Spain, Andalusia, Sierra Arana, 25 km NE Granada, near cueva del Agua, 16.x.2003, 1700m, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Acupalpus (Acupalpus) meridianus (Linnaeus, 1761)
Male adult
length: 3-4 mm
Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1993
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Anthracus consputus (Duftschmid, 1812)
Male adult
length: 3,8-5,1 mm<br /> Austria, Burgenland, Schützen a. Gebirge<br /> leg.det., G.Rößler, 1977<br /> Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Bradycellus (Bradycellus) harpalinus (Audinet-Serville, 1821)
Male adult
length: 3,6-5,1 mm
Germania, Oberfranken, Marktredwitz
leg.det. G.Rößler, 1992
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Dicheirotrichus (Dicheirotrichus) gustavii Crotch, 1871
Male adult
Karelia, Loukhi Distr., N shore of Olenevskij Isl. 4-24.VIII.2006 I.Melnik leg.<br /> L=7.0 mm
Copyright © Kirill Makarov

Dicheirotrichus (Dicheirotrichus) gustavii Crotch, 1871
Male adult
length: 5,2-7,2 mm
Holland, Goes
leg.det. Veldkamp, 1978
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Dicheirotrichus (Trichocellus) cognatus (Gyllenhal, 1827)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Stenolophus (Stenolophus) mixtus (Herbst, 1784)
Male adult
Poland: EB56
Piotrawin
13.VI.2008
leg. G. Bistu&a-Prószy&ski
Copyright © Mieczyslaw Stachowiak

Stenolophus (Stenolophus) mixtus (Herbst, 1784)
Female adult
length: 5-6 mm
Germania, Thüringen, Harra
leg.det. U.Schmidt, 2000
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Stenolophus (Stenolophus) teutonus (Schrank, 1781)
Male adult
Azerbaijan,
near Khachmaz, 16.VII.2008, Pljushch I.G. leg.
L=7mm
Copyright © Ruslan Panin

Dicheirotrichus (Trichocellus) placidus (Gyllenhal, 1827)
Male adult
length: 4,2-5 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2006
Photo: U.Schmidt, 2008
Copyright © U.Schmidt

Bradycellus (Bradycellus) caucasicus Chaudoir, 1846: 187 (Acupalpus)
Male adult
L=4,0 mm
RUSSIA, Tver reg., Likhoslavlskiy distr., Domanikha village, in the garden, in soil. H=160 m. N57*13.629’ E035*45.675’ 21.VII.2005
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Acupalpus (Acupalpus) flavicollis (Sturm, 1825)
Male adult
L=3,6 mm
RUSSIA, Moscow reg., Orekhovo-Zuevskiy reg., 3 km to S from Anciferovo settl., grassland. 16.V.2001
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Acupalpus (Acupalpus) parvulus (Sturm, 1825)
Male adult

Copyright © Kirill Makarov

Acupalpus (Acupalpus) exiguus Dejean, 1829
Male adult

Copyright © Kirill Makarov

Acupalpus (Acupalpus) meridianus (Linnaeus, 1761)
Male adult

Copyright © Kirill Makarov

Bradycellus (Bradycellus) verbasci (Duftschmid, 1812)
Female adult
det. R.Panin & D.Khrapov
Copyright © Kirill Makarov

Anthracus consputus (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bradycellus (Bradycellus) ruficollis Stephens, 1828a: 168 (Trechus)
Male adult
Czech Republic, Bohemia occ., Kraslice, Tisovec hill, 720-760 m a.s.l., 11.x.1987, lgt. et det. J. Hejkal
Copyright © Pavel Krasensky

Bradycellus (Bradycellus) csikii Laezo, 1912
Female adult
W Ukraine, Lviv city, Sykhiv, waste-ground, 49°47'29"N, 24°3'18"E, 7.IV.2018, Panin R leg., L=4.2mm
Copyright © Denis Khrapov

Bradycellus (Bradycellus) verbasci (Duftschmid, 1812)
Male adult
W Ukraine, Lviv city, Holosko Cemetery, 3.IX.2014, Panin R. leg. L=5.1 mm
Copyright © Denis Khrapov

Bradycellus (Bradycellus) harpalinus (Audinet-Serville, 1821)
Male adult
W Ukraine, Lviv city, Holosko, bank of Natural lake, R.Panin leg., 3.VI.2017, L=4,2 mm
Copyright © Denis Khrapov

Acupalpus (Acupalpus) dubius Schilsky, 1888
Male adult
W-Ukraine, Roztochia, Lviv Reg., Yavoriv distr., N of Ivano-Frankovo vill., Site «Zalyvky» peat bog, 12.V - 8.VI.2021, R.Panin leg., L=3.3 mm
Copyright © Denis Khrapov