Carabus (Ohomopterus) yamato Nakane, 1953
Female adult
Japan, Ohkubo, 740-770m, Shirakawa-mura
Ohno-gun (Gifu) 20-25.6.80 Ishikawa leg.
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) yaconinus yamaokai Ishikawa & Kubota, 1994
Female adult
Japan, Ehime pref., Taisanji, 14.5.2002, Matsuyama city, T. Kurihara leg. (cAA)
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) yaconinus yaconinus Bates, 1873
Male adult
Japan, Kagawa pref, Shin-ike
Awai town, Kannonji city, 4.3.1993
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) maiyasanus maiyasanus Bates, 1873
Female adult
Japan, Shiga pref.,
Lake Yogo 18.8.1995 Shimizu leg.
24 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Yamanashi pref., Kodachi, Kawaguchiko-cho, Minamitusru distr., 2.IX.2000 T. Kurihara leg. (cAA). 29 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Female adult
15.VII.1998 JAPAN, Ichihara, env. Chiba
Copyright © Nil Belonogov

Carabus (Ohomopterus) arrowianus murakii Ishikawa & Kubota, 1984
Male adult
Sho, alt 3m, Futami-T, Watarai-Co. Mie pref. Japan, 17.5.1987 Y. Imura leg. (cAA) 27 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) arrowianus arrowianus Breuning, 1934
Male adult
Japan, Shizuoka pref., Mt. Mitake, Inasa town 18.10.1998 T. Kurihara leg.
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) albrechti esakianus Nakane, 1961
Male adult
Japan, Tokyo pref, Mt. Takao, Nachioji City, 11.8.1990 K. Kurihara leg. (cAA)
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
Male adult
Japan, Yamanashi pref., Koganesawa-rindo, Otsuki city, 13.6.1990 K. Kurihara leg. (cAA). 20 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Female adult
Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, Yamanashi pref., Otawa-rindo,
Narusawa vill., 9.IX.2002 T. Kurihara leg
male, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) arboreus exarboreus (Ishikawa, 1992)
Male+Female adult
E.Russia, Far East, Sakhalin Isl., forest of M.Tym river, 11-IX-2000 . Egorov leg. male 27mm, female 30mm.
Copyright © Safronov

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan: Niigata pref., Chayagahara Jyouetsu City,
18.XI.2006 A.Toba leg.
Copyright © Alexander Anichtchenko

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
L=34 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=33 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=21 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus arboreus Lewis, 1882
Male adult
N-Japan, SW-Hokkaido, Chitose sity, 25.V-8.VI.2013, T. Hirabayashi leg.<br /> 32 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
Male adult
Japan: C Honshu: Niigata pref., Mysato: Sakae Abiko-City; Iwafune distr., Awashima Isl. (LABEL ERROR)
Copyright © Igor Solodovnikov

Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
Female adult
Japan: C Honshu: Niigata pref., Mysato: Sakae Abiko-City; Iwafune distr., Awashima Isl. (LABEL ERROR)
Copyright © Igor Solodovnikov

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. 23 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) kaempferi Thomson, 1875 Holotype
Male adult
Image courtesy of Biological Museum, Lund University
Copyright © - - -

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Female adult
Japan, S Honshu, Hyogo Pref., Nishiwaki city, 20.VI.2010, Shinichi S. leg., L=37 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C Honshu, Nagano Prefecture, Nagano city, 3.IV.2012, S. Shinichi leg. L=42 mm. det. Anichtchenko A. 2021
Copyright © Ruslan Panin

Carabus (Leptocarabus) harmandi Lapouge, 1909 Lectotype
Male adult
Honshu, Tochigi, Mts near Nikko. Muséum national d’Histoire naturelle, Paris (France) Collection: Insects - Coleoptera (EC) Specimen MNHN-EC-EC549
Copyright © MNHN

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan, C Honshu, Nagano Pre., Toyooka town, 28.XII.2019, S. Shinichi leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
Male adult
Hagyu., Iide Town, Nishiokitama Co., Yamagata pref., Japan, 17 3.2016.T. Nakamura leg. (DUBC). 35mm
Copyright © Alexander Anichtchenko

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, Kiyose, Tokyo, 20.1.1961 (DUBC). 40 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Male adult
Japan, S Honshu, Mie Pref., Inabe city, 21.II.2021, Shinichi S. leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) kyushuensis cerberus Ishikawa, 1962
Female adult
Fuchu (Mitsugi,Hiroshima) JAPON, VII-2001. 34 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) kyushuensis nakatomii Ishikawa, 1966
Male adult
Tombara (Honshu,Shimane) JAPON, VII-2001. 36 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) kyushuensis kyushuensis Nakane, 1960
Female adult
Izumi, (Honshu, Kumamoto) JAPON, VII-2001. 31 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus akitanus (Ishikawa, 1992)
Male adult
Muriyoshi (Akita), JAPON, VIII-2001; 26 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus babai Ishikawa, 1968
Male adult
Hiychiyama (Niigata, Honshu), JAPON, XI-1984; 20 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus gracillimus Bates, 1883
Female adult
Mt.Tate-yama, 2000m (Toyama, Honshu), JAPON, X-1994; 20 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus karibanus (Ishikawa, 1992)
Female adult
Kuromatsunai (Hokkaido), JAPON, X-1994; 30 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus kitakamisanus (Ishikawa, 1992)
Male adult
Niisato (Iwate, Honshu), JAPON, VIII-2001; 30 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus nepta (Ishikawa, 1984)
Male adult
North, Hakkoda-san (Aomori, Honshu), JAPON, VI-1986 ; 25 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus ogurai Ishikawa, 1969
Female adult
Tabayama (Yamanashi, Honshu), JAPON, X-1990; 24 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus pronepta (Ishikawa, 1992)
Male adult
Hakkode-san (Aomori, Honshu), JAPON, XI-1990; 24 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus shinanensis Born, 1922
Male adult
Mt. Yatsugatake (Nagano, Honshu), JAPON, VII-1985; 23 mm
Copyright © Alain Mollard

Carabus (Ohomopterus) albrechti okumurai (Ishikawa, 1966)
Male+Female adult
Mt. Daibosatzu (Yamanashi, Honshu) JAPON
Copyright © Alain Mollard

Carabus (Ohomopterus) albrechti tohokuensis Ishikawa, 1984
Male+Female adult
Hashimori Pass (Iwate) JAPON
Copyright © Alain Mollard

Carabus (Ohomopterus) albrechti freyi van Emden, 1932
Male+Female adult
Sado Island (Niigata) JAPON
Copyright © Alain Mollard

Carabus (Ohomopterus) arrowianus nakamurai (Ishikawa, 1966)
Female adult
Mt.Takamoriyama (Tida-City, Nagano) JAPON VI-1987
Copyright © Alain Mollard

Carabus (Ohomopterus) chugokuensis chugokuensis (Nakane, 1961)
Male+Female adult
Mt.Takasho (Shiman) JAPON VI-1987
Copyright © Alain Mollard

Carabus (Ohomopterus) lewisianus awakazusanus Ishikawa, 1981
Male adult
Mt. Iyogatake 280m, Tomiyama town, Awa Co., Chiba pref, 4.2.1996 K. Watanabe leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) albrechti Morawitz, 1862
Male adult
Horoman, Samani town, Samani Co., Hokkaido, 16.6.2015 T. Nakamura leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola nishikawai (Ishikawa, 1966)
Male adult
Yamada, Isumi city, Chiba pref. Japan 20.5.2007 K. Osozawa leg. (cAA). 28mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Kiyose, Tokyo 20.1.1961 (cAA). 29 mm
Copyright © Alexander Anichtchenko

Cylindera (Apterodela) ovipennis (Bates, 1883)
Male adult
Japan, Nagano pref., Iiyama city, Ichiyama, Sekida pass, 1100 m, S. Kakinuma leg. 02.06.2013
Copyright © Fabian Bötzl

Abroscelis anchoralis punctatissima (Schaum, 1863)
Female adult
Japan, Ishikawa (JWGC). det. J. Wiesner
Copyright © Peter Schüle

Cylindera (Eugrapha) elisae novitia (Bates, 1883)
Male adult
ZIN
Copyright © Igor Solodovnikov

Cylindera (Apterodela) ovipennis (Bates, 1883)
Female adult
Japan, Niigata pref., Itoigawa city., Agero, 800-1000m, 28.5.2016 leg. S. Kakinuma (cAA)
Copyright © Alexander Anichtchenko

Cylindera (Apterodela) ovipennis (Bates, 1883)
Male adult
Japan, Niigata pref., Itoigawa city., Agero, 800-1000m, 28.5.2016 leg. S. Kakinuma (cAA)
Copyright © Alexander Anichtchenko

Cicindela (Cicindela) sachalinensis niohozana Bates, 1883
Female adult
Japan, Hokkaido, Sapporo-shi, Jozankei, 42.999N 141.073E, 27.VI.2021, Masaki Endo leg.
Copyright © Ruslan Panin

Callytron yuasai yuasai
Male adult

Copyright © Michio Hori

Trichotichnus (Trichotichnus) fukuharai Habu, 1957 Holotype
Male adult
Tokura, Oze, GUNMA, 04. IX. 1952, N. Fukuhara. COL-137
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) fukuharai Habu, 1957 Holotype
Male adult
Tokura, Oze, GUNMA, 04. IX. 1952, N. Fukuhara. COL-137
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) fukuharai Habu, 1957 Holotype
Male adult
Tokura, Oze, GUNMA, 04. IX. 1952, N. Fukuhara. COL-137
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Male adult
Collection ground Ashikaga city,Tochigi Pref.Ashikaga～shi 36079
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Male adult
Daegu-si, Republic of Korea. Scale bar: 3 mm.
Copyright © Dooyoung Kim

Trichotichnus (Bottchrus) nanus Habu, 1954c: 233
Male adult
Daegu-si, Republic of Korea. Scale bar: 2 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) fukuharai Habu, 1957
Male adult
Yunnan. Image from Kataev et al, 2022
Copyright © - - -

Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Female adult
Primorye Territory, Khasan Distr., vicinity of Slavyanka Vill., Cape Busse, 42°52'42"N 131°27'58"E, 13.VI.2023, I. Melnik leg. L=10.0mm
Copyright © Kirill Makarov

Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Male adult
Primorye Territory, Khasan Distr., vicinity of Slavyanka Vill., Cape Busse, 42°52'42"N 131°27'58"E, 13.VI.2023, I. Melnik leg. L=11.0mm
Copyright © Kirill Makarov

Odacantha (Odacantha) hagai Nemoto, 1989b: 101
Male adult
Khasansky Distr., mouth of Kedrovaya River, 22–24.VIII 2018, M. Korepanov, K. Makarov, A. Matalin & Yu. Sundukov leg.
Copyright © Kirill Makarov

Odacantha (Odacantha) hagai Nemoto, 1989b: 101
Male adult
Khasansky Distr., mouth of Kedrovaya River, 22–24.VIII 2018, M. Korepanov, K. Makarov, A. Matalin & Yu. Sundukov leg.
Copyright © Kirill Makarov

Odacantha (Odacantha) hagai Nemoto, 1989b: 101
Female adult
Khasansky Distr., mouth of Kedrovaya River, 22–24.VIII 2018, M. Korepanov, K. Makarov, A. Matalin & Yu. Sundukov leg.
Copyright © Kirill Makarov

Odacantha (Odacantha) hagai Nemoto, 1989b: 101
Female adult
Khasansky Distr., mouth of Kedrovaya River, 22–24.VIII 2018, M. Korepanov, K. Makarov, A. Matalin & Yu. Sundukov leg.
Copyright © Kirill Makarov

Lebia (Poecilothais) hikosana Habu, 1955
Male adult
Dôgo Is., Fuse (Mt. Daimanji-san), 25.V.2023. Image from Takashi Shimizu and Ryo Nakamura, 2023
Copyright © - - -

Lebia (Poecilothais) idae Bates, 1873
Female adult
Dôgo Is., Fuse (Mt. Daimanji-san), 16.IX.2022. Image from Takashi Shimizu and Ryo Nakamura, 2023
Copyright © - - -

Nebria (Sadonebria) hakusana Sasakawa, 2022 Holotype Paratype
Male+Female adult
Holotype, male (KS), Shinmata Pass, Nomuki, Katsuyama, Fukui Prefecture, Japan, 1.vi.2020, S. Inoue leg.; paratype: female, KAGA-Hakusan Nakahanba ~ Bettonozoki (1,500~1,750 m) 1.VIII. 1973 M. Miyatake
Copyright © - - -

Jujiroa ohminensis Wada & Ashida, 2022 Holotype
Male adult
(OMNH TI 535), 23.V.2021 (bated trap set on 3.V.2021), H. Ashida leg. (Asahi, 540 m in altitude, Totsukawa-mura, Nara Prefecture, Japan). Image from Wada & Ashida, 2022
Copyright © - - -

Jujiroa yorikoae Morita & Inagaki, 2023 Holotype
Male adult
Cave Fujigano-no-ana (= Kurotengu-no-ana), 8−16.II.2023, M. Inagaki leg. Image from Morita & Inagaki, 2023
Copyright © - - -

Jujiroa elongata Ueno, 1955
Male adult
Cave Fujigano-no-ana (= Cave Kurotengu-no-ana), Aso, Taiki-chô, Watarai-gun, Mie Prefecture, central part of Honshu, Japan, 8.II.2023, M. Inagaki leg. Image from Morita & Inagaki, 2023
Copyright © - - -

Jujiroa ohminensis Wada & Ashida, 2022 Paratype
Male adult
“Paratype Jujiroa ohminensis Wada et Ashida, 2022”// “ 奈良県十津川村 旭 ドロゴボワ谷(540m) 12–VI–2021 (23–V trap) 芦田 久 採集 ” // “11474 Hisashi Ashida”. Image from Morita & Inagaki, 2023
Copyright © - - -

Atranodes uenoi Habu, 1971
Female adult
Mt. Shirakusa-yama,Norimasa, Gero-shi, Gifu Prefecture, 22.IX–4.X.2009, N. Toda leg. Image from Uchida, Morita & Y. Kurosa, 2025
Copyright © - - -

Amara (Bradytus) macra Bates, 1883
Male adult

Copyright © Kirill Makarov

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trephionus abiba Sasakawa & Itô, 2018 Holotype
Male adult
Mt. Ishikiri (entrance of the Ishikiri Cave), Haguro, Tainai-shi, Niigata Pref. (38.051754°N, 139.435703°E), 23.V.2013, H. Itô leg. Image from Kôji Sasakawa & Hirotarô Itô, 2018
Copyright © - - -

Trephionus inexpectatus Sasakawa & Itô, 2018 Holotype
Male adult
Kitazawa Pass, alt. ca. 2000 m, Hase-mura, Nagano Pref., 7–15.X.1998, M. Hama leg. Image from Kôji Sasakawa & Hirotarô Itô, 2018
Copyright © - - -

Trephionus niumontanus Sasakawa, 2018 Holotype
Male adult
Mt. Kunimi, asl. ca. 480 m, Fukui-shi, Fukui Pref., 27.VI.2012, S. Inoue leg. Image from Kôji Sasakawa & Hirotarô Itô, 2018
Copyright © - - -

Trephionus cylindriphallus Sasakawa, 2018
Male adult
12 - Holotype, 13 - Paratype from Abe Pass. Image from Kôji Sasakawa & Hirotarô Itô, 2018
Copyright © - - -

Trephionus cylindriphallus Sasakawa, 2018 Holotype
Male adult
Kawamata, University Forest in Chichibu, The University of Tokyo, Ôtaki, Chichibu-shi, Saitama Pref., 16.X.2009, N. Nishiyama leg. Image from Kôji Sasakawa & Hirotarô Itô, 2018
Copyright © - - -

Trephionus bifidilobatus Sasakawa & Itô, 2018 Holotype
Male adult
Kotosawa, alt. ca. 310 m, Mitake, Oume-shi, Tokyo (35.795931°N, 139.168746°E), 10–12.IX.2009, K. Sasakawa leg. Image from Kôji Sasakawa & Hirotarô Itô, 2018
Copyright © - - -

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
South Primorye, Kedrovaya Pad nature reserve, mouth of Kedrovaya River, swamp 43°04'47"N 131°36'42"E 1.VI.2019 leg. K. Makarov L= 7.1
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
South Primorye, Kedrovaya Pad nature reserve, mouth of Kedrovaya River, swamp 43°04'47"N 131°36'42"E 1.VI.2019 leg. K. Makarov L= 7.1
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Female adult
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Female adult
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Female adult
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.8 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.2 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.6 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Amara (Bradytus) macra (Bates, 1883)
Female adult
Kunashir Isl., Tret’yakovo Vill., 43°59'09"N, 145°39'15"E, h ~ 26 m 20–25.IX.2009 leg. I. Melnik, A. Zaitsev L=10.8 mm
Copyright © Kirill Makarov

Amara (Bradytus) macra (Bates, 1883)
Male adult
Kunashir Isl., Tret’yakovo Vill., 43°59'09"N, 145°39'15"E, h ~ 26 m 20–25.IX.2009 leg. I. Melnik, A. Zaitsev L=10.8 mm
Copyright © Kirill Makarov

Pterostichus (Epinialoe) fumikoae Y.Kurosa, 2019 Holotype
Male adult
Kiyozasa-tôge, Fujieda-shi, 11.XI.2017, Y. Kurosa leg. (National Museum of Nature and Science, Tsukuba). Image from Kurosa, 2019
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Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Kunashiri Isl., mouth of Medny Spring 44°16'31"N 145°56'29"E 27.VI.2008 leg. K.Makarov
L=4.3
Copyright © Kirill Makarov

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033. (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html). Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu. Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Thalassoduvalius masidai Ueno, 1956a: 65
Male adult
Japan, Kioto, leg. S. Kawai
Copyright © Vittorino Monzini

Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,7
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,8
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5,2
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5.0
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,75 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,6 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,85 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,45
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=5.0 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=4,7 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,55 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,75
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,9 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,8
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,3 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,6 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=5 mm
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum sanatum
Male adult
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum sanatum
Female adult
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov

Epaphiopsis (Epaphiopsis) narukawai Morita, 2021 Paratype
Female adult
Mt. Gozaisho-dake, alt. 1,000 m, Komonochô, Mie-gun, Mie Prefecture, Honshu, Japan. Image from Morita, 2021
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