Acinopus (Acinopus) picipes (Olivier, 1795)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr., near Bogatoe Ushchele vill. 04.VIII.2009 Shehovcov A. leg.
L=16 mm
Copyright © Ruslan Panin

Acinopus (Acinopus) picipes (Olivier, 1795)
Female adult

Copyright © Kirill Makarov

Acinopus (Oedematicus) gutturosus Buquet, 1840
Male adult
Spain, Algeciras, San Roque,4.2014, Anichtchenko A. leg.; det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Acinopus (Acinopus) giganteus Dejean, 1831
Male adult
Spain, Andalucia, Sierra de las Nieves, Los Quejigales, 6-8.4.2010, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Acinopus (Acmastes) haroldii Schaum, 1863
Male adult
Spain, Cadiz, Tarifa, 10.9.1989, J.L. Torres (cAA) 15 mm
Copyright © Alexander Anichtchenko

Acinopus (Acinopus) almeriensis Mateu, 1954
Male adult
Spain, Almeria, Cabo de Gata, playa de San Jose 20.4.2004 Anichtchenko A. leg & det. (cAA) 14mm
Copyright © Alexander Anichtchenko

Daptus vittatus Fischer von Waldheim, 1823: pl.46
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus rufipes (Degeer, 1774)
Male adult
Russia, S.Ural
Copyright © Sunvas

Harpalus (Harpalus) affinis (Schrank, 1781)
Male adult
length: 9-12 mm
Helvetia, Graubünden, Ilanz
leg.det. U.Schmidt, 2004
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Harpalus (Harpalus) honestus (Dufischmid, 1812)
Male adult
length: 7,4-10,6 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2007
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Harpalus (Pseudoophonus) rufipes (Degeer, 1774)
Male adult
W-UKRAINE,
Lviv Region, Javorov distr., near Ivano-Frankovo 1-20.VI.07 Panin R. leg
long = 17mm
Copyright © Ruslan Panin

Harpalus (Harpalus) distinguendus (Duftschmid, 1812)
Female adult
POLAND; CE 20 (Lower Vistula Valey)
Bruki Unislawskie
25.04.2009
leg. M. Traczyk
Photo of Mieczyslaw Stachowiak
Copyright © Mieczyslaw Stachowiak

Harpalus (Harpalus) latus (Linne, 1758)
Male adult
length: 10,2 mm
Russia., Sverdlovskaya area, the vicinities of Ekaterinburg city, Verkhnemakarovo vill., floodplain meadow
6.07.2005
Zinovyev E. leg.
Copyright © Evgeniy Zinoviev

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Harpalus (Harpalus) distinguendus (Duftschmid, 1812)
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Harpalus (Harpalus) laevipes (as ainus) Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) serripes (Quensel, 1806)
Female adult

Copyright © Kirill Makarov

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Male adult
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) dispar Dejean, 1829
Male adult
France, L’Albaron, 13123 Pascal 9/10/2001, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) smaragdinus (Duftschmid, 1812)
Male+Female adult
France, forêt de Saint Germain en Laye 78100, 30/8/85, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) anxius (Duftschmid, 1812)
Male+Female adult
France, Forêt de Fontainebleau 77300 18/6/88, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) oblitus Dejean, 1829a: 273
Male+Female adult
France, Brétignolles sur mer 85470 17/5/85, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) neglectus Audinet-Serville, 1821
Female adult
France, Ile d’Oléron (17) 12/6/2006, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) pumilus Sturm, 1818: 77
Male+Female adult
France, forêt de Fontainebleau (77300), 15/7/1985, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) tardus (Panzer, 1796)
Male+Female adult
France, forêt de Fontainebleau (77300), 18/4/1987, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Male adult
Portugal, Algarve Portimao, 25/9/1983, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) honestus (Dufischmid, 1812)
Male adult
CZ, NE Moravia, Moravskoslezské Beskydy - Smrk; 9mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) rubripes (Dufischmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) autumnalis (Duftschmid, 181)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) oblitus Dejean, 1829a: 273
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Baeticoharpalus) lopezi Serrano & Lencina, 2008
Male adult
Spain, Jaen prov., Sierra Magina, 2000m, 37.728701, -3.460772 3-4.05.2014, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) anxius (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) attenuatus Stephens, 1828a: 152
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) latus (Linne, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) atratus Latreille, 1804a: 360
Male adult
Germany, Bavaria, Eußenheim, F. Boetzl leg. & det. 19.07.14
Copyright © Fabian Bötzl

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) pumilus Sturm, 1818: 77
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) rubripes (Dufischmid, 1812)
Female adult

Copyright © Kirill Makarov

Harpalus (Semiophonus) signaticornis (Duftschmid, 1812)
Male adult
France, 77860, Quincy-Voisins, 12-05-2018, leg. & det. P. Deschamps
Copyright © Pascal Deschamps

Harpalus (Harpalus) fuscicornis Menetries, 1832: 134
Female adult
Germany, Iphofen, 2018, leg. & det. Boetzl
Copyright © Fabian Bötzl

Harpalus (Harpalus) pygmaeus Dejean, 1829
Female adult
Greece bor. occ., 15 km W of Metsovo, Petra, 9.6.2008, leg. Orszulik
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) modestus Dejean, 1829
Female adult
Russia, 13 km from Irkutsk by Aleksandrovskii road, 20.8.1992, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Cryptophonus) melancholicus Dejean, 1829
Male adult
Italy, Patro di Moncalvo (Asti).
Copyright © Gianni Allegro

Harpalus (Harpalus) semipunctatus Dejean, 1829
Male adult
Spain, Guadalajara, pto de Maranchon, 28.4.2006, 1300m, Anichtchenko A. leg. & det. (cAA) 11mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) contemptus Dejean, 1829
Male adult
Spain, Sierra Nevada, 5 km N Capileira, 1800-2000m, 25.4.2003, Anichtchenko A. leg. & det. (cAA) 10 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) ebeninus Heyden, 1870
Male adult
Spain, Picos de Europa, Macizo central, Llagu Bajero, 1890m, 6-7.8.2003, Anichtchenko A. leg & det. (cAA) 11mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) microthorax (Motschulsky, 1849)
Male adult
Spain, Toledo, lagunas de Quero, 28.7.2008, Anichtchenko A. leg & det. (cAA) 10 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) nevadensis K. Daniel & J. Daniel, 1898
Male adult
Spain, Sierra Nevada, 5km N Capileira, 1800-2000m, 25.4.2003, Anichtchenko A. leg & det. (cAA) 9 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) punctatostriatus Dejean, 1829
Female adult
Spain, Teruel, Montes Universales, pto del Portillo, 1780m, 31.5.2006, Anichtchenko A. leg & det. (cAA) 10mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) sulphuripes goudotii Dejean, 1829
Male adult
Algeria, Mostaganem, 30.12.1972, V. Orlov leg. (cAA) 7mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) zabroides Dejean, 1829
Male adult
E Siberia, Irkutsk, downtown, 20.6.2001, Anichtchenko A. leg & det. (cAA) 15 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) angustitarsis Reitter, 1887
Male adult
Spain, Segovia, Valsain, 3.5.2003 (cAA) det. Anichtchenko A. 7.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) rufipalpis wagneri Schauberger, 1926
Male adult
Spain, Sierra Nevada, rio Trevelez, 1200m, 6km abajo Trevelez, 7.4.2004, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) albanicus Reitter, 1900
Male adult
Spain, Granada, Castell del Ferro, 5.4.2002 Anichtchenko A. leg. (cAA) 8.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) dimidiatus (P. Rossi, 1790)
Male+Female adult
France 21410 Pont de Pany Sainte-Marie-sur-Ouche 30-06-1984 et 18130 Dun sur Auron 18-05-2012
Copyright © Pascal Deschamps

Harpalus (Harpalus) decipiens decipiens
Male+Female adult
Espagne Alto de Morredero sierra de Teleno 21-05-2001 et Cabeza de Manzaneda 24-05-2001
Copyright © Pascal Deschamps

Harpalus (Harpalus) decipiens decipiens
Male adult
Portugal, Serra Estrela, 5.VI.1993
Copyright © Gianni Allegro

Harpalus (Cryptophonus) fulvus Dejean, 1829
Male adult
Italy, Sardinia, Chia (CA), Stagno Campana, 29.VIII.1994
Copyright © Gianni Allegro

Harpalus (Harpalus) punctatostriatus Dejean, 1829
Male adult
Italy, Sardinia, Arborea (OR), 9.VI.1978
Copyright © Gianni Allegro

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
Italy, Piedmont, Sambuco (CN), m 1400, 4.VI.2006
Copyright © Gianni Allegro

Harpalus (Licinoderus) franzi Mateu, 1954
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Epiharpalus) ebeninus Heyden, 1870
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Drymoharpalus) atratus Latreille, 1804
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Cryptophonus) fulvus Dejean, 1829
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Kirill Makarov

Harpalus (Cryptophonus) litigiosus Dejean, 1829
Male adult
Spain, Andalusia, Sierra Arana, 25 km NE Granada, near cueva del Agua, 16.x.2003, 1700m, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) puncticeps Stephens, 1828
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Male adult
length: 6,0-9,1 mm Germania, Bayern, Pegnitz<br /> leg.det. U.Schmidt, 1972<br /> Photo: U.Schmidt, 2008
Copyright © U.Schmidt

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Ophonus (Ophonus) ardosiacus (Lutshnik, 1922)
Female adult
CZ, S Moravia, Mikulov; 10,5mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) diffinis (Dejean, 1829)
Male adult
CZ, SE Moravia, Bílé Karpaty Mts.; 11mm
Copyright © Aleš Sedláček

Ophonus (Hesperophonus) cribricollis (Dejean, 1829)
Male adult
SK, S Slovakia, Kamenica nad Hronom; 8mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) stictus Stephens, 1828
Male adult
CZ, SE Moravia, Uherský Brod; 12mm
Copyright © Aleš Sedláček

Ophonus (Hesperophonus) azureus (Fabricius, 1775)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) cordatus (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) puncticeps Stephens, 1828
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Male adult
det. A. Matalin
Copyright © Maksim Smirnov

Ophonus (Ophonus) stictus Stephens, 1828
Female adult
W Ukraine, NW Podolian Upland, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.V-15.VI.2017, Panin R. & Kanarsky Yu. leg. L=15 mm
Copyright © Ruslan Panin

Ophonus (Hesperophonus) subquadratus (Dejean, 1829)
Male adult
Greece, Corfu, Benitses env.,23-29.6.2003. det. D Wrase (det confirmed by Anichtchenko 2021); 7 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) melletii (Heer, 1837)
Male adult
Slovakia, Sahy-Tesmak. 7.5 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) schaubergerianus (Puel, 1937)
Male adult
Mor. b. Stramberk. det Wrase. 9 mm
Copyright © Alexander Anichtchenko

Ophonus (Incisophonus) incisus (Dejean, 1829)
Female adult
France Gard dep., Nimes 3.11.2013 (cAA) 14 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) cunii Fairmaire, 1880
Male adult
Spain, Andalicia, Sierra de Grazalema, Puerto de las Palomas, 10.4.2010, Anichtchenko A. leg. & det. (cAA) 9.5 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) parallelus (Dejean, 1829)
Male adult
Spain, Andalucia, Sierra de las Nieves, Los Quejigales, 6-8.4.2010 Anichtchenko A. leg & det. (DUBC) 7 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) rupicola (Sturm, 1818)
Male adult
Russia, NW Caucasus, Krasnodar reg., Taman env., 23.4.2000 (DUBC)
Copyright © Alexander Anichtchenko

Ophonus (Ophonus) opacus (Dejean, 1829)
Male adult
Spain, Cadiz, Paterna de Ribera 14.11.1999 (DUBC) det. A. Anichtchenko, 2021; 11.5mm
Copyright © Alexander Anichtchenko

Ophonus (Hesperophonus) subquadratus (Dejean, 1829)
Male adult
Bulgaria, Chernomorets, 24.11.2006, Anichtchenko A. leg. & det. (DUBC)
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) subsinuatus Rey, 1886
Male adult
WС Ukraine, Vinnytsa reg., Lityn distr., Petryk vill., to light, 5.VI.2020, S. Kokhan leg.,/det.R.Panin, L=7,6mm
Copyright © Denis Khrapov

Pangus scaritides (Sturm, 1818)
Male adult
SK, S Slovakia, Velké Kosihy; 8mm
Copyright © Aleš Sedláček

Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)
Male adult
S Slovakia, Jurský Chlm; 10mm
Copyright © Aleš Sedláček

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) maculicornis (Duftschmid, 1812)
Male adult
Spain, Guipuzcoa, Irun, Playaundi, 28.5.2006, Anichtchenko A leg. & det. (cAA) 6.5 mm
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) hispanus (Rambur, 1838)
Male adult
Spain, Andalicia, San Pedro de Alcantara, 7.4.2012, Anichtchenko A. leg. & det. (cAA) 7.5 mm
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) hespericus Jeanne, 1985
Male adult
Spain, Cadiz, Paterna de Rivera, 9.11.1996 (DUBC) 9 mm
Copyright © Alexander Anichtchenko

Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)
Male adult
S Tadzhikistan, Kurgan-Tube reg., 20km S Dzhilikul, 11.5.1999 (DUBC) 8.5mm
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) antoinei (Schauberger, 1931)
Male adult
Spain, Andalucia, Ubrique, 7.1.2000 (DUBC) 6.5mm
Copyright © Alexander Anichtchenko

Trichotichnus (Trichotichnus) laevicollis (as carpathicus Schauberger, 1936)
Male adult
W Ukraine, NE Carpathians Mts.,
Transcarpathian reg., Rakhov Distr., Chornogora range, S slope of Petros Mt., h~1650m, 28.VI.2009, Panin R.leg.
L=8 mm
Copyright © Ruslan Panin

Trichotichnus (Trichotichnus) laevicollis (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Typsiharpalus bonvouloiri (Vuillefroy, 1866)
Female adult
Spain, Sierra de Guadarrama, Cotos, 15-7-2009, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko