Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Endophallus
endophalus
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Endophallus
Jaen reg., Monte Javalcuz h=1436 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Endophallus
NC-Spain, Soria, Puerto de Oncala
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186
Endophallus
CW Spain, Province Huelva, Arroyomolinos de Leon.
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Endophallus
WC Spain, Cáceres, Hervás, h=1000m
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840
Endophallus
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m
det.Panin R.
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri baguenai Breuning, 1926
Endophallus
SE Spain, Almería Province, Carboneras env.
det. R.Panin
Copyright © Ruslan Panin

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Endophallus
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m,
Copyright © Ruslan Panin

Carabus (Carabus) arvensis deyrollei Gory, 1839
Endophallus
NW-Spain, Cantabria, Udías env., Monte Corona h=100m
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus latus Dejean, 1826
Endophallus
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Endophallus
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Endophallus
CW-Spain:Salamanca, Brincones env., h=750m
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Endophallus
NE Spain, Catalonia,Girona Prov., San Hilario Sacalm env.<br />
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996)
Endophallus
NW-Spain, Leon prov., Sierra de Gistredo, Pto. de Magdalena,
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Endophallus
NC-Spain, Basque Co, Álava, Sierra de Gorbea.
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
Endophallus
SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h=1300 m,
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Endophallus
NW Portugal, Viana do Castelo distr., Sierra de Agra
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Endophallus
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Endophallus
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Endophallus
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m
Copyright © Ruslan Panin

Carabus (Hemicarabus) nitens Linne, 1758
Endophallus
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Endophallus
Collection ground Mukawa cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) semipunctatus Dejean, 1829
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) nevadensis K. Daniel & J. Daniel, 1898d: 65
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) ebeninus Heyden, 1870
Endophallus

Copyright © Alexander Anichtchenko

Harpalus (Cryptophonus) litigiosus Dejean, 1829
Endophallus
Spain, Andalusia, Sierra Arana, 25 km NE Granada, near cueva del Agua, 16.x.2003, 1700m, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Platytarus gracilis gracilis (Dejean, 1831)
Endophallus

Copyright © Alexander Anichtchenko

Dromius (Dromius) agilis (Fabricius, 1787)
Endophallus
Karachay-Cherkessia, gorge. Dukka, h~2100 3-5.VII.2020 leg. M. Nabozhenko
Copyright © Kirill Makarov

Platyderus (Platyderus) barsevskisi Anichtchenko, 2010
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) zaballosi Anichtchenko, 2011 Holotype
Endophallus
Hontanar, Montes de Toledo, Toledo, Toribio
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) zaballosi Anichtchenko, 2011 Holotype
Endophallus
Hontanar, Montes de Toledo, Toledo, Toribio
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) zaballosi Anichtchenko, 2011 Holotype
Endophallus
Hontanar, Montes de Toledo, Toledo, Toribio
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) zaballosi Anichtchenko, 2011 Holotype
Endophallus
Hontanar, Montes de Toledo, Toledo, Toribio
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) varians Schaufuss, 1862
Endophallus
Sierra de Guadarrama, rio de la Guarramillas, 1500-1800m, 28.8.2000
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) varians Schaufuss, 1862
Endophallus
Sierra de Guadarrama, rio de la Guarramillas, 1500-1800m, 28.8.2000
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) varians Schaufuss, 1862
Endophallus
Sierra de Guadarrama, rio de la Guarramillas, 1500-1800m, 28.8.2000
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) varians Schaufuss, 1862
Endophallus
Sierra de Guadarrama, rio de la Guarramillas, 1500-1800m, 28.8.2000
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) lencinai Anichtchenko, 2011 Holotype
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) gazureki Anichtchenko, 2011 Holotype
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) bolivari Anichtchenko, 2011 Holotype
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) itziarae Anichtchenko, 2005 Holotype
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) sagrensis Anichtchenko, 2005 Holotype
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) migelangeli Anichtchenko, 2005 Holotype
Endophallus
Sierra de Cazorla
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) rotundatus Chaudoir, 1866: 111
Endophallus
Granada, Sierra Arana
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) berlovorum Anichtchenko, 2005 Holotype
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) mateui Anichtchenko, 2005 Holotype
Endophallus
Sierra Aitana, Alicante
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) valencianus Anichtchenko, 2005 Holotype
Endophallus
Tavernes de la Valldigna, Valencia
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) marduk Anichtchenko, 2003 Paratype
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) montanellus Graells, 1851
Endophallus
Berzosa de Lozoya, 11.XI.2000, Madrid
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) gallaecus Jeanne, 1970
Endophallus
Burgueira, Pontevedra
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) salmantinus Jeanne, 1996b: 400
Endophallus
Candelario, Sra. Bejar, 25.3.2002, Salamanca
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) incertans Mateu, 1952
Endophallus
Espejo de Tera, 18,VIII, 1997, Soria, M.Toribio leg.
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) incertans Mateu, 1952
Endophallus
Cameros
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) saezi Vuillefroy, 1868: 293
Endophallus
Hervas (Cc.), 1.VI.1990, E. Carabajal leg. (cAA).
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) speleus Cobos, 1961
Endophallus
rio Verde, 3 km. O de Otivar, Sierra Almijara, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) marianicus Ruiz-Tapiador & Anichtchenko, 2007 Holotype
Endophallus
Puerto de Valderrepisa, Sierra Madrona
Copyright © Alexander Anichtchenko

Calathus (Neocalathus) melanocephalus melanocephalus
Endophallus
Left - Spain, Montes Cantabricos; right – paratype of C. (N.) anistschenkoi syn.
Copyright © Alexander Anichtchenko

Platyderus (Platyderus) coiffaiti Jeanne, 1996 Paratype
Endophallus

Copyright © Alexander Anichtchenko

Platyderus (Platyderus) toribioi Anichtchenko, 2005 Paratype
Endophallus
Estepona, Los Reales
Copyright © Alexander Anichtchenko

Amara (Zezea) plebeja (Gyllenhal, 1810)
Endophallus
Collection ground Chitose city, Hokkaido Pref. pref. 34882
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) familiaris (Duftschmid, 1812)
Endophallus
Collection ground Kiyosato cho, Hokkaido Pref. pref. 38785
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) communis (Panzer, 1797)
Endophallus
det. K. Makarov
Copyright © Kirill Makarov

Poecilus (Ancholeus) baeticus baeticus
Endophallus
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Poecilus (Poecilus) crenulatus crenulatus (Dejean, 1828)
Endophallus
Spain, Leon, Qiuntanilla de Samoza, 3.IV.2004, J.L. Zapata leg.
Copyright © Alexander Anichtchenko

Poecilus (Parapedius) decipiens decipiens
Endophallus
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Poecilus (Poecilus) cupreus (Linne, 1758)
Endophallus
Turkije : Ardahan; Ardahan 7KM E Karst 13-7-1992 1900m. Heinz W. leg.
Copyright © Jan Muilwijk

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Endophallus
Agri, Balık Gölü, Taşlıçay, 2250 m, 17.07.76; W. Heinz leg.
Copyright © Jan Muilwijk

Orthomus (Orthomus) perezii (Martinez y Saez, 1873)
Endophallus
Spain, Cuenca, Montes Universales, pto del Cubillo 1620m, 23.5.2006, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Amara (Leironotus) rotundicollis (Schaufuss, 1862)
Endophallus
Spain, Sierra Guadarrama, 2 km E Navacerrada 1900m, 1.9.2002, Anichtchenko A. leg. & det. (cAA).
Copyright © Alexander Anichtchenko

Trechus (Trechus) bouilloni Faille, Bourdeau & Fresneda, 2012 Paratype
Endophallus
Spain, Navarra, Sierra de Urbasa–Andнa, Lizarraga, Puerto de Lizarraga, MSS, trap: 1–5–1980/15–8–1980, Bourdeau and Fresneda leg.
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---

Trechus (Trechus) grenieri Pandelle, 1867
Endophallus
Nistos, Bas–Nistos, Grotte de l’Eglise (Bonadona 1971, Corbaz & Jauzion 1988)
Photo Faille, Bourdeau & Fresneda, 2012
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Trechus (Trechus) uhagonii uhagonii s.str.
Endophallus
Cueva de Orobe
Photo Faille, Bourdeau & Fresneda, 2012
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Trechus (Trechus) sharpi Jeannel, 1921c: 179
Endophallus
Quijas, Cueva la Cuevona, 13–IX–1995
Photo Faille, Bourdeau & Fresneda, 2012
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Trechus (Trechus) beusti (Schaufuss, 1863)
Endophallus
Zegama, Cueva de San Adrián
Photo Faille, Bourdeau & Fresneda, 2012
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Trechus (Trechus) pieltaini Jeannel, 1920e: 155
Endophallus
Cueva de Mairuelegorreta
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---

Trechus (Trechus) bruckii Fairmaire, 1862a: 548
Endophallus
Gourette, Lac d’Anglas, 20–8–2000, Bourdeau leg.
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---