Poecilus (Poecilus) cupreus Linne, 1758
Female adult
W Ukraine, Lviv Reg., Javorov distr. near Ivano-Frankovo vill., Panin R. leg., 20.IV-4.V.2008 L=13mm
Copyright © Ruslan Panin

Poecilus (Poecilus) lepidus (Leske, 1785)
Male+Female adult

W. UKRAINE, NW.Podolian,
Ternopil Reg., Pidvolochisk distr., near Rosokhuvatets vill., in steppe, 24.V-15.VI.08 Panin R. leg.
female-15 mm

Copyright © Ruslan Panin

Poecilus (Poecilus) versicolor Sturm, 1824
Male adult

Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornogora range, h~1850m.29.VI-28.VII.07 Panin R. leg.
L=12mm

Copyright © Ruslan Panin

Poecilus (Poecilus) cupreus Linne, 1758
Female adult
<p> length: 9-13 mm<br /> Germania, Bayern, Oberfranken, Bindlach<br /> leg.det. U.Schmidt, 1978</p>
Copyright © U.Schmidt

Poecilus (Poecilus) versicolor Sturm, 1824: 99 (Platysma)
Male adult
L=10,3 mm
RUSSIA, Moscow reg., Istrinskiy distr., Snegiri settl. 23.V.2009
leg. ?
det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Male adult

Copyright © Kirill Makarov

Poecilus (Poecilus) kugelanni (Panzer, 1797)
Male adult
SE Slovakia, Slovenský kras; 13mm
Copyright © Aleš Sedláček

Poecilus (Poecilus) quadricollis (Dejean, 1828)
Male adult
S Spain, Granada; 11mm
Copyright © Aleš Sedláček

Poecilus (Poecilus) zaballosi Jeanne & Ruiz-Tapiador, 1996: 464
Male adult
Spain, 29.09. 2014, alrededores de la laguna de las Yeguas, Alcázar de San Juan (Ciudad Real). Image of Víctor Manuel Ramos Sánchez-Mateos
Copyright © - - -

Poecilus (Poecilus) cupreus Linne, 1758: 416 (Carabus)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Poecilus (Carenostylus) purpurascens (Dejean, 1828)
Female adult
Spain, Andalucia, San Pedro de Alcantara, 7.4.2010, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Poecilus (Ancholeus) baeticus baeticus
Male adult
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Poecilus (Poecilus) crenulatus crenulatus (Dejean, 1828)
Male adult
Spain, Leon, Qiuntanilla de Samoza, 3.IV.2004, J.L. Zapata leg.; det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Poecilus (Poecilus) tyrrhenicus Csiki, 1930
Female adult
Spain, Cadiz, Chiclana, 28.X.1999, P. Coello leg. det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Poecilus (Parapedius) decipiens decipiens
Male adult
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Poecilus (Ancholeus) gisellae gisellae
Female adult
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Poecilus (Ancholeus) baeticus baeticus
Endophallus
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Poecilus (Poecilus) crenulatus crenulatus (Dejean, 1828)
Endophallus
Spain, Leon, Qiuntanilla de Samoza, 3.IV.2004, J.L. Zapata leg.
Copyright © Alexander Anichtchenko

Poecilus (Parapedius) decipiens decipiens
Endophallus
Spain, San Roque, Cortijo de la Ventilla, 4.IV.2003, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Poecilus (Poecilus) cupreus (Linne, 1758)
Male adult
Turkije : Ardahan; Ardahan 7KM E Karst 13-7-1992 1900m. Heinz W. leg.
Copyright © Jan Muilwijk

Poecilus (Poecilus) cupreus (Linne, 1758)
Endophallus
Turkije : Ardahan; Ardahan 7KM E Karst 13-7-1992 1900m. Heinz W. leg.
Copyright © Jan Muilwijk

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Endophallus
Agri, Balık Gölü, Taşlıçay, 2250 m, 17.07.76; W. Heinz leg.
Copyright © Jan Muilwijk

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Male genitalia
1 - Iran, Kodir. 2 - Iran, Chaharmahal and Bakhtiari, Sulegan lagoon, Cheshmeye Gholamhossein, 28.v.2019, 31° 57ˊ57 ̋ N, 50° 49ˊ27 ̋ E, J. Muilwijk leg.
Copyright © Jan Muilwijk

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Female adult
Israel
Copyright © Jan Muilwijk

Poecilus (Ancholeus) aeneolus (Chaudoir, 1868) Holotype
Male adult
MNHN
Copyright © Jan Muilwijk

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Male adult
Iran, Chaharmahal and Bakhtiari, Sulegan lagoon, Cheshmeye Gholamhossein, 28.v.2019, 31° 57ˊ57 ̋ N, 50° 49ˊ27 ̋ E, J. Muilwijk leg
Copyright © Jan Muilwijk

Pterostichus oblongopunctatus (Fabricius, 1787: 202)
Male adult
Pterostichus (Bothriopterus) oblongopunctatus
Chelyabinskaya obl., Katav-Ivanovskiy prov., mountain Maliy Iremal, alt. 1180 m. 19/08/2008.
Leg. E.Chibilov
Copyright © Sunvas

Pterostichus (Adelosia) macer Marsham, 1802
Male adult

E. Ukraine,
Kharkov reg., 1972
16mm.

Copyright © Ruslan Panin

Pterostichus (Adelosia) macer Marsham, 1802
Male adult

Length: 15.4 mm
West Kazakhstan,
Uralskaya area, nearby Volodarskoye village, sands.
15.04.2000
F.G.Bidashko leg. and det.

Copyright © Alexander Anichtchenko

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult
SE Siberia, Irkutsk Area, near Bratsk city, 30.VII.07, Fominyh D. leg. long=13mm.
Copyright © Ruslan Panin

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult
<p> long=11.5mm</p>
Copyright © Ruslan Panin

Pterostichus (Feronidius) melas (Creutzer, 1799)
Male adult

W-Ukraine,
Lviv Region, Zolochiv distr., near Gologirky, in steppe, 16-29.V.2007 Panin R. leg.
long-17mm

Copyright © Ruslan Panin

Pterostichus (Platysma) niger niger s.str.
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.2007 Panin R. leg.

Copyright © Ruslan Panin

Pterostichus (Platysma) niger niger s.str.
Male adult

length: 22,8 mm
West Siberia, KHMAO-Yugra, the vicinities of Khanty-Mansiysk town
1.07.2004.
Zinoviev E.leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Platysma) niger niger s.str.
Male adult

Chelyabinsk, 4.03.2011

Chibilov leg.

Copyright © Sunvas

Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)
Male adult
W UKRAINE, Lviv Reg., near Kamenka-Bugskaya, in forest under the bark oak 29.III.08 Panin R. leg. L=12mm
Copyright © Ruslan Panin

Pterostichus (Pseudomaseus) gracilis (Dejean, 1828)
Male adult

length: 9,3 mm
Russia, West Siberia, KHMAO-Yugra, Khanty-Mansiysk distr., Ob river, Bolshoy Tchukhtinsiy island
4.08.2007.
Zinoviev E.leg

Copyright © Evgeniy Zinoviev

Pterostichus (Troglorites) breuili mendizabali Jeannel, 1921
Male adult
Spain, Guipuzcoa, Albistur, (Tolosa) cueva Mendicute, M.Toribio leg.

Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) melanarius (Illiger, 1798)
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Pterostichus (Morphnosoma) melanarius (Illiger, 1798)
Female adult
W Ukraine,<br /> Ternopol Reg., Pidvolochisk distr., forest Bogdanivskyi 24.V-15.VI.08 Panin R. leg.<br /> long =19 mm
Copyright © Ruslan Panin

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Female adult
photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Pterostichus (Bothriopterus) oblongopunctatus (as honshuensis Habu et Baba, 1958) Holotype
Male adult
Mt. Monnai, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-038
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Bothriopterus) oblongopunctatus (as honshuensis Habu et Baba, 1958) Holotype
Male adult
Mt. Monnai, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-038
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Alexander Anichtchenko

Pterostichus (Bothriopterus) oblongopunctatus (as honshuensis Habu et Baba, 1958) Holotype
Male adult
Mt. Monnai, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-038
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Bothriopterus) oblongopunctatus (as honshuensis Habu et Baba, 1958) Holotype
Male adult
Mt. Monnai, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-038
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Steropus) ebenus (Quensel, 1806)
Male adult
SC Spain, Jaen provincia, Linares, 30.IX.2001, M.A. Lopez Vergara leg.
L=21 mm
Copyright © Ruslan Panin

Pterostichus (Troglorites) breuili mendizabali Jeannel, 1921
Male adult
Spain, Guipuzcoa, Albistur, (Tolosa) cueva Mendicute, M.Toribio leg.
10 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Female adult
France, Sainte Engrace, (64) 27-05-2007, Pascal Deschamps coll. & det.
Copyright © Pascal Deschamps

Pterostichus (Argutor) vernalis Panzer, 1796
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Argutor) cursor Dejean, 1828: 243
Female adult
det. K. Makarov
Copyright © Alexander Anichtchenko

Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) strenuus (Panzer, 1796)
Female adult
Russia, Moscow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Oreophilus) xatartii (Dejean, 1828)
Male adult
France : Pyrénées or., Ft. du Carcanet-9.x.1971, P. Picard Leg. - Coll.RBINS, N. Mal
Copyright © Noël Mal

Pterostichus (Cryobius) abaxoides abaxoides (Dejean, 1828)
Male adult
Spain, C Pirineos, Pleta Llardana, 2600m, 23.8.2005, Anichtchenko A. leg. & det. ; 10mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) pumilio (Dejean, 1828)
Male adult
Spain, Navarra, Sierra de Aralar, 9.5.2006, Altxueta, 1000m, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) aralarensis Espanol & Mateu, 1945
Female adult
Spain, Mts, Cantabricos, Soto de Sajambre, Canto Cabronero Mt., 1850m, 12.VIII.2008, Anichtchenko A. leg. & det. 7.5mm
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) cristatus cristatus
Male adult
France, 77860, Quincy-Voisins, 20-06-2009
Copyright © Pascal Deschamps

Pterostichus (Troglorites) breuili breuili Jeannel, 1919
Male adult
Alli, Navarra, Spain, Marc Curti leg.
Copyright © Jean Michel Lemaire

Pterostichus (Cryobius) pusillus sagittalis Jeannel, 1937a: 18
Female adult
Spain, C Pirineos, Pleta Llardana, 2600 m, 23.8.2005, Anichtchenko A. leg; 5.8 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) ehlersi (Heyden, 1881)
Male adult
Spain, Mt. Cantabricos, Picos de Europa, 2200 m, 3 km NW mirador Fuente De, 22.07.2019, A. Anichtchenko & R. Sciaky leg. 9 mm. det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) aralarensis asturicus Jeanne, 1969a: 3
Male adult
Spain, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg. det. Anichtchenko A. 7.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) cantabricus cantabricus (Schaufuss, 1862)
Male adult
Spain, Cantabria, Pico Tres Mares, 2000-2100m, 24.07.2019, A. Anichtchenko & R. Sciaky leg.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) procerulus (Heyden, 1880)
Male adult
Spain, Peña Ubiña, Montes Cantabricos, 2000 m, 28.6.2009, Anichtchenko A. leg. 8 mm.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) cantabricus vasconicus Aubry, 1963
Male adult
Spain, Sierra de Urbasa, 900-1000 m, 27.5.2006, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Oreophilus) franzi Negre, 1955
Male adult
SPAIN, Cantabria, 3 km SW Fuente De, Cantijon riv., 1400 m, 21.07.2019, A. Anichtchenko & R. Sciaky leg.
Copyright © Alexander Anichtchenko

Pterostichus (Pseudomaseus) gracilis (Dejean, 1828)
Female adult

Copyright © Kirill Makarov

Pterostichus (Feronidius) melas italicus Dejean, 1828: 274
Male adult

Copyright © Alberto Narro

Pterostichus (Feronidius) melas italicus Dejean, 1828: 274
Male genitalia

Copyright © Alberto Narro

Pterostichus (Troglorites) breuili mendizabali Jeannel, 1921
Female adult

Copyright © Vittorino Monzini

Pterostichus (Troglorites) breuili breuili Jeannel, 1919
Male adult

Copyright © Vittorino Monzini

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult

Copyright © Kirill Makarov

Pterostichus (Pterostichus) dux L.Schaufuss, 1862
Female adult
Spain, Mts Cantabricos, Macizo Central, Picu Urriellu, 2000m, 13.8.2008 (cAA) leg & det. Anichtchenko A.; 18 mm;
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) insidiatrix (Brulerie, 1872)
Male adult
Spain, Teruel, Montes Universales, Pto del Portillo 1780m 31.5.2006, Anichtchenko A. leg & det. (cAA) 19mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) ghilianii Putzeys, 1846
Male adult
Spain, Sierra de Guadarrama, pto Somosierra, 1500m 3.6.2006, Anichtchenko A. leg & det. (DUBC) 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) ghilianii Putzeys, 1846
Female adult
Spain, Sierra de Guadarrama, pto Somosierra, 1500m 3.6.2006, A. Anichtchenko leg & det. (DUBC). 16 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Female adult
Spain, Navarra, Bera de Bidasoa, Larrun Mt. 850m 9-15.9.2001, A. Anichtchenko leg. & det. (cAA) 17 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) madidus (Fabricius, 1775)
Male adult
Belgium, Limburg, Forest reserve of Kolmont, 24.4.2016 (DUBC). det. Anichtchenko A., 2021. 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Male adult
Spain, Navarra, Bera de Bidasoa, Larrun Mt. 850m 9-15.9.2001, A. Anichtchenko leg. & det. (cAA) 19 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Male adult
Spain, Asturias, Rioseco, 11.8.87 (DUBC) 16 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) pusillus Dejean, 1828
Female adult
Hautes-Pyrénées Department, Vallon d'Estaragne, 42°48'15.93"N, 0° 8'1.36"E, 2580 m, July 2021, P. Moret leg. & det.
Copyright © Pierre Moret

Pterostichus (Cryobius) pumilio (Dejean, 1828)
Male adult
Hautes-Pyrénées Department, Vallon d'Estaragne, 42°48'49.70"N, 0° 8'45.07"E, 2130 m, July 2021, P. Moret leg. & det.
Copyright © Pierre Moret

Pterostichus (Cryobius) amoenus (Dejean, 1828)
Male adult
Hautes-Pyrénées Department, Cirque de Troumouse, 42° 42' 48.7" N, 0° 06' 43.0" E, 2370 m, July 2017, P. Moret leg. & det.
Copyright © Pierre Moret

Pterostichus (Oreophilus) cantaber (Chaudoir, 1868)
Male adult
Spain, Santander, Loredo, 8.8.1993 (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) colasi Jeannel, 1937
Male adult
Spain, Lleida, Vielha, Panta de Senet, Barranco de Salenca. Image from Sedlmeier and Faille 2022: First insights into the phylogeny of Cryobius
Copyright © - - -

Pterostichus (Lianoe) arribasi Ortuño, 1991
Male adult
Ezcaray, 16.8.97, Valdezcaray, La Rioja, Spain, M. Toribio leg. & det. (cAA). 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) amaroides Dejean, 1828
Female adult
Andorra, 6.1983, Monzini leg. (cAA). 8 mm
Copyright © Alexander Anichtchenko

Pterostichus (Feronidius) brevipennis brevipennis
Female adult
Spain, Bueu, Pontevedra, 6.8.1996 (cAA). 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Feronidius) brevipennis sousae (Vuillefroy, 1868)
Male adult
Spain, Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.7.2008, Anichtchenko A. leg & det. (cAA). 13 mm
Copyright © Alexander Anichtchenko

Pterostichus (Melanius) elongatus (Duftschmid, 1812)
Female adult
Marruecos, Gada de Debdou, 9.6.1993 (cAA)
Copyright © Alexander Anichtchenko

Stomis (Stomis) pumicatus (Panzer, 1796)
Male adult
Canada; Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Stomis (Stomis) pumicatus (Panzer, 1796)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Styracoderus atramentarius Rosenhauer, 1856: 34
Female adult
Spain, Sierra Nevada, Puerto de la Ragua, 26.5.2004, 2000m, Anichtchenko A. leg. & det.
13 mm
Copyright © Alexander Anichtchenko

Styracoderus martinezi (Vuillefroy, 1868)
Female adult
Spain, Sierra Aitana, 1200m, Alcoleja, Alicante, 23.4.1984 (cAA) 15mm
Copyright © Alexander Anichtchenko

Styracoderus azarai Perez Arcas, 1865
Male adult
Spain, Candelario, Sierra de Bejar, Salamanca 15.8.1987 (cAA) 12 mm
Copyright © Alexander Anichtchenko