Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Endophallus
France 39, Foncina le Haut 300m
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Endophallus

Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Endophallus
W Ukraine, NW Podolian, Lviv reg., Javoriv distr.,near Stradch vill.,
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Endophallus
Czech, S Moravia, Pouzdrany
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis hortensis Linne, 1758
Endophallus
W-Ukraine, Male Polissya, near Kamenka-Buzka vill.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus germarii Sturm, 1815
Endophallus
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Endophallus
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh.
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei linnaei Panzer, 1812
Endophallus
W.UKRAINE, Lviv Reg., NE Carpathians, Skolivski Beskidy Mts. Zelemyanka Mt. Rng
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Endophallus
Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) nitens Linne, 1758
Endophallus
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov

Carabus (Orinocarabus) sylvestris sylvestris (as micklitzi Sokolar, 1910)
Endophallus
W Austria, Styria, Mt Hochwechsel W slope, 47.531667, 15.906000 h=1640 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri Panzer, 1799
Endophallus
E-Austria, Wienerwald, Hochstraß am Lengbachl, h=575 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri (as floriani Penecke, 1905)
Endophallus
SE-Austria, Styria, Rein bei Graz VI.1970, R.Kenyery leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri (as floriani Penecke, 1905)
Endophallus
SE-Austria, Styria, Gratwein Kehrerwaldh=580 m, 47.130800, 15.270800
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri bergeri Peeters & I.Rapuzzi, 2025
Endophallus
W-Austria,Tyrol, Vergötschen (Kaunertal), Feichten, h= 1300 m
Copyright © Ruslan Panin

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Endophallus
Collection ground Mukawa cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Endophallus
South Dagestan, 5 km SE of Dzhalgan village, h~50 S of the Sabnovo-Dzhalgansky ridge, 41°59'11"N 48°17'47"E 7.1.2026 leg. O. and K. Makarov's
Copyright © Kirill Makarov

Dromius (Dromius) agilis (Fabricius, 1787)
Endophallus
Karachay-Cherkessia, gorge. Dukka, h~2100 3-5.VII.2020 leg. M. Nabozhenko
Copyright © Kirill Makarov

Calathus (Neocalathus) micropterus (Dufischmid, 1812)
Endophallus
Sakhalin isl.
Copyright © Alexander Anichtchenko

Amara (Zezea) plebeja (Gyllenhal, 1810)
Endophallus
Collection ground Chitose city, Hokkaido Pref. pref. 34882
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Bradytus) majuscula (Chaudoir, 1850)
Endophallus
Collection ground Monbetsu cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) familiaris (Duftschmid, 1812)
Endophallus
Collection ground Kiyosato cho, Hokkaido Pref. pref. 38785
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) communis (Panzer, 1797)
Endophallus
det. K. Makarov
Copyright © Kirill Makarov

Poecilus (Poecilus) cupreus (Linne, 1758)
Endophallus
Turkije : Ardahan; Ardahan 7KM E Karst 13-7-1992 1900m. Heinz W. leg.
Copyright © Jan Muilwijk

Amara (Amara) pulpani Kult, 1949
Endophallus
W Ukraine, NE Carpathians, Lviv reg., Skole Beskids Mts, Grabnyk Range h=670 m,
Copyright © Denis Khrapov

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Endophallus
Agri, Balık Gölü, Taşlıçay, 2250 m, 17.07.76; W. Heinz leg.
Copyright © Jan Muilwijk

Amara (Bradytus) majuscula (Chaudoir, 1850)
Endophallus
Buryatia, Tunkinskii Distr., environs of Tunka Vill. 3.VIII.2012 leg. A. Prosvirov
Copyright © Kirill Makarov

Amara (Bradytus) fulva (O.F. Muller, 1776)
Endophallus
Altai Terr., Smolenskii Distr., 5 km S of Smolenskoe Vill., Peschanaya River bank, 31.VII–2.VIII.2000 leg. A. Matalin
Copyright © Kirill Makarov

Amara (Bradytus) crenata Dejean, 1828: 507
Endophallus
Stavropol Terr., Izobil’nenskii Distr., Moskovskoe Vill. 4(17).III.1922 (collector unknown)
Copyright © Kirill Makarov

Amara (Bradytus) consularis (Duftschmid, 1812)
Endophallus
Armenia, mountains W of Lake Arpi, h ~ 2000 m, 8.VI.2002 leg. A. Rubenyan
Copyright © Kirill Makarov

Amara (Bradytus) apricaria (Paykull, 1790)
Endophallus
Kyrgyzstan, Moldy-Too Mt. Range, Kurtka River, h ~ 2200 m, 20–21.VII.1998 leg. A. Klimenko
Copyright © Kirill Makarov