Amblystomus metallescens (Dejean, 1829a)
Female adult

Copyright © Kirill Makarov

Amblystomus metallescens (Dejean, 1829)
Male adult
France, Camargue, Bouche du Rhône. Image from Wrase & Magrinii, 2012
Copyright © - - -

Amblystomus niger (Heer, 1841)
Male adult
Italy, Toscana, Padule di Fucecchio, Pistoia. Image from Wrase & Magrinii, 2012
Copyright © - - -

Carterus (Pristocarterus) angustus (Menetries, 1832)
Male adult
Azerbaijan, Bozdagh range,Turianchai Nature Reserve, 07.VII.2008 Pljushch I.G. leg. L=8mm
Copyright © Ruslan Panin

Ditomus calydonius oriens (Dvorak, 1993)
Male adult
S Kazakhstan, Karatau mts.,<br /> Buylikkul lake 26.6.1999<br /> 16 mm
Copyright © Alexander Anichtchenko

Ditomus calydonius oriens (Dvorak, 1993)
Male adult
length: 15 mm
Turcia, Troja
leg. U.Schmidt, 1997; det. A.Skale
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Dixus eremita Dejean, 1825
Male adult
Volgogrd Area, Pallasovka Distr., 5 km NNW Elton Vill., Sorochya Balka 8.IV.2007 leg. A.Matalin
L=9.0
Copyright © Kirill Makarov

Eucarterus sparsutus Reitter, 1898f: 338
Male adult

Copyright © Kirill Makarov

Carterus (Pristocarterus) angustipennis lutshniki Zamotajlov, 1988
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carterus (Carterus) dama (P.Rossi, 1792)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ditomus calydonius oriens (Dvorak, 1993)
Male adult

Copyright © Kirill Makarov

Carterus (Pristocarterus) angustus (Menetries, 1832)
Female adult
Iran, Lorestan prov., near Dorud, 5 km SW Chabdar vill. 33.501061°N 49.003441°E 14-15.V.2017 leg. D.Kasatkin L=8.5 mm
Copyright © Kirill Makarov

Carterus (Pristocarterus) angustus (Menetries, 1832)
Male adult
Iran, Lorestan prov., near Dorud, 5 km SW Chabdar vill. 33.501061°N 49.003441°E 14-15.V.2017 leg. D.Kasatkin L=8.5 mm
Copyright © Kirill Makarov

Oedesis caucasicus (Dejean, 1831)
Female adult
Italy, Veneto, Rosara di Codevigo, alla lampada UV, 10-12.V.2017; 8.1 mm. Image from Zanella & Uliana, 2022
Copyright © - - -

Harpalus rufipes (Degeer, 1774)
Male adult
Russia, S.Ural
Copyright © Sunvas

Acinopus (Acinopus) laevigatus Menetries, 1832: 128
Male adult
Azerbaijan, 01.VII.2008, Pljushch I.G. leg.
female,14.5mm
Copyright © Ruslan Panin

Acinopus (Acinopus) picipes (Olivier, 1795)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr., near Bogatoe Ushchele vill. 04.VIII.2009 Shehovcov A. leg.
L=16 mm
Copyright © Ruslan Panin

Acinopus (Osimus) ammophilus Dejean, 1829
Male adult
S Ukraine, Krym Peninsula,
Shchelkino distr., near Mysovoe vill., 03.VIII.2008V. Shaporinskiy leg.
23mm
Copyright © Ruslan Panin

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
NW Podolian Upland, W UKRAINE,<br /> Lviv Reg., Zolochiv distr., near Gologirki vill., 30.04-19.05.07 Panin R.leg.<br /> L= 15mm
Copyright © Ruslan Panin

Harpalus (Harpalus) affinis (Schrank, 1781)
Male adult
length: 9-12 mm
Helvetia, Graubünden, Ilanz
leg.det. U.Schmidt, 2004
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Harpalus (Harpalus) caspius (Steven, 1806)
Male adult
W UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa vill., Panin R. leg. 1.V-16.VI.2007
male, 13mm
Copyright © Ruslan Panin

Harpalus (Harpalus) fuscipalpis Sturm, 1818
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Harpalus) honestus (Dufischmid, 1812)
Male adult
length: 7,4-10,6 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2007
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Harpalus (Harpalus) mitridati Pliginskiy, 1915
Male adult
Rostov Area, Kumyska Vill. 24.III.2006 leg. M.Nabozhenko
L=8.5
Copyright © Kirill Makarov

Harpalus (Pseudoophonus) rufipes (Degeer, 1774)
Male adult
W-UKRAINE,
Lviv Region, Javorov distr., near Ivano-Frankovo 1-20.VI.07 Panin R. leg
long = 17mm
Copyright © Ruslan Panin

Harpalus (Harpalus) distinguendus (Duftschmid, 1812)
Female adult
POLAND; CE 20 (Lower Vistula Valey)
Bruki Unislawskie
25.04.2009
leg. M. Traczyk
Photo of Mieczyslaw Stachowiak
Copyright © Mieczyslaw Stachowiak

Harpalus (Harpalus) latus (Linne, 1758)
Male adult
length: 10,2 mm
Russia., Sverdlovskaya area, the vicinities of Ekaterinburg city, Verkhnemakarovo vill., floodplain meadow
6.07.2005
Zinovyev E. leg.
Copyright © Evgeniy Zinoviev

Harpalus (Harpalus) progrediens Schauberger, 1922
Male adult
length: 8,4 mm
Russia., Sverdlovskaya area, Beloyarskyi distr., Chernousovo vill.
2.09.2007
Vlasov V. leg.
Copyright © Evgeniy Zinoviev

Ophonus (Metophonus) gammeli (Schauberger, 1932)
Male adult
Stavropol Reg., env. Zheleznovodsk 1.06.2007 S.Svetlov leg.
male
L=11,0 mm
Copyright © Maksim Smirnov

Ophonus (Metophonus) puncticeps Stephens, 1828
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Male adult
length: 6,0-9,1 mm Germania, Bayern, Pegnitz<br /> leg.det. U.Schmidt, 1972<br /> Photo: U.Schmidt, 2008
Copyright © U.Schmidt

Trichotichnus (Trichotichnus) nishioi Habu, 1961
Male adult
Far East, Lazo Distr., midd. course of Socolovka Riv. 42°58'35"N 133°59'29E 28.VII.2005 K.Makarov leg.
L=8.5 mm
Copyright © Kirill Makarov

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Harpalus (Harpalus) foveiger Tschitscherine, 1895
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) egorovi Lafer, 1989
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) amplicollis Menetries, 1848
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) brevicornis Germar, 1824
Male adult

Copyright © Kirill Makarov

Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) distinguendus (Duftschmid, 1812)
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Harpalus (Loboharpalus) rubefactus Bates, 1873
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) coruscus (as sachalinensis) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) nishioi Habu, 1961 Holotype
Male adult
Kotoni, HOKKAIDO, 03. V. 1955, Y. Nishio. COL-138
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) nishioi Habu, 1961 Holotype
Male adult
Kotoni, HOKKAIDO, 03. V. 1955, Y. Nishio. COL-138
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) tarsalis (as hokkaidensis) Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) tarsalis Mannenheim, 1825: 28 Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) tarsalis Mannenheim, 1825: 28 Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes (as ainus) Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) serripes (Quensel, 1806)
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) lumbaris Mannerheim, 1825: 27
Male adult

Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) lucidus (A. Morawitz, 1863)
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Male adult
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Male adult
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Male adult
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Male adult
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Male adult
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Male adult
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) tarsalis (as hokkaidensis Habu et Baba, 1963)
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39250
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883: 239
Male adult
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Male adult
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Female adult
Russia
Copyright © Kirill Makarov

Harpalus (Loboharpalus) platynotus Bates, 1873
Male adult
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Male adult
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) calathoides Motschulsky, 1844: 200
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Male adult
NW Caucas, Krasnodar reg., SE Goryachii Kluch, леса между пос. Асфальтовая Гора и Садовый (нежилой) 21.V.2012 leg. М. Смирнов
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Daptus pictus Fischer von Waldheim, 1823
Male adult
Kazakhstan, 20 km NW Akkuduk vill., N 43*06'58 E 054*11'46, h=90m, 24.V.2011, Ivanov A. leg.
det. Anichtchenko A. 2013
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) dispar Dejean, 1829
Male adult
France, L’Albaron, 13123 Pascal 9/10/2001, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) smaragdinus (Duftschmid, 1812)
Male+Female adult
France, forêt de Saint Germain en Laye 78100, 30/8/85, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) anxius (Duftschmid, 1812)
Male+Female adult
France, Forêt de Fontainebleau 77300 18/6/88, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) oblitus Dejean, 1829a: 273
Male+Female adult
France, Brétignolles sur mer 85470 17/5/85, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) neglectus Audinet-Serville, 1821
Female adult
France, Ile d’Oléron (17) 12/6/2006, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) pumilus Sturm, 1818: 77
Male+Female adult
France, forêt de Fontainebleau (77300), 15/7/1985, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) tardus (Panzer, 1796)
Male+Female adult
France, forêt de Fontainebleau (77300), 18/4/1987, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Male adult
Portugal, Algarve Portimao, 25/9/1983, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Harpalus) crates Bates, 1883
Male adult
SE Maritime Prov., Lazovsky dist., Lazo env., 17-20.VI.2010, leg. М. Smirnov Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Pangus scaritides (Sturm, 1818)
Male adult
SK, S Slovakia, Velké Kosihy; 8mm
Copyright © Aleš Sedláček

Harpalus (Cephalophonus) cephalotes Fairmaire & Laboulbene, 1854
Male adult
SE Bulgaria, Primorsko; 16mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Female adult
SK, S Slovakia, Veľký Krtíš; 12mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) saxicola Dejean, 1829
Male adult
SE Slovakia, Slovenský kras; 7mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) honestus (Dufischmid, 1812)
Male adult
CZ, NE Moravia, Moravskoslezské Beskydy - Smrk; 9mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) ardosiacus (Lutshnik, 1922)
Female adult
CZ, S Moravia, Mikulov; 10,5mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) diffinis (Dejean, 1829)
Male adult
CZ, SE Moravia, Bílé Karpaty Mts.; 11mm
Copyright © Aleš Sedláček

Ophonus (Hesperophonus) cribricollis (Dejean, 1829)
Male adult
SK, S Slovakia, Kamenica nad Hronom; 8mm
Copyright © Aleš Sedláček

Ophonus (Metophonus) gammeli (Schauberger, 1932)
Male adult
SK, S Slovakia, Štúrovo; 10mm
Copyright © Aleš Sedláček

Ophonus (Metophonus) gammeli (Schauberger, 1932)
Female adult
SK, S Slovakia, Galanta; 11,5mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) stictus Stephens, 1828
Male adult
CZ, SE Moravia, Uherský Brod; 12mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) flavescens (Piller & Mitterpacher, 1783)
Male adult
CZ, S Moravia, Bzenec; 11mm
Copyright © Aleš Sedláček

Acinopus (Acinopus) laevigatus Menetries, 1832: 128
Male adult

Copyright © Kirill Makarov

Acinopus (Acinopus) picipes (Olivier, 1795)
Female adult

Copyright © Kirill Makarov

Acinopus (Osimus) ammophilus Dejean, 1829
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) pulvinatus lubricus Reitter, 1900
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) metallinus Menetries, 1836: 149
Female adult
det. B. Kataev, 2013
Copyright © Kirill Makarov

Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Female adult

Copyright © Kirill Makarov

Acinopus (Oedematicus) megacephalus (P. Rossi, 1794)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
Moravia, Kobylí; 11 mm
Copyright © Aleš Sedláček

Microderes (Microderes) brachypus (Steven, 1809)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) froelichii Sturm, 1818: 117
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) petri Tschitscherine, 1902e: 596
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) servus (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) rubripes (Dufischmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Parophonus (Parophonus) planicollis (Dejean, 1829)
Female adult

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) autumnalis (Duftschmid, 181)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) chrysopus chrysopus
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) circumpunctatus Chaudoir, 1846a: 171
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) oblitus Dejean, 1829a: 273
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) stevenii Dejean, 1829a: 242
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) torridoides Reitter, 1900a: 81
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) terrestris (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Hesperophonus) azureus (Fabricius, 1775)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Hesperophonus) convexicollis Menetries, 1832
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalobrachys leiroides (Motschulsky, 1844)
Female adult
Russia, Burjatia, Tunka valley, Tunka vill. vicinity, 17.VII.1995. Anichtchenko A. leg. 12 mm det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) anxius (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) attenuatus Stephens, 1828a: 152
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) flavicornis Dejean, 1829a: 366
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) cordatus (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) hospes hospes
Male adult
Slovakia, Horné Strháre; 12 mm
Copyright © Aleš Sedláček

Ophonus (Metophonus) puncticeps Stephens, 1828
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) hystrix Reitter, 1894
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) latus (Linne, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) major (Motschulsky, 1850)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) atratus Latreille, 1804a: 360
Male adult
Germany, Bavaria, Eußenheim, F. Boetzl leg. & det. 19.07.14
Copyright © Fabian Bötzl

Harpalus (Harpalus) politus Dejean, 1829a: 370
Male adult
Germany, Bavaria, Reiterswiesen, F. Boetzl leg. 21.05.14, M. Fritze det. 2014
Copyright © Fabian Bötzl

Daptus vittatus Fischer von Waldheim, 1823: pl.46
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) ussuricus Mlynar, 1979: 97
Male adult
Ю Приморье, Хасанский р-н, 7.5 км Ю п.Барабаш, устье руч. Олений 43°07'01"N 131°29'15"E 27.V.2015 leg. К.Макаров L=8.7 mm
Copyright © Kirill Makarov

Harpalus (Harpalus) pusillus (Motschulsky, 1850)
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) pumilus Sturm, 1818: 77
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) rubripes (Dufischmid, 1812)
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) angulatus scytha Tschitscherine, 1899
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) subcylindricus Dejean, 1829a: 374
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) viridanus viridanus (as fuscipalpoides)
Male adult

Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Male adult
det. A. Matalin
Copyright © Maksim Smirnov

Harpalus (Harpalus) hirtipes (Panzer, 1796)
Female adult
13.5 mm; Russia, Volgograd, Kamyshensky, Golubaja river valley, 150 m, leg. E.Alexeev, 15.-19.IV.2000; det. Wrase, 2016. Photo: U.Schmidt, 2017
Copyright © Udo Schmidt

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) distinguendus distinguendus
Female adult

Copyright © Kirill Makarov

Ophonus (Ophonus) stictus Stephens, 1828
Female adult
W Ukraine, NW Podolian Upland, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.V-15.VI.2017, Panin R. & Kanarsky Yu. leg. L=15 mm
Copyright © Ruslan Panin

Neophygas microcephalus (Faldermann, 1835)
Male adult
Mongolia, Ovorhangai aimak, near Hayrhandulaan, 18.7.2006, Anichtchenko A. leg.; 9 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) solitaris Dejean, 1829a: 337
Female adult
W-Ukraine, Transcarpathian reg., Chornagora Massif, 8 km NE Bohdan vill., h=800m, 9-29.VI..2014, Panin R. leg. L=10.5 mm
Copyright © Ruslan Panin

Harpalus (Semiophonus) signaticornis (Duftschmid, 1812)
Male adult
France, 77860, Quincy-Voisins, 12-05-2018, leg. & det. P. Deschamps
Copyright © Pascal Deschamps

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) fuscicornis Menetries, 1832: 134
Female adult
Germany, Iphofen, 2018, leg. & det. Boetzl
Copyright © Fabian Bötzl

Harpalus (Harpalus) amariformis Motschulsky, 1844
Female adult
Buryatia, Dzhida distr., Belozersk vic., 4-9.7.97, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) pygmaeus Dejean, 1829
Female adult
Greece bor. occ., 15 km W of Metsovo, Petra, 9.6.2008, leg. Orszulik
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) brevis Motschiulsky, 1844
Male adult
Russia, Irkutsk reg., Zalari distr., Tungui vill. 20-24.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) amputatus obtusus (Gebler, 1833)
Male adult
Russia, Buryatia, Dzhida distr., Belozersk vic., 4-9.7.1997, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) rufiscapus Gebler, 1833
Male adult
Russia, Irkutsk reg., Alarskii distr., vic. Alar vill. 26-28.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) macronotus Tschitscherine, 1893
Male adult
Russia, Irkutsk reg., Zalari distr., Tungui vill. 20-24.6.1997, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) aequicollis Motschulsky, 1844
Female adult
Russia, 14 km from Irkutsk by Aleksandrovskii road, 9.7.1994, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) lederi Tschitscherne, 1899
Female adult
Buryatia, Hamar-Daban Mts., Taltsy riv., 1700m 19-25.5.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) modestus Dejean, 1829
Female adult
Russia, 13 km from Irkutsk by Aleksandrovskii road, 20.8.1992, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) nigrans A. Morawitz, 1862
Female adult
Russia, Irkutsk vic., Mamony vill., Irkut riv., 11.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) sinuatus Tschitscherine, 1893
Male adult
Russia, 14 from Irkutsk by Aleksandrovskii road, 9.7.1994, Anichtchenko A. leg. & det. 6.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) udege Lafer, 1989: 189
Male adult
Russia, Baikal lake, Listvianka vill., 2.8.1992, Anichtchenko A. leg. & det. 6.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) quasianxius Kataev, 1989 Paratype
Male adult
Image courtesy of Biological Museum, Lund University
Copyright © - - -

Harpalus (Cryptophonus) melancholicus Dejean, 1829
Male adult
Italy, Patro di Moncalvo (Asti).
Copyright © Gianni Allegro

Ophonus (Hesperophonus) subquadratus (Dejean, 1829)
Male adult
Greece, Corfu, Benitses env.,23-29.6.2003. det. D Wrase (det confirmed by Anichtchenko 2021); 7 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) melletii (Heer, 1837)
Male adult
Slovakia, Sahy-Tesmak. 7.5 mm
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) planicollis (Dejean, 1829)
Female adult
Bulgaria, Pirin Sandanski, 30.7.2012 (DUBC) 6 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) parallelus (Dejean, 1829)
Male adult
Spain, Andalucia, Sierra de las Nieves, Los Quejigales, 6-8.4.2010 Anichtchenko A. leg & det. (DUBC) 7 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) rupicola (Sturm, 1818)
Male adult
Russia, NW Caucasus, Krasnodar reg., Taman env., 23.4.2000 (DUBC)
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) laticollis Mannerheim, 1825
Male adult
Russia, NW Caucasus, Novorossiisk distr., Markoth Mts., 23.5.2005 (DUBC) det. Anichtchenko A. 2021; 10.5mm
Copyright © Alexander Anichtchenko

Ophonus (Hesperophonus) subquadratus (Dejean, 1829)
Male adult
Bulgaria, Chernomorets, 24.11.2006, Anichtchenko A. leg. & det. (DUBC)
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) erosus Mannerheim, 1825
Male adult
E Siberia, lake Baikal, Listvyanka vill. 2.8.1992, Anichtchenko A. leg & det. (cAA) 11.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) vittatus Gebler, 1833
Male adult
Tuva, Tes-Hem13.6.1948 (cAA) 6.5 mm; det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) zabroides Dejean, 1829
Male adult
E Siberia, Irkutsk, downtown, 20.6.2001, Anichtchenko A. leg & det. (cAA) 15 mm
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) laeviceps (Menetries, 1832)
Female adult
Bulgaria, near Chernomorets, XI.2006 Anichtchenko A. leg. & det. (cAA) 7.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) heyrovskyi Jedlicka, 1928
Male adult
Russia, Altaiskiy reg., Zmeinogorsk, 11.6.1984 (cAA) 10.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) acupalpoides Reitter, 1900
Male adult
Mongolia, 2000m, 40km SW Arvayheer, 15-16.7.2006 Anichtchenko A. leg. & det. (cAA) 4.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) longipalmatus Mordkovitsh, 1969
Male adult
Russia, Chita reg., Daurian NP., Zun-Torey lake, 1.6.2002 Anichtchenko A. leg. (cAA) 4.3 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) albanicus Reitter, 1900
Male adult
Spain, Granada, Castell del Ferro, 5.4.2002 Anichtchenko A. leg. (cAA) 8.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) optabilis Dejean, 1829
Male adult
Burjatia, Selenga riv., Deben vic., 15.6.2003 Anichtchenko A. leg & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) tichonis Jakobson, 1907
Male adult
Siberia, Chita reg., 30 km SE Hilok vill., 28.5.2002 Anichtchenko A. leg & det. (cAA) 11.5mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) compressus Motschulsky, 1844: 213
Male adult
SW Ukraine, Odessa reg., Ivanovka distr., Gudevichevo vill., steppe, 13.VI-11.VII.2022, A.Gontarenko leg. L=12 mm
Copyright © Ruslan Panin

Harpalus (Harpalus) flavicornis Dejean, 1829
Male adult
Italy, Piedmont, Ottiglio (Cave di Moleto), 24.V.2009
Copyright © Gianni Allegro

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
Italy, Piedmont, Sambuco (CN), m 1400, 4.VI.2006
Copyright © Gianni Allegro

Harpalus (Harpalophonus) stevenii Dejean, 1829a: 242
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Heteroharpalus) metallinus Menetries, 1836
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Baryharpalus) caspius (Steven, 1806)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Brachyharpalus) kazanensis Jedlicka, 1958
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Hypsinephus) salinus Dejean, 1829
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Nephoharpalus) pallidipennis A. Morawitz, 1862
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Diaharpalus) vittatus Gebler, 1833
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalobius) viridanus Motschulsky, 1844
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Homaloharpalus) tangutorum Kataev, 1993
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Asioharpalus) nigrans A. Morawitz, 1862
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Anamblystus) torridoides Reitter, 1900
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Haploharpalus) macronotus Tschitscherine, 1893a: 372
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Caucasoharpalus) aeneipennis (Faldermann, 1836)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Drymoharpalus) atratus Latreille, 1804
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Anamblystus) luteicornis (Dufischmid, 1812)
Female adult
Latvia, Ilukste, 6.2010 M. Janovska leg (DUBC). det. Anichtchenko A. 2023
Copyright © Alexander Anichtchenko

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest, on the sands, 17.V-25.VI.2024, R.Panin leg., L=13 mm
Copyright © Ruslan Panin

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Kirill Makarov

Harpalus (Amblystus) rufipalpis rufipalpis
Male adult
Latvia, Ilgas 10.6.92 (DUBC) 9.8 mm
Copyright © Alexander Anichtchenko

Harpalus (Actephilus) picipennis (Duftschmid, 1812)
Male adult
Latvia, Ilgas 18.6.96 (DUBC). 6.3 mm
Copyright © Alexander Anichtchenko

Harpalus (Actephilus) picipennis (Duftschmid, 1812)
Female adult
Latvia, Ilgas 18.6.96 (DUBC) 6.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Cryptophonus) litigiosus Dejean, 1829
Male adult
Spain, Andalusia, Sierra Arana, 25 km NE Granada, near cueva del Agua, 16.x.2003, 1700m, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Harpalus (Harpalophonus) hospes ciscaucasicus Lutshnik, 1921
Male+Female adult
Kalmanskiy Distr., Ob River near Kalistratikha Vill., 52°58.5ʹN, 83°37ʹE, 12–14.VIII.2013, R. and E. Dudko leg. Image from Kataev & Dudko, 2029
Copyright © - - -