Carabidae image gallery
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Apotominae taxon page (3 photos)
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Brachininae taxon page (36 photos)
Brachinus (Brachynolomus) stenoderus Bates, 1873
Endophallus
Collection ground Hobetsu cho, Hokkaido Pref. 33485
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Pheropsophus (Stenaptinus) agnatus (Chaudoir, 1876)
Endophallus
1-3 — Vietnam, Cat Tien; 4-5 Vietnam, Kon Ka Kinh.
Copyright © Dmitri Fedorenko -
Broscinae taxon page (9 photos)
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Carabinae taxon page (982 photos)
Endofallus Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Endophallus
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822
Endophallus
Lviv Region, Brody distr., near Yasenov vill.,
Copyright © Ruslan Panin
Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Endophallus
endophallus
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Endophallus
endophallus sibiricus obliteratus Fischer, 1827
NE Kazakhstan,
S Altai mts.,S Ulbinsky mt.h=500m.,Turgusun vill., 20.VI.2001
male 25mm.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Endophallus
endophallus
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Endophallus
endophallus sibiricus errans
SW-Russia,
Rostov reg. env. stanitsa Razdorskaya, A.Ponomarev leg
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Endophallus
endophallus sibiricus errans- Lugansk reg., near Slavyanoserbk
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795
Endophallus
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Endophallus
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus </b> S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Endophallus
Moldova,<br /> Anenii Noi distr., near Garbovets vill.
Copyright © Ruslan Panin
Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Endophallus
endophallus
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Endophallus
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
Endophallus
endophallus
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
Endophallus
<p> endophallus odoratus odoratus (as melleus Lapouge, 1909)<br /> <br /> <br /> </p>
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Endophallus
endophallus
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Endophallus
endophallus
Copyright © Ruslan Panin
Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Endophallus
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Endophallus
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Endophallus
endophallus
Copyright © Ruslan Panin
Carabus (Microplectes) convallium convallium Starck, 1889
Endophallus
Endophallus
Copyright © Ruslan Panin
Carabus (Microplectes) convallium argonautarum Semenov, 1898
Endophallus
Endophallus
Copyright © Ruslan Panin
Carabus (Pachystus) glabratus glabratus Paykull, 1790
Endophallus
Endophallus
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as nubicola Zolotarev, 1913)
Endophallus
Endophallus prometheus prometheus (as nubicola Zolotarev, 1913)
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Endophallus
Endophallus prometheus prometheus Reitter, 1887
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Endophallus
aedeagus
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Endophallus
aedeagus miroshnikovi Psebai env., Nikitino
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Endophallus
Endophallus
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Endophallus
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill.,
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus (as gvozdevae Obydov, 1997)
Endophallus
W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill.,
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Endophallus
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Endophallus
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Endophallus
Copyright © Ruslan Panin
Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Endophallus
Steppe Reserve “Provalsky",
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians (as dvorshaki Mandl, 1975)
Endophallus
Copyright ©
Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Endophallus
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk,
Copyright © Ruslan Panin
Carabus (Carabus) arvensis arvensis Herbst, 1784
Endophallus
W Ukraine, NW Podolian, Lviv reg., Javoriv distr.,near Stradch vill.,
Copyright © Ruslan Panin
Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Endophallus
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan,
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Endophallus
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996
Copyright © Ruslan Panin
Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Endophallus
N Japan:Hokkaido, Hamamasu env.,
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Endophallus
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
Copyright © Ruslan Panin
Carabus (Pachystus) hortensis hortensis Linne, 1758
Endophallus
W-Ukraine, Male Polissya, near Kamenka-Buzka vill.
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Endophallus
Luxembourg, Bollendorf-Pont env., 3.III.1999
Copyright © Ruslan Panin
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Endophallus
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m,
Copyright © Ruslan Panin
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Endophallus
NС-Caucasus, Karachaevo-Cherkessia, right bank Pkhiya river, Abishir-Akhuba Mt.Range h=2250-2600 m,
Copyright © Ruslan Panin
Carabus (Pachycarabus) staehlini Adams, 1817
Endophallus
S Russia, E Caucasus, Dagestan, Nukatl Mt Rng h=2600m,
Copyright © Ruslan Panin
Carabus (Carabus) stscheglowi Mannerheim, 1827
Endophallus
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka,
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Sphodristocarabus) adamsi Adams, 1817
Endophallus
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env.,
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype
Endophallus
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Endophallus
Russia, Chita Region, Udokansky district. Kodar Range, 56 °34'42,9 " N, 118 °29'34,95" E, 6/13/2002, leg. A. Pokhorov
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Endophallus
S Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m
Copyright © Ruslan Panin
Carabus (Tribax) nacharensis Rost, 1893
Endophallus
Abkhazia, vall. Kluch River, 13.08.1928
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Endophallus
Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Endophallus
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m
Copyright © Ruslan Panin
Carabus (Cechenochilus) heydenianus Starck, 1889
Endophallus
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Megodontus) obovatus Fischer, 1827
Endophallus
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg. (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Endophallus
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Endophallus
Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) N-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Endophallus
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Endophallus
Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill.
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Endophallus
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain,
Copyright © Ruslan Panin
Carabus (Hemicarabus) nitens Linne, 1758
Endophallus
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Endophallus
Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
Endophallus
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
Endophallus
Russia: Kabardino-Balkaria: u.r. Adylsu Riv.
Copyright © Igor Solodovnikov
Carabus (Pachystus) cribratus cribratus Quensel, 1806
Endophallus
NW Armenia, Amasia, (Shirak prov.)
Copyright © Ruslan Panin
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Endophallus
E Siberia, Sokhondinskii NP, Ingoda riv., 49.50.207N 111.10.825E, 1700m, 23.7.2009 Shavrin A. leg.
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) kurilensis sichotensis Born, 1914
Endophallus
Bureinski NP, up Bureya river, Karbohon lake, tundra 28.7.2006
Copyright © Alexander Anichtchenko
Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Endophallus
NW Caucasus, Republic Adygea, Mt.Tybga h=2100 m,
Copyright © Ruslan Panin
Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Endophallus
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha”
Copyright © Ruslan Panin
Carabus (Tomocarabus) marginalis Fabricius, 1794
Endophallus
N Ukraine, Kiev reg., Vyshgorod disrt., near Khotyanivka vill.,
Copyright © Ruslan Panin -
Cicindelinae taxon page (124 photos)
Cicindela (Cicindela) monticola monticola
Endophallus
Karachaevo-Cherkessia, Hudes riv., near Tsarskie vorota
Copyright © Alexander Anichtchenko -
Dryptinae taxon page (11 photos)
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Elaphrinae taxon page (55 photos)
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Harpalinae taxon page (336 photos)
Trichotichnus (Trichotichnus) lucidus (A. Morawitz, 1863)
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Stenolophus (Stenolophus) propinquus A.Morawitz, 1862
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Pseudoophonus) roninus Bates, 1873
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Endophallus
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Zangoharpalus) tinctulus Bates, 1873
Endophallus
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Endophallus
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Harpalus) crates Bates, 1883: 239
Endophallus
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Endophallus
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Loboharpalus) platynotus Bates, 1873
Endophallus
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Endophallus
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Bradycellus (Tachycellus) subditus (Lewis, 1879)
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 39249
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Anisodactylus (Pseudanisodactylus) punctatipennis A.Morawitz, 1862
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Endophallus
Collection ground Mukawa cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Acupalpus (Palcuapus) inornatus Bates, 1873
Endophallus
Collection ground Kamogawa city, ChibaPref. 34922
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Acupalpus (Setacupalpus) hilaris Tschitscherine, 1899
Endophallus
Russia, Evreiskaya AO, Birobidzhan vic., 3.11.2013. leg. Budilov. P.
Copyright © Pavel Budilov
Bradycellus (Tachycellus) glabratus Reitter, 1894d: 125
Endophallus
Russia, Evreiskaya AO, Birobidzhan vic., 3.11.2013. leg. Budilov. P.
Copyright © Pavel Budilov
Acupalpus (Acupalpus) inouyei (Habu, 1980)
Endophallus
Kunashir Island, west coast of Lake Peshchanoe 29.IX.2014 leg. Yu. Sundukov
Copyright © Kirill Makarov
Carterus (Pristocarterus) angustus (Menetries, 1832)
Endophallus
Iran, Lorestan prov., near Dorud, 5 km SW Chabdar vill. 33.501061°N 49.003441°E 14-15.V.2017 leg. D.Kasatkin
Copyright © Kirill Makarov
Anisodactylus (Anisodactylus) arsenjevi Kataev, Makarov & Sundukov, 2024 Paratype
Endophallus
S Primorye Territory, Yakovlevka Distr., 1 km S Beltsovo Vill., Arsenjevka R. floodplain, h~120 m, 44°48'57"N 133°34'11"E, 21.V.2024, K. Makarov leg.
Copyright © Kirill Makarov
Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Endophallus
Primorye Territory, Khasan Distr., vicinity of Slavyanka Vill., Cape Busse, 42°52'42"N 131°27'58"E, 13.VI.2023, I. Melnik leg. L=11.0mm
Copyright © Kirill Makarov
Anisodactylus (Pseudanisodactylus) punctatipennis A.Morawitz, 1862
Endophallus
Primorye Territory, Khasan Distr., 12.5 km WNW of Barabash Vill., Ovchinnikovo, 43°13'56"N 131°21'23"E, h~110 m, 10–13.VIII.2020, K. Makarov, A. Matalin leg. L=11.1 mm
Copyright © Kirill Makarov
Harpalus (Cryptophonus) litigiosus Dejean, 1829
Endophallus
Spain, Andalusia, Sierra Arana, 25 km NE Granada, near cueva del Agua, 16.x.2003, 1700m, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko -
Lebiinae taxon page (222 photos)
Cymindis (Menas) daimio Bates, 1873
Endophallus
Collection ground Tomakomai city, Hokkaido Pref. 37172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Demetrias (Demetrias) marginicollis Bates, 1883: 285
Endophallus
Collection ground Kawakami gun, OkayamaPref. 33401
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Demetrias (Demetrias) amurensis Motschulsky, 1860: 39
Endophallus
Collection ground Atsuma cho, Hokkaido Pref. 36996
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Lebia (Poecilothais) bifenestrata A. Morawitz, 1862a: 245
Endophallus
Collection ground Misugi machi, Mie pref. 34126
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Lebia (Poecilothais) retrofasciata Motschulsky, 1864: 227
Endophallus
Collection ground Toyoura cho, Hokkaido Pref. 34482
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Perigona (Perigona) exigua (A.Morawitz, 1863)
Endophallus
Kunashir Island, Cape Ivanovskii, 20.IX.2013, Yu. & L. Sundukov leg.
Copyright © Kirill Makarov
Dromius (Dromius) agilis (Fabricius, 1787)
Endophallus
Karachay-Cherkessia, gorge. Dukka, h~2100 3-5.VII.2020 leg. M. Nabozhenko
Copyright © Kirill Makarov -
Licininae taxon page (149 photos)
Badister (Baudia) nakayamai Morita, 1992c: 156
Endophallus
Collection ground Ebetsu city , Hokkaido Pref. 35604
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Licinus (Licinus) yezoensis Habu, 1947b: 91
Endophallus
Collection ground Pippu cho, Hokkaido Pref. 36636
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Badister (Badister) pictus Bates, 1873
Endophallus
Collection ground Noda city, iChiba pref. 34517
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Chlaenius (Agostenus) alutaceus Gebler, 1829: 48
Endophallus
South of Primorsky Territory, near Lake Khanka, Turiy Rog, 15.VII.1998, V. Patrikeev leg.
Copyright © Kirill Makarov
Chlaenius (Agostenus) quadrisulcatus Paykull, 1790: 109
Endophallus
South Primorye, Khasansky district, shore of Melkovodnaya Bay 3.VI.2019 leg. K.Makarov
Copyright © Kirill Makarov
Chlaenius (Agostenus) quadrisulcatus Paykull, 1790: 109
Endophallus
South Primorye, Khansan district, middle flow of Narva River, left bank, swamp 43°04'05"N 131°20'26"E 24.V.2021 leg. Yu.Sundukov L=13.0 mm
Copyright © Kirill Makarov
Chlaenius (Eochlaenius) suvorovi Semenov, 1912
Endophallus
South Primorye, Khasansky district, 6.5 km ESE of Khasan, bank of the channel 42°25'07"N 130°43'36"E 14.V.2023 leg. K. Makarov
Copyright © Kirill Makarov -
Loricerinae taxon page (3 photos)
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Melaeninae taxon page (2 photos)
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Nebriinae taxon page (107 photos)
Nebria (Catonebria) sajana sarlyk Dudko et Shilenkov, 2001
Endophallus
Altai, Seminskiy Mt.R., Mt. Sarlyk, h~1950-2250 13.VI.2013 leg. R. et E. Dudko
Copyright © Kirill Makarov
Nebria (Nebria) bonellii M.F. Adams, 1817
Endophallus
NW Caucas, Adygea reg., Lago Naki plato, Murzikal mt., 2100 m, 28.6.07
Copyright © Alexander Anichtchenko -
Omophroninae taxon page (3 photos)
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Panagaeinae taxon page (15 photos)
Panagaeus (Panagaeus) japonicus Chaudoir, 1861c: 356
Endophallus
Collection ground Kaminokuni cho, Hokkaido Pref. 35988
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Panagaeus (Panagaeus) robustus A. Morawitz, 1862b: 323
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 36670
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Tinoderus singularis Bates, 1873a: 245
Endophallus
Collection ground Fujioka machi, TochigiPref. 34784
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Peronomerus auripilis Bates, 1883: 235
Endophallus
Collection ground Taiki cho, Hokkaido Pref. 34157
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro -
Paussinae taxon page (9 photos)
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Platyninae taxon page (205 photos)
Metacolpodes buchanani Hope, 1831: 21
Endophallus
Russia, Evreiskaya AO., Birobidjan 31.07.2013
Copyright © Pavel Budilov
Calathus (Neocalathus) micropterus (Dufischmid, 1812)
Endophallus
Sakhalin isl.
Copyright © Alexander Anichtchenko
Calathus (Neocalathus) melanocephalus melanocephalus
Endophallus
Left - Spain, Montes Cantabricos; right – paratype of C. (N.) anistschenkoi syn.
Copyright © Alexander Anichtchenko
Anchomenus (Anchomenus) leucopus Bates, 1873
Endophallus
South Primorye, Khasansky district, middle flow Narva River, right coast 43°03'52"N 131°20'45"E 24.V.2021 leg. A. Zaitsev L=8.0 mm
Copyright © Kirill Makarov -
Psydrinae taxon page (1 photos)
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Pterostichinae taxon page (521 photos)
Amara (Zezea) plebeja (Gyllenhal, 1810)
Endophallus
Collection ground Chitose city, Hokkaido Pref. pref. 34882
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Bradytus) sinuaticollis A. Morawitz, 1862
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Bradytus) simplicidens A. Morawitz, 1863: 60
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Bradytus) majuscula (Chaudoir, 1850)
Endophallus
Collection ground Monbetsu cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Curtonotus) macronota Solsky, 1875
Endophallus
Collection ground Tomakomai city, Hokkaido Pref. 37172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Curtonotus) hiogoensis (Bates, 1873)
Endophallus
Collection ground Tsushima, NagasakiPref. 35695
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Curtonotus) gigantea (Motschulsky, 1844)
Endophallus
Collection ground Kagoshima city, Kagoshima Pref. 38979
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Reductocelia) lucens Baliani, 1943: 44
Endophallus
Collection ground Sadamitsu cho, Tokushima Pref. 38262
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Reductocelia) chalcophaea Bates, 1873a: 292
Endophallus
Collection ground Tomakomai city, Hokkaido Pref. Tomakomai～shi 37172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Amara) ussuriensis Lutshnik, 1935a: 260
Endophallus
Collection ground Otobe cho, Hokkaido Pref. 35273
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Amara) obscuripes Bates, 1873a: 294
Endophallus
Collection ground Shibecha cho, Hokkaido Pref. 38493
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Amara) familiaris (Duftschmid, 1812)
Endophallus
Collection ground Kiyosato cho, Hokkaido Pref. pref. 38785
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Amara) congrua A. Morawitz, 1862b: 326
Endophallus
Collection ground Ise city, Mie Pref. 37125
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Amara (Amara) chalcites Dejean, 1828: 462
Endophallus
Collection ground Ebetsu city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Pterostichus (Eosteropus) japonicus (Motschulsky, 1860)
Endophallus
Collection ground Tomakomai, Hokkaido Pref. 30926
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Endophallus
Collection ground Rebun Is. Hokkaido Pref. 27191
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Pterostichus (Morphnosoma) chechcirensis Lafer, 1979: 28 Paratype
Endophallus
Copyright © Pavel Budilov
Pterostichus (Phonias) kutensis Poppius, 1905
Endophallus
Хабаровский край, Верхнебуреинский район, окр. пос. Софийск. Лиственничная тайга, h~1000м, 4.09.2013 (Habarovsk prov., Verhnebureinsk distr., Sofiisk vill.)
Copyright © Pavel Budilov
Amara (Amara) silvestrii Baliani, 1937: 179
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -
Pterostichus (Eosteropus) orientalis nigromontanus Lafer & Budilov, 2015 Holotype
Endophallus
Россия: Приморский край, Хасанский р-н, заповедник «Кедровая Падь», 27.VIII 2004, В.Н. Кузнецов
Copyright © Pavel Budilov
Amara (Curtonotus) gigantea (Motschulsky, 1844)
Endophallus
Russia, Primorskii reg.
Copyright © Alexander Anichtchenko
Poecilus (Poecilus) cupreus (Linne, 1758)
Endophallus
Turkije : Ardahan; Ardahan 7KM E Karst 13-7-1992 1900m. Heinz W. leg.
Copyright © Jan Muilwijk
Poecilus (Poecilus) cupreus erythropus Dejean, 1828
Endophallus
Libanon J.Sannin Sud 16-5-2000 1300 I. Rapuzzi leg.
Copyright © Jan Muilwijk
Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Endophallus
Agri, Balık Gölü, Taşlıçay, 2250 m, 17.07.76; W. Heinz leg.
Copyright © Jan Muilwijk
Pterostichus (Badistrinus) neglectus A.Morawitz, 1862
Endophallus
Khabarovsk Territory, Khawar Stream below cordon Bichi, 24.VII.2021, Yu. Sundukov & L. Sundukova leg.
Copyright © Kirill Makarov
Pterostichus (Badistrinus) goschi Jedlicka, 1930
Endophallus
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov
Pterostichus (Badistrinus) goschi Jedlicka, 1930
Endophallus
Primorye Territory, Khasansky district, 10.7 km east of Khasan, Cape Ostrovok Falshivy, 42°26’54”N 130°46’47”E 23.V.2022, K. Makarov leg.
Copyright © Kirill Makarov
Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
South Primorye, Kedrovaya Pad nature reserve, mouth of Kedrovaya River, swamp 43°04'47"N 131°36'42"E 1.VI.2019 leg. K. Makarov L= 7.1
Copyright © Kirill Makarov
Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
Hirotuka Kanagawa-Ken 23.jul.1958 S.Nomura L= 7.4 mm
Copyright © Kirill Makarov
Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.2 mm
Copyright © Kirill Makarov
Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov -
Rhysodinae incertae sedis taxon page (7 photos)
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Scaritinae taxon page (41 photos)
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Siagoninae taxon page (3 photos)
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Trechinae taxon page (379 photos)
Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014
Endophallus
Male genital apparatus of Bembidion (?Nipponobembidion) ruruy sp. n., holotype. Scale bars 0.25 mm. A aedeagus, right side B aedeagus, left side C right paramere D left paramere E sternite 9, ventral view F sternite 8 G tergite 8.
Copyright © Kirill Makarov
Bembidion (Plataphodes) tetraporum Bates, 1883: 270
Endophallus
Aedeagus of Bembidion (Plataphodes) tetraporum Bates, 1883. Scale bars 0.25 mm. A right side B left side.
Copyright © Kirill Makarov
Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Endophallus
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov
Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Endophallus
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Endophallus
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov
Copyright © Kirill Makarov
