Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
NW Podolian
Copyright © Safronov

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
Moldova, 2003
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
Male adult
W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., W Ukraine, Lysa Gora steppe massif, 19.V.-7.VI.2007, Panin R. leg. 25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Male adult
Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Endophallus
Moldova,<br /> Anenii Noi distr., near Garbovets vill.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian, Ternopil reg., Pidvilochisk distr., near Rosokhuvatets & Kamyanki vill., steppe sites, 24.V-15.VI.2008, Panin R. leg.
male, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (f.i. frivaldszkyi Kraatz, 1887)
Female adult
W-UKRAINE, NW Podolian Upland,<br /> Lviv Reg., Zolochiv Distr., Lysa Gora steppe massif, 14-30.V.2008, Panin. R. leg./det.<br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
NC Ukraine, Kiev, Lysa Gora, on the fringes forest, 5-24.VI.2007, Gerasimov R.P.. leg.<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
C- UKRAINE, NW Odessa reg., Kodyma distr., near Aleksandrivka vill.,18.V.2003, Demidov G.N. leg.<br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Female adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Steppe Reserve Lysa Gora, 19.V-7.VI.2007, Panin R. leg.<br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens ( as jasilkowskii Born,1905)
Male adult
NE Romania, 5 km W Suceava, 47°63′940″N 26°18′795″E,h=340m, agricultural field, 15-20.V.2011, B. Brezina leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as moldaviensis Born, 1903)
Female adult
E-Romania, Vaslui reg.,15 km SE Vaslui, Crasna env., V-VI,2013, M,Pasek leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens paradoxus Lapouge, 1906 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC539
Copyright © - - -

Carabus (Morphocarabus) excellens paradoxus
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC539
Copyright © - - -

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 4-24.V.2010, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicusn (as rareulensis Born, 1907)
Male adult
NE-Romania, Rarau Mts. h-1300m, VI.2014, J. Kynkor leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
Male adult
SW Poland: Przemyśl env. photo by Mieczyslaw Stachowiak
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
Female adult
W Ukraine, Lviv Region, Zhydahciv distr., Borodchytsi vill.,the right bank of Dniester river, 11-16.VII.2012, Panin. R leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype
Endophallus
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 18.V-12.VI.2014, L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
Habitat
Biletsky flood-plain forest,
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 31 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 30 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 12.V-2.VI.2009, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 31 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (ab.goldeggi Duftschmid, 1812)
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Female adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
W Ukraine, Vinogradov distr.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
endophallus hampei
Vinogradov distr.
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male+Female adult
W.Ukraine.
Transcarpathian region,Vinogradiv distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.2008. Panin R. leg.
female 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
W.Ukraine.
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male, 28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as ormayi Reitter, 1896) loc.typ.
Female adult
W.UKRAINE,
Transcarpathian reg., near Beregovo,15.V-16.VI.2009 Kanarsky Yu. & Geryak Yu. leg.
female, 30 mm
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
Carabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
arabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
Male+Female adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
female, 32 mm
photo endophallus Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus Dejean, 1831
Male adult
Carabus (Morphocarabus) rothi comptus Dejean, 1831
Carpathians
male, 23mm
coll.SMNH Ukraine ,Lvi
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male adult
C Romania,
Olt River: Rimnica Vilcea VII.1971, Savulescu N. leg.

Copyright © Igor Solodovnikov

Carabus (Morphocarabus) rothi rothi Dejean, 1829
Male+Female adult
Romania
Banat, Nemeseti h=250m. VI.1989 Lie P. leg.

photo Solodovnikov I.A.
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
Male adult
NW Romania , Bistrita Nasaud, Mts. Tibles, 6 km SE Bichigiu, 17.VI.2009, S.Budka leg.
L=35 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as dacicus Csiki, 1906)
Female adult
NW Romania , Bistrita Nasaud, Beclean env., 17.VI.2009, S.Budka leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Male adult
W. UKRAINE,
Transcarpathian reg., near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Endophallus
near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
C Romania, Târgu Mureş, Ernei, 46°35′471″N 24°37’351″E, h=300 m,20.VIII.2004, Sl.Budka leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
C Romania, Tвrgu Mureş,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
Male+Female adult
NW Romania, Zalău env., 31.VII.2003, Sl.Budka leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Female adult
SW Romania, Tarcu Mts, vf. Matania Mt. h=2000 m, 12.VI.2008, S.Budka leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Muntii Metaliferi, Hunedoara reg., near Sacaramb vill., 6.VI.2011, S.Budka leg.
L=37 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei as telekii Csiki, 1937 (? hampei )
Female adult
CW Romania, Arad distr., Zarandulu Mt Range, near Ilteu vill., h=276 m, 46°66′487″N 22°20’455″E, 25.VI.2005, Sl.Budka leg.
L=30 mm

Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi Dejean, 1829
Male+Female adult
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m, 19.VIII.2001, S.Budka leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi Dejean, 1829
Endophallus
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Endophallus
SW Romania, Tarcu Mts, vf. Nedeia h=2000m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Male adult
SW Romania, Tarcu Mts, vf. Nedeia h=2000m, 12.VI.2008, J.Kynkor leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Male adult
N Romania, E Carpathian, Maramureş Reg., Gutâi Mts, cabana Mogosa,4 km N Baia Sprie, 9-17.VI.2010, J.Kynkor leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Endophallus
N Romania, E Carpathian, Maramureş Reg., Gutai Mts, cabana Mogosa, 4 km N Baia Sprie,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Endophallus
C Romania, Brașov County, Cristian vill., h=380 m
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Male+Female adult
C Romania, Brașov County, Cristian vill., h=380 m,V-VI.2008, M.Pasek leg.
female, 27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi (? incompsus as spectabilis Csiki, 1906)
Female adult
NC Romania, Bistrita Nasaud reg., Călimani Mts, Colibita vill., mixed forest, meadows h=800m, 15-21.V.2011, M.Pasek leg.<br /> L=34 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus (as mendax Csiki, 1906)
Female adult
W Ukraine, NE Carpathians,
Transcarpathian reg., Chornogora Mt Range, h=1100m,13.VI.2011, Panin R. leg./det.
L=33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Endophallus
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male+Female adult
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts., VII.2007, I. Marvan leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Male adult
SW Romania, S Carpathians,
Banat , Tarcu Mts, Borlova env., 19-31.V.2005, P. Sima leg.
L=24 мм
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Male adult
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,. 6.VIII.2008
L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Endophallus
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill., 45°97'472"N/23°02'630"E, h=680 m, mixed forest, meadows, 15-20.V.2011, B.Brezina leg.
L=36 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Endophallus
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
Female adult
C Romania, Sibiu reg., left bank Olt River, Avrig env., 12.VI.2013, V. Slovak leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Endophallus
SW Romania,, Banat, Borlova env.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii Kraatz, 1854
Male adult
Ukrainian Carpathians
Lviv Reg., Skole Distr., near Grebeniv vill., h=1000m 9 - 20.VII.07, Panin R. leg.
male, 31 mm
Copyright © Safronov

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
W-UKRAINE, Transcarpathian region, Rakhiv distr., Chornogora Mt Range, h=800 m, near Bogdan vill., 10.VI.2012, Panin R. leg.
L= 33 мм
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
W Ukraine,Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., near Lug (Lunca la Tisa) vill., h=400 m,12-15.VII.2011
L=32 mm
det.Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Endophallus
W Ukraine,Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., near Lug vill., h=400 m.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Female adult
W Ukraine, NE Carpathians,
Transcarpathian Reg., Marmarosh Mts., Rakhiv distr., 5 km W Dilowe vill., Site Kuziy, 5-15.VII.2010, Demidov G.N. leg./det.Panin R.
L= 35 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov distr., Svidovec Mt Rng., Mt. Podpula h=1500-1600m. Lazutkin G. leg, 13-14.VII.1978, det. Panin R.
L=28mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Female adult
W Ukraine, Transcarpathian, Marmarosh Mts., near Lug vill., h=400 m, 28.IV -26.VII.2017, Panin R. leg., L= 38 mm
Copyright © Ruslan Panin