Anisodactylus (Hexatrichus) poeciloides pseudoaeneus Dejean, 1829
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Anisodactylus (Anisodactylus) binotatus (Fabricius, 1787)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus rufipes (Degeer, 1774)
Male adult
Russia, S.Ural
Copyright © Sunvas

Carterus (Pristocarterus) angustus (Menetries, 1832)
Male adult
Azerbaijan, Bozdagh range,Turianchai Nature Reserve, 07.VII.2008 Pljushch I.G. leg. L=8mm
Copyright © Ruslan Panin

Ditomus calydonius oriens (Dvorak, 1993)
Male adult
S Kazakhstan, Karatau mts.,<br /> Buylikkul lake 26.6.1999<br /> 16 mm
Copyright © Alexander Anichtchenko

Ditomus calydonius oriens (Dvorak, 1993)
Male adult
length: 15 mm
Turcia, Troja
leg. U.Schmidt, 1997; det. A.Skale
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Dixus eremita Dejean, 1825
Male adult
Volgogrd Area, Pallasovka Distr., 5 km NNW Elton Vill., Sorochya Balka 8.IV.2007 leg. A.Matalin
L=9.0
Copyright © Kirill Makarov

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
NW Podolian Upland, W UKRAINE,<br /> Lviv Reg., Zolochiv distr., near Gologirki vill., 30.04-19.05.07 Panin R.leg.<br /> L= 15mm
Copyright © Ruslan Panin

Harpalus (Harpalus) affinis (Schrank, 1781)
Male adult
length: 9-12 mm
Helvetia, Graubünden, Ilanz
leg.det. U.Schmidt, 2004
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Harpalus (Harpalus) caspius (Steven, 1806)
Male adult
W UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa vill., Panin R. leg. 1.V-16.VI.2007
male, 13mm
Copyright © Ruslan Panin

Harpalus (Harpalus) fuscipalpis Sturm, 1818
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Pseudoophonus) rufipes (Degeer, 1774)
Male adult
W-UKRAINE,
Lviv Region, Javorov distr., near Ivano-Frankovo 1-20.VI.07 Panin R. leg
long = 17mm
Copyright © Ruslan Panin

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Male adult
length: 6,0-9,1 mm Germania, Bayern, Pegnitz<br /> leg.det. U.Schmidt, 1972<br /> Photo: U.Schmidt, 2008
Copyright © U.Schmidt

Dixus obscurus Dejean 1825
Male adult
S Ukraine, Crimea
Cape Tarhankut, Site Djangul, 2-3.VII.2010, Shaporinskiy V. leg./det.Panin R.
L=11.5 mm

Copyright © Ruslan Panin

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) calceatus (as itoshimanus) Holotype
Male adult
Mt. Raizan, FUKUOKA, 22. VIII. 1946, A. Habu. COL-132
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Male adult
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Male adult
NW Caucas, Krasnodar reg., SE Goryachii Kluch, леса между пос. Асфальтовая Гора и Садовый (нежилой) 21.V.2012 leg. М. Смирнов
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Daptus pictus Fischer von Waldheim, 1823
Male adult
Kazakhstan, 20 km NW Akkuduk vill., N 43*06'58 E 054*11'46, h=90m, 24.V.2011, Ivanov A. leg.
det. Anichtchenko A. 2013
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) tardus (Panzer, 1796)
Male+Female adult
France, forêt de Fontainebleau (77300), 18/4/1987, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Male adult
Portugal, Algarve Portimao, 25/9/1983, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Pangus scaritides (Sturm, 1818)
Male adult
SK, S Slovakia, Velké Kosihy; 8mm
Copyright © Aleš Sedláček

Harpalus (Cephalophonus) cephalotes Fairmaire & Laboulbene, 1854
Male adult
SE Bulgaria, Primorsko; 16mm
Copyright © Aleš Sedláček

Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)
Male adult
S Slovakia, Jurský Chlm; 10mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Female adult
SK, S Slovakia, Veľký Krtíš; 12mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) ardosiacus (Lutshnik, 1922)
Female adult
CZ, S Moravia, Mikulov; 10,5mm
Copyright © Aleš Sedláček

Ophonus (Ophonus) stictus Stephens, 1828
Male adult
CZ, SE Moravia, Uherský Brod; 12mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) pulvinatus lubricus Reitter, 1900
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) metallinus Menetries, 1836: 149
Female adult
det. B. Kataev, 2013
Copyright © Kirill Makarov

Acinopus (Oedematicus) megacephalus (P. Rossi, 1794)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
Moravia, Kobylí; 11 mm
Copyright © Aleš Sedláček

Microderes (Microderes) brachypus (Steven, 1809)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Chilotomus arnoldii Kryzhanovskij, 1962b : 541
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Parophonus (Parophonus) planicollis (Dejean, 1829)
Female adult

Copyright © Alexander Anichtchenko

Harpalus (Harpalus) circumpunctatus Chaudoir, 1846a: 171
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Hesperophonus) convexicollis Menetries, 1832
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) reflexus anadoluensis Kataev, 1993
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) flavicornis Dejean, 1829a: 366
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) politus Dejean, 1829a: 370
Male adult
Germany, Bavaria, Reiterswiesen, F. Boetzl leg. 21.05.14, M. Fritze det. 2014
Copyright © Fabian Bötzl

Ditomus calydonius oriens (Dvorak, 1993)
Male adult

Copyright © Kirill Makarov

Dixus obscurus Dejean 1825
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Male adult
det. A. Matalin
Copyright © Maksim Smirnov

Ophonus (Ophonus) stictus Stephens, 1828
Female adult
W Ukraine, NW Podolian Upland, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.V-15.VI.2017, Panin R. & Kanarsky Yu. leg. L=15 mm
Copyright © Ruslan Panin

Carterus (Pristocarterus) angustus (Menetries, 1832)
Female adult
Iran, Lorestan prov., near Dorud, 5 km SW Chabdar vill. 33.501061°N 49.003441°E 14-15.V.2017 leg. D.Kasatkin L=8.5 mm
Copyright © Kirill Makarov

Carterus (Pristocarterus) angustus (Menetries, 1832)
Male adult
Iran, Lorestan prov., near Dorud, 5 km SW Chabdar vill. 33.501061°N 49.003441°E 14-15.V.2017 leg. D.Kasatkin L=8.5 mm
Copyright © Kirill Makarov

Harpalus (Harpalus) smyrnensis medicus Kataev, 1993
Female adult
Iran, East Azerbaijan prov., near Kandovan vill. 37.748092°N 46.286895°E 21-22.V.2017 leg.D.Kasatkin L=10.0
Copyright © Kirill Makarov

Harpalus (Harpalus) smyrnensis medicus Kataev, 1993
Male adult
Iran, East Azerbaijan prov., near Kandovan vill. 37.748092°N 46.286895°E 21-22.V.2017 leg.D.Kasatkin L=9.5 mm
Copyright © Kirill Makarov

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) fuscicornis Menetries, 1832: 134
Female adult
Germany, Iphofen, 2018, leg. & det. Boetzl
Copyright © Fabian Bötzl

Harpalus (Harpalus) quasianxius Kataev, 1989 Paratype
Male adult
Image courtesy of Biological Museum, Lund University
Copyright © - - -

Harpalus (Cryptophonus) melancholicus Dejean, 1829
Male adult
Italy, Patro di Moncalvo (Asti).
Copyright © Gianni Allegro

Ophonus (Metophonus) melletii (Heer, 1837)
Male adult
Slovakia, Sahy-Tesmak. 7.5 mm
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) maculicornis (Duftschmid, 1812)
Male adult
Spain, Guipuzcoa, Irun, Playaundi, 28.5.2006, Anichtchenko A leg. & det. (cAA) 6.5 mm
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) planicollis (Dejean, 1829)
Female adult
Bulgaria, Pirin Sandanski, 30.7.2012 (DUBC) 6 mm
Copyright © Alexander Anichtchenko

Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)
Male adult
S Tadzhikistan, Kurgan-Tube reg., 20km S Dzhilikul, 11.5.1999 (DUBC) 8.5mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) rupicola (Sturm, 1818)
Male adult
Russia, NW Caucasus, Krasnodar reg., Taman env., 23.4.2000 (DUBC)
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) zabroides Dejean, 1829
Male adult
E Siberia, Irkutsk, downtown, 20.6.2001, Anichtchenko A. leg & det. (cAA) 15 mm
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) laeviceps (Menetries, 1832)
Female adult
Bulgaria, near Chernomorets, XI.2006 Anichtchenko A. leg. & det. (cAA) 7.5 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) dimidiatus (P. Rossi, 1790)
Male+Female adult
France 21410 Pont de Pany Sainte-Marie-sur-Ouche 30-06-1984 et 18130 Dun sur Auron 18-05-2012
Copyright © Pascal Deschamps

Harpalus (Harpalus) flavicornis Dejean, 1829
Male adult
Italy, Piedmont, Ottiglio (Cave di Moleto), 24.V.2009
Copyright © Gianni Allegro

Harpalus (Harpalus) tardus (Panzer, 1796)
Male adult
Italy, Piedmont, Sambuco (CN), m 1400, 4.VI.2006
Copyright © Gianni Allegro

Amblystomus niger (Heer, 1841)
Male adult
Italy, Toscana, Padule di Fucecchio, Pistoia. Image from Wrase & Magrinii, 2012
Copyright © - - -

Amblystomus rectangulus Reitter, 1883a: 143
Male adult
Italy, Asciano, Siena. Image from Wrase & Magrinii, 2012
Copyright © - - -

Harpalus (Heteroharpalus) metallinus Menetries, 1836
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Baryharpalus) caspius (Steven, 1806)
Male adult
Image from Kataev, 2023
Copyright © - - -

Oedesis caucasicus (Dejean, 1831)
Female adult
Italy, Veneto, Rosara di Codevigo, alla lampada UV, 10-12.V.2017; 8.1 mm. Image from Zanella & Uliana, 2022
Copyright © - - -

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Kirill Makarov

Harpalus (Amblystus) rufipalpis rufipalpis
Male adult
Latvia, Ilgas 10.6.92 (DUBC) 9.8 mm
Copyright © Alexander Anichtchenko

Harpalus (Cryptophonus) litigiosus Dejean, 1829
Male adult
Spain, Andalusia, Sierra Arana, 25 km NE Granada, near cueva del Agua, 16.x.2003, 1700m, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Acupalpus (Acupalpus) meridianus (Linnaeus, 1761)
Male adult
length: 3-4 mm
Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1993
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Anthracus consputus (Duftschmid, 1812)
Male adult
length: 3,8-5,1 mm<br /> Austria, Burgenland, Schützen a. Gebirge<br /> leg.det., G.Rößler, 1977<br /> Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Dicheirotrichus (Dicheirotrichus) ustulatus (Dejean, 1829)
Male adult
Volgogrda Area, Pallasovka Distr., Elton Vill., on light 6.X.2006 leg. K.Makarov
L=6.5
Copyright © Kirill Makarov

Stenolophus (Stenolophus) mixtus (Herbst, 1784)
Male adult
Poland: EB56
Piotrawin
13.VI.2008
leg. G. Bistu&a-Prószy&ski
Copyright © Mieczyslaw Stachowiak

Stenolophus (Stenolophus) mixtus (Herbst, 1784)
Female adult
length: 5-6 mm
Germania, Thüringen, Harra
leg.det. U.Schmidt, 2000
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Stenolophus (Stenolophus) teutonus (Schrank, 1781)
Male adult
Azerbaijan,
near Khachmaz, 16.VII.2008, Pljushch I.G. leg.
L=7mm
Copyright © Ruslan Panin

Dicheirotrichus (Trichocellus) placidus (Gyllenhal, 1827)
Male adult
length: 4,2-5 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2006
Photo: U.Schmidt, 2008
Copyright © U.Schmidt

Bradycellus (Bradycellus) caucasicus Chaudoir, 1846: 187 (Acupalpus)
Male adult
L=4,0 mm
RUSSIA, Tver reg., Likhoslavlskiy distr., Domanikha village, in the garden, in soil. H=160 m. N57*13.629’ E035*45.675’ 21.VII.2005
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Acupalpus (Acupalpus) parvulus (Sturm, 1825)
Male adult

Copyright © Kirill Makarov

Acupalpus (Acupalpus) exiguus Dejean, 1829
Male adult

Copyright © Kirill Makarov

Stenolophus (Stenolophus) steveni Krynicki, 1832
Female adult
SE Slovakia, Slovenský kras; 6,5mm
Copyright © Aleš Sedláček

Acupalpus (Acupalpus) meridianus (Linnaeus, 1761)
Male adult

Copyright © Kirill Makarov

Anthracus consputus (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Stenolophus (Egadroma) marginatus Dejean, 1829
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Acupalpus (Acupalpus) suturalis Dejean, 1829
Male adult
Turkey, Yahyah, Ağcaşar Baranjı.
Copyright © Bernd Jaeger

Dicheirotrichus (Dicheirotrichus) lacustris (L.Redtenbacher, 1858)
Female adult
Dagestan: 20 км E Mahachkala, Syrykum, 2.VI.2019, И.А. Забалуев leg. det. B.Kataev, 2020. Image from www.zin.ru by I. Zabaluev
Copyright © - - -

Acupalpus (Acupalpus) dubius Schilsky, 1888
Male adult
W-Ukraine, Roztochia, Lviv Reg., Yavoriv distr., N of Ivano-Frankovo vill., Site «Zalyvky» peat bog, 12.V - 8.VI.2021, R.Panin leg., L=3.3 mm
Copyright © Denis Khrapov

Acupalpus (Acupalpus) luteatus (Duftschmid, 1812)
Male adult
France, 85270 Saint-Hilaire-de-Riez, 02-09-2023
Copyright © Pascal Deschamps