Brachinus (Brachynolomus) scotomedes L. Redtenbacher, 1867
Female adult
Japan, Higashihagio Town, Omuta City, Fukuoka Pref., 19.6.2011, S. Nishimoto leg.
Copyright © Alexander Anichtchenko

Broscus punctatus Dejean, 1828: 431
Female adult
India, Uttarakhand, 17-18.04.2012, Uttarkashi distr., N30*45'15.72" E78*34'32.80, 1374 m, 14 km E Uttarkashi, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Broscodera (Sinobrosculus) chukuai Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1026168; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China, scale line = 1.0 mm. Image from Kavanaugh & Liang, 2021
Copyright © David Kavanaugh

Carabus (Apotomopterus) delavayi medea Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) pustulifer guerryi Born, 1903
Female adult
N Yunnan: Gaoligongshan, h =1300 m, 06.2003 Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) gemmifer gemmifer (as loczyi Csiki, 1927)
Female adult
S-China, W-Yunnan: Dali city, Mt Jizushan, VI.2018, L=34 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus vicarius Cavazzuti, 1996
Male adult
China, S Sichuan, / Yunnan border, W of Lugu lake, S of Yongning vill., h=2700m, 11-18.VI.2017, L=31 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) ludivinae ludivinae Deuve, 1996
Male adult
S-China, SW-Yunnan: Lincang city, Mt. Daxue, 18.IV.2017, L=29 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
Endophallus
S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036
Copyright © Ruslan Panin

Carabus (Apotomopterus) madefactus Cavazzuti, 1997
Male adult
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=27 mm
Copyright © Ruslan Panin

Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
Copyright © Ruslan Panin

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
China, N Yunnan: Baimaxue Shan, S of Deqen, h=4300m, 28°20'N 99°03'E, 16-20.VI.2016, C.Reuter leg., L=18 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
Copyright © Ruslan Panin

Cylindera (Eugrapha) minuta (Olivier, 1790)
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 7-11.06.2011 Anichtchenko A. leg.
det. J. Wiesner 2011
Copyright © Alexander Anichtchenko

Cicindela (Calochroa) holzschuhi Wiesner, 2012 Holotype
Male adult
NE Laos, Hua Phan prov., Ban Saleui, Phou Pan (Mt), 20°12’N, 104°01’E, 10.V.-16.VI.2009, 1300-1900 m, leg. C. Holzschuh (in coll. Wiesner).
photo Peter SCHЬLE
Copyright © Peter Schüle

Neocollyris (Leptocollyris) variitarsis
Female adult
Vietnam North, Cao Bang Prov., vic. Vin Den, Nui Pia Oac National Res., 900-1300 m, leg. A.Skale, 6.-10.V.2013; det. J.Wiesner, 2013<br /> Photo: U.Schmidt, 2013<br /> Size: 11,5 mm
Copyright © Udo Schmidt

Cylindera (Eugrapha) mutata (Fleutiaux, 1893)
Female adult
Vietnam North, Bac Kan Prov., Ba Bè National Park, 180-220 m, leg. A.Skale, 16.-20.V.2014; det. J.Wiesner, 2014; Photo: U.Schmidt, 2014. 10,7 mm
Copyright © Udo Schmidt

Heptodonta ferrarii ferrarii Gestro, 1893
Male adult
Laos, Xiang Khuang Prov. leg. local collector, VI.2013; det. J.Wiesner, 2014. Photo: U.Schmidt, 2014. 13,3 mm
Copyright © Udo Schmidt

Cylindera (Eugrapha) minuta (Olivier, 1790)
Female adult
Indonesia, East Java, Mt. Argopuro, Aug. 2011. 8 mm; det. F. Bötzl, 2020
Copyright © Alexander Anichtchenko

Neocollyris (Pachycollyris) bipartita (Fleutiaux, 1897)
Male adult
NE Laos, Hua Phan prov., Mt. Phu Pane, 900-1500m, 10-21.6.2010. det. A. Anichtchenko 2017
Copyright © Alexander Anichtchenko

Myriochila (Myriochila) specularis specularis
Male adult
China, Yunnan (JWGC). det. J. Wiesner
Copyright © Peter Schüle

Cylindera (Cylindera) delavayi (Fairmaire, 1886)
Female adult
N Laos, 20km NW Louang Namtha. 900-1100m, 5-30.v.1997 (cJW) 8.5 mm
Copyright © Alexander Anichtchenko

Cylindera (Eugrapha) mutata (Fleutiaux, 1893)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. & Mainda T. leg. (cAA)
Copyright © Alexander Anichtchenko

Zuphium olens (P.Rossi, 1790)
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 27-30.VI.2006 K.Makarov leg.
L=8.7 mm
Copyright © Kirill Makarov

Planetes (Planetes) puncticeps Andrewes, 1919d: 480
Male adult
Collection ground Tsushima, Nagasaki pref. 35695
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Dendrocellus nepalensis (Hope, 1831)
Male adult
det. R. Sciaky
Copyright © Riccardo Sciaky

Megadrypta maozhoui Chen, Shi & Liang, 2025 Holotype
Male adult
“Sichuan, Qionglai, Mt. Tiantaishan, Sandaowan, 1246 m, 2024.V.1-5, N30.2674 E103.1199, Maozhou XU & Tianxuan GU leg.”; “HOLOTYPE Megadrypta maozhoui sp. nov., det. Chen & Shi. 2024” [red label]; IZAS.
Copyright © - - -

Chydaeus (Chydaeus) asetosus Kataev & Kavanaugh, 2012 Holotype
Male adult
China, W Yunnan, NE Fugong, 26°57'09"N, 98°54'00"E – 26°56'30"N, 98°55'14"E, 2160–2690 m,<br /> 29.V.2006, I. Belousov & I. Kabak leg. <br /> (Deposited in IOZ)
Copyright © - - -

Acupalpus (Stenolophidius) inornatus Bates, 1873
Male adult
Acupalpus (Palcuapus) inornatus Bates, 1873 Collection ground Kamogawa city, Chiba Pref. 34922 (C) Working Group for Biological Indicator Ground Beetles Database, Japan Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Stenolophus (Egadroma) nitens (Motschulsky, 1864)
Female adult
4,7 mm, India, Sьdgoa, District Canacona, Raj Baga Beach, am Licht
leg. U.Schmidt, 16.-29.XI.2010; det. Kataev, 2011
Photo: U.Schmidt, 2012

Copyright © Udo Schmidt

Dioryche yunnana Kataev, 2002
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Rhysopus klynstrai Andrewes, 1929 Paratype
Male adult
Java. Cotype. Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -

Harpalus (Nephoharpalus) pallidipennis A. Morawitz, 1862
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Hyloharpalus) tibeticus Andrewes, 1930
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Zangoharpalus) praticola Bates, 1891
Male adult
Image from Kataev, 2023
Copyright © - - -

Kareya laosensis (Jedlicka, 1966)
Male+Female adult
Laos. Scale 1 mm. Image from Kataev, 2015
Copyright © - - -

Physodera eschscholtzii Parry, 1849
Male adult
Laos, Khammouang Prov., Ban Khounkham (Nahin), 18o13'N 194o31'E h~200 m, 16-17.IV.2005 ltg. O.Gorbunov, A.Zamesov
L=11.0 mm
Copyright © Kirill Makarov

Ophionea (Ophionea) indica (Thunberg, 1784)
Male adult
Thailand, 10 km W Han-Kha, N Suphan-Bur V.1991
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Dromius (Klepterus) hauserianus Lorenz, 1998 Holotype
Male adult
Lebidromius hauseri Jedlicka, Type
China, Prov. Yunnan, Vallis flumin, Soling-Ho (NMPC)
Copyright © Alexander Anichtchenko

Catascopus (Catascopus) facialis facialis (Wiedemann, 1819)
Male adult
N INDIA, Uttarakhand, 23-26.4.2012, 5 km N Ramnagar valley of Kosi riv. N29°25'56.50", E79°8'21.40" Anichtchenko A. leg. & det.; 14 mm
Copyright © Alexander Anichtchenko

Ophionea (Ophionea) indica (Thunberg, 1784)
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 19-23.06.2011 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Physodera dejeani Eschscholtz, 1829
Male adult
China, Yunnan<br /> scale bars = 2.0 mm<br /> Image from Shi H., Zhou H. & Liang H., 2013
Copyright © - - -

Lebidia bioculata A. Morawitz, 1863: 29
Male adult
India. det. A. Anichtchenko, 2015
Copyright © Alexander Anichtchenko

Acoptodera (Acoptodera) piligera (Chaudoir, 1883)
Male adult
Above Tura, Garo Hills, Assam, 3500-3900ft, 15.vii-30.viii1917 (NHMUK)
Copyright © Alexander Anichtchenko

Amphimenes (Amblops) marginicollis Fedorenko, 2019
Male adult
China, Yunnan/Honghe, Qibaofeng, 1350m, 23°4’4.2N 103°23’46.9E, 17.vi.2018, A. Weigel leg. (NME)
Copyright © Alexander Anichtchenko

Lachnoderma biguttatum Bates, 1892
Male adult
S Vietnam, Bin Phuoc pr., Bu Gia Map NP., 16-20.iv.2024, 12.11.38N, 107.12.21E, 450-600m, A. Weigel leg. (NME). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Chlaenius (Achlaenius) micans (Fabricius, 1792)
Male adult
China, North Guizhou prov., Mayanghe, Daheba village, 2007.10.1, Liu ye leg.;
photo & det. by liuye
Copyright © Ye Liu

Chlaenius (Chlaeniostenus) circumdatus Brulle, 1835
Male adult
China, Shichuan pro., 2007, Bi W.X. coll.;
photo by Bi Wenxuan
det. by liuye
Copyright © Bi W.X.

Chlaenius (Chlaeniostenus) punctatostriatus Chaudoir, 1856: 244
Male adult
China, Yunnan pro., Gaoligongshan
photo by liuye
det. by Liang H.B.
Copyright © Ye Liu

Chlaenius (Chlaeniostenus) rugulosus Nietner, 1857
Female adult
China, Yunnan, Hekou, Jinshahe, 22°53'23.7" 103°52'00.7" ,176m,2003.12.19, Liang Hongbin & Kataev coll.
det. by Yu
photo by liuye
Copyright © Ye Liu

Chlaenius (Chlaenius) binghami Andrewes, 1919
Male adult
China, Yunnan pro., Jinkan
liuye photo
det. by Yu peiyu
Copyright © Ye Liu

Hololeius ceylanicus Nietner, 1856
Male adult
China, Yunnan pro., Jinhong
det & photo by liuye
Copyright © Ye Liu

Chlaenius (Haplochlaenius) costiger pecirkai Jedlicka, 1932
Male adult
<p> China,Yunnan Pro., Fugong town, 26.88897N,98.86558E, 1230m, 2004.4.21, Liang H.B. coll.;<br /> photo & det. by liuye</p>
Copyright © Ye Liu

Lachnocrepis japonica Bates, 1873
Male adult

Copyright © Kirill Makarov

Chlaenius (Lithochlaenius) propeagilis Liu & Kavanaugh, 2011 Holotype
Male adult
Southwest Yunnan, Gongshan, Dulongjiang, 0.5km N of Dizhengdang, 28.08442,98.32652”/ "1880 m, 2004.10.29, David Kavanaugh Coll.
(IZCAS)
Copyright © Ye Liu

Chlaenius (Chlaenioctenus) rotundithorax Liu & Kavanaugh, 2010 Holotype
Male adult
“Yunnan Prov., Tengchong, Jietou, Datang, Dahelingganjiao, 25.73947°N,98.69630°E’’/ “2010m, 2006.5.15 day, Kavanaugh D., Brett R. California Academy & IOZ, Chinese Acad.Sci.”/ “HOLOTYPE, Chlaenius (Chlaenioctenus) rotundithorax n. sp., designated by Y. Liu & D. Kavanaugh 2009”
Copyright © Ye Liu

Paropisthius indicus chinensis Bousquet et Smetana, 1996: 225 Paratype
Male adult
China, Centr. Sichuan, Luding Co., Moxi env., 22.V-10.VI. 1993, Benes (cAA)
Copyright © Alexander Anichtchenko

Trichisia violacea Jedlicka, 1935a: 2
Male adult
Philippines, E Visayas, Samar, Lope de Vega, XII.2016
Copyright © Alexander Anichtchenko

Itamus castaneus Schmidt-Goebel, 1846
Male adult
Thailand, Nakhon Prov., Ratchasima (Korat), Wat Soeng Sang, near Non Sambun Village, 260 m asl, 14°16,5' N, 102°23,3' E, 4.08.2009, V.K. Zinchenko leg
D. Fedorenko det. 2011
Copyright © Roman Dudko

Paussus (Scaphipaussus) hystrix Westwood, 1850
Male adult
NE Laos Phu Pane Mt. (DUBC). det. Anichtchenko A. 2023
Copyright © Alexander Anichtchenko

Dicranoncus femoralis Chaudoir, 1850c: 393
Male adult
SE Maritime Prov., Lazovsky dist., Lazo env., attracted to light, 18.VI.2010 leg. M.Smirnov
Copyright © Maksim Smirnov

Dolichus halensis Schaller, 1783: 317
Male adult
S Ukraine, Krym Peninsula,
Simpheropil, 15-30.VIII.2009, Shaporinskiy V. leg.
male, 19 mm
Copyright © Ruslan Panin

Agonum (Olisares) impressum (Panzer, 1796)
Female adult

Copyright © Kirill Makarov

Sericoda quadripunctata (Degeer, 1774)
Male adult

Copyright © Kirill Makarov

Metacolpodes buchanani Hope, 1831: 21
Male adult

Copyright © Kirill Makarov

Xestopus gutangensis Zhu & Kavanaugh, 2021 Holotype
Male adult
Xizang, Nyingchi Prefecture, Mêdog County, Gutang Township, Xingkai village, 29.46414°N, 95.73563°E”; “2025 m, 2019.VIII.15 N, Liang H.B. & Xu Y. lgt., (Institute of Zoology, IZAS). 16.9 mm
Copyright © David Kavanaugh

Onycholabis stenothorax Liang & Kavanaugh, 2005
Male adult
S Vietnam, Lam Dong Pr., Lac Duong Distr., Bidoup Nui Ba NP, 1450-1650m, 22-28.4.2024 A. Scale leg. (Mus. Erfurt). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Nirmala odelli Andrewes, 1930
Male adult
China, Yunnan pro., Gaoligong Mountain, Liang H.B. coll.;
Photo by Liuye
Copyright © Ye Liu

Amara (Amara) ovata (Fabricius, 1792)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Trigonotoma loeffleri Kirschenhofer, 2007 Holotype
Male adult
N. Thailand (Chiang Mai prov.) Doi Pha Hom Pok, Mt. Mae Ai, 2000 m, 22.-27.IV. 2004, S. Löffler“ (CDW). Image from KIRSCHENHOFER, 2007
Copyright ©

Pterostichus (Gutta) kongshuhensis Gueorguiev, 2015 Holotype
Male adult
CHINA, Yunnan Prov., Tengchong County, vil. Kongshuhe near cave Wu Shi Shan 1-2-3 N25.73111 E98.66459, ca. 2500-2600m, 11.11.2013, B. Petrov leg.
Copyright © Borislav Gueorguiev

Trigonognatha delavayi (Fairmaire, 1888)
Female adult
China, NW Yunnan, Lijiang pref., S Yulongxue Shan 3200 m., 16.7.2016, V. Patrikeev leg.; 16 mm; det. R. Sciaky, 2018
Copyright © Alexander Anichtchenko

Trigonognatha fairmairei Sciaky, 1995
Male adult
China, W Sichuan, Kanding env. Xinduchuao 3900 m, 15.7.2013, V.Patrikeev leg.; 20 mm ; det. R. Sciaky, 2018
Copyright © Alexander Anichtchenko

Myas (Trigonognatha) becvari Sciaky, 1995
Male adult
China, NW Yunnan, Shangri La, 3400 m. 17.5 mm. det. R. Sciaky, 2018
Copyright © Alexander Anichtchenko

Myas (Trigonognatha) echarouxi Lassalle, 2010
Male adult
China, N Yunnan, Baima pass between Dequen and Shangri La 4700m, 15.6.2016, V. Patrikeev leg. (cAA). det. R. Sciaky, 2019
Copyright © Alexander Anichtchenko

Aristochroa venusta Tschitscherine, 1898
Male adult
China, NW Sichuan, Luhuo town, 3500 m, leg. V. Patrikeev (cAA) 14 mm; det. Hongliang Shi, 2020
Copyright © Alexander Anichtchenko

Amara (Amara) ovata (Fabricius, 1792)
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. & det. (cAA). 8.5mm
Copyright © Alexander Anichtchenko

Yamatosa reuteri Hovorka, 2018 Holotype
Male adult
China, Yunnan, SW / Jingdong, Maotou Shan / 24°23’ N, 100°45’ E / 1300-1600 m, 16.-18. VI. / 2014, leg. C. Reuter (OHPC). Image from Oldřich HOVORKA, 2018
Copyright © - - -

Clivina (Leucocara) tranquebarica Bonelli, 1813
Female adult
Size: 6,1 mm
Location: India, South Goa, Distr. Canacona, Raj Baga Beach
leg. U.Schmidt, 16.-29.XI.2010; det. P.Bulirsch, 2011
Photo: U.Schmidt, 2013
Copyright © Udo Schmidt

Scarites (Parallelomorphus) semicircularis W.S. MacLeay, 1825
Male adult
Vietnam, Da Nang prov., 1450m, Ba Na Mt., 5.2015. 16 mm; det. P. Bulirsch, 2018
Copyright © Alexander Anichtchenko

Kozlovites major Belousov et Kabak, 2016 Paratype
Male adult
pass 25 km SE Dequin
Copyright © - - -

Queinnectrechus (Gaoligongtrechus) balli Deuve and Kavanaugh, 2016 Holotype
Male adult
CHINA, Yunnan Province, Gaoligong Shan, Nujiang Prefecture, Nujiang State Nature Reserve, Dulong/Gongshan Yakou [= Qiqi/Dulong divide] area, 21 airkm w of Gongshan,”/ “N27.69655°/ E98.45389°, 3300-3680m, 16-17 July 2000, Stop#00-24C, D.H. Kavanaugh, C.E. Griswold, Liang H.-B., D. Ubick, & Dong D.-Z. collectors (IOZ). Image from Deuve et al, 2016
Copyright © - - -

Epaphiotrechus fortipesoides Deuve & Kavanaugh, 2016 Holotype
Female adult
China, Yunnan, Tengchong County, Houqiao Township, 8.5 airkm NNE of Houqiao at Gaoshidong, N25.39858°/ E98.30533°, 2580 m. Image from Deuve et al, 2016
Copyright © - - -

Trechepaphiama gaoligong Deuve & Kavanaugh, 2016 Holotype
Female adult
China, Yunnan, Gaoligong Shan, Lushui County, 9.3 km ESE of Pianma, N25.99363°/E98.66651°, 2460-2470 m. Image from Deuve et al, 2016
Copyright © - - -

Trechepaphiopsis monochaeta Deuve & Kavanaugh, 2016 Holotype
Male adult
China, Yunnan, Gongshan County, Qiqi Trail at No. 12 Bridge Camp area, 16.3 airkm W of Gongshan, N27.71503°/E98.50244°, 2775 m. Image from Deuve et al, 2016
Copyright © - - -

Sinechostictus (Pseudolimnaeum) alesmetana Toledano, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Yunotrechus diannanensis Tian & S.Huang, 2014 Holotype
Male adult
cave named Shiyakou Dadong, 23°06′54″N, 104°17′58″E, 1320 m, Shiyakou Cun, Pojiao Xiang, Maguan County, 3. VII.2013, leg. Mingyi Tian, Wei Lin, Weixin Liu, Haomin Yin and Sunbin Huang (SCAU). Image from Tian & S.Huang, 2014
Copyright © Mingyi Tian

Shilinotrechus zhangfani Tian, Huang & Jia, 2023 Holotype
Male adult
cave Xianren Dong, Xi’er Township, Mile Shi, Honghe Hani & Yi Autonomous Prefecture, Yunnan Province, 103.20 E,24.48 N, 1901 m above sea level, 2020-VIII-17, Mingyi Tian & Jingli Cheng leg., in SCAU.
Copyright © Mingyi Tian

Yunaphaenops graciliphallus Tian & He, 2023 Holotype
Male adult
Fengwei Dong Cave, Wenliu Xiang, Qiubei Xian, Wenshan Zhuang & Miao Autonomous Prefecture, eastern Yunnan, 24.33° N / 104.29° E, 1302 m, 3.IX.2020, leg. Chenghai Fu (SCAU).
Copyright © Mingyi Tian

Aspidaphaenops qujingensis Deuve & Tian, 2020 Holotype
Male adult
Chine, Yunnan, Préfecture de Qujing, Luoping Xian, Machang, grotte Shuiyuan Dong, 1530 mètres, 24°49’33’’N 104°21’48’’E (Tian M. Y., Lui W. X., Wang X. H., Tang M. R. leg., 18.VI.2015), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve