Apotomus rufithorax Pecchioli, 1837
Male adult

Copyright © Kirill Makarov

Brachinus (Cnecostolus) bayardi Dejean, 1831
Female adult
E Turkmenistan, Charjev env., V.1996, V.Perepel leg.
12 mm
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sclopeta (Fabricius, 1792)
Male adult
S Bosnia, Mostar; 6mm
Copyright © Aleš Sedláček

Brachinus (Brachynolomus) nigricornis Gebler, 1829
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinus) elegans Chaudoir, 1842
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Cnecostolus) bipustulatus Quensel In Schoenherr, 1806
Male adult
Greece, Etolia Akarnania, Neohori, 1.5.1999. F. Angelini leg.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sichemita Reiche & Saulcy, 1855
Male adult
Greece, Ahaia, Kalavryta, 28-29.4.1999, leg. F. Angelini
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) alexandri F. Battoni, 1984
Male adult
SE Azerbajan, Talysh, Mistan v., 7-10.10.1993. det. A. Anichtchenko
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) explodens Duftschmid, 1812
Female adult
Bulgaria Chernomorets, 24.11.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Female adult
W Ukraine, Lviv city, Sykhiv (Zubra) forest, 49.791889, 24.047417, 13.VII.2024, Panin R. leg.
Copyright © Ruslan Panin

Broscus nobilis Dejean, 1828: 432
Male adult
Greece, Isl. Rhodos, 10 km NNW Lindos 10-20m, 18.4.1994 A Korell leg.
17 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt., 6-17.VII.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Procrustes) banoni Dejean, 1829
Male adult
Crete, Agia Irini (Sélinos); 31mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Female adult
SW Romania, Banat; 15mm
Copyright © Aleš Sedláček

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
Copyright © Noël Mal

Carabus (Pachystus) hortensis preslii Dejean, 1830
Male adult
S-Greece:, N Peloponnes, Erymanthos Reg., Kalentzi, 26.VI.2007, Manfred Egger leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Female adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 38 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Pachystus) graecus morio Mannerheim, 1830
Female adult
W Turkey, Manisa Prov., 35 rm SEE Salihc, 38°20'N, 28°24'E, 30.VI.2006, J.Halada leg. L=25 mm
Copyright © Ruslan Panin

Calomera littoralis nemoralis Olivier, 1790
Male adult
UKRAINE,
Kherson Reg.,Genitchesk Distr., near Tehongar st., Kujuktuk isl., on saline 8-9.V.1995 Mishustin R. leg.
16 mm.
Copyright © Ruslan Panin

Calomera aulica aulica s.str.
Male adult
Oman, Dhofar, Jabal al Qamar, 5 km NE Dhalqut, 16°43'22.48’’N 53°16’27.26’’E, 22.9.2011, Richard Ambrus lgt.
11 mm
Copyright © Alexander Anichtchenko

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
CZ, C Moravia, Přerov; 9mm
Copyright © Aleš Sedláček

Calomera concolor concolor Dejean, 1822
Female adult
14,0 mm; Greece, Crete, Nom. Hanion. Georgioupolis, 4 km E, Kavros, leg. J. Scheuern, 4.-12.X.1999; det. J.Schoenfeld, 2000; Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Megacephala (Grammognatha) euphratica euphratica
Unsexed
Spain : Alicante, Salina la Mata, 15.vii.1979 - E. Vives leg. (Size : 24 mm) Coll. N. Mal
Copyright © Noël Mal

Cylindera (Eugrapha) trisignata hellenica (Cassola, 1973)
Male adult
SW Ukraine, Odessa region, Tatarbunary distr., Lebedivka vill., sea beach, 29.VII-6.VIII.2021, Yu. Kanarsky leg. /det. R.Panin, L=10.5 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg., L=12 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Male adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg.
Copyright © Ruslan Panin

Polistichus fasciolatus (P. Rossi, 1790)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr., near Bogatoe Ushchele vill., 04.VIII.2009, Shehovcov A. leg.
L=8mm
photo Panin R.
Copyright © Ruslan Panin

Zuphium olens (P.Rossi, 1790)
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 27-30.VI.2006 K.Makarov leg.
L=8.7 mm
Copyright © Kirill Makarov

Drypta (Drypta) dentata (P.Rossi, 1790)
Male adult

Copyright © Kirill Makarov

Parazuphium (Parazuphium) damascenum (Fairmaire, 1897)
Male adult
Morocco, Tagounite vill area 28.6.2010, Anichtchenko A. leg. & det. (aedeagus studied)
Copyright © Alexander Anichtchenko

Parazuphium (Parazuphium) chevrolatii chevrolatii Laporte, 1833a: 254
Male adult
Iran Gilan Elburz 1000m 5.viii.1968 W.Heinz leg. at light. J. Muilwijk det.
Copyright © Jan Muilwijk

Odontocarus robustus (Dejean, 1831)
Male adult
Greece, Peloponnes, Oros Menalon
occ. Dimitsama 900-1000m 12-23.5.1992 A.Korell leg.
17 mm
Copyright © Alexander Anichtchenko

Bradycellus (Bradycellus) caucasicus Chaudoir, 1846: 187 (Acupalpus)
Male adult
L=4,0 mm
RUSSIA, Tver reg., Likhoslavlskiy distr., Domanikha village, in the garden, in soil. H=160 m. N57*13.629’ E035*45.675’ 21.VII.2005
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Pangus scaritides (Sturm, 1818)
Male adult
SK, S Slovakia, Velké Kosihy; 8mm
Copyright © Aleš Sedláček

Carterus (Pristocarterus) angustipennis lutshniki Zamotajlov, 1988
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Penthus tenebrioides (Waltl, 1838)
Male adult

Copyright © Kirill Makarov

Harpalus (Semiophonus) signaticornis (Duftschmid, 1812)
Male adult
France, 77860, Quincy-Voisins, 12-05-2018, leg. & det. P. Deschamps
Copyright © Pascal Deschamps

Ophonus (Metophonus) laticollis Mannerheim, 1825
Male adult
Russia, NW Caucasus, Novorossiisk distr., Markoth Mts., 23.5.2005 (DUBC) det. Anichtchenko A. 2021; 10.5mm
Copyright © Alexander Anichtchenko

Bradycellus (Bradycellus) verbasci (Duftschmid, 1812)
Male adult
W Ukraine, Lviv city, Holosko Cemetery, 3.IX.2014, Panin R. leg. L=5.1 mm
Copyright © Denis Khrapov

Dicheirotrichus (Pelagophilus) punicus Bedel, 1899
Female adult
Espagne, Guardias Viejas, Almeria 04-05-1991
Copyright © Pascal Deschamps

Lebia (Lebia) trimaculata (Villers, 1789)
Male adult
S Ukraine, Krym Peninsula,
Simpheropil, 1-20.VI.2009, Shaporiskiy V. leg.
L.= 5.5 mm
Copyright © Ruslan Panin

Metadromius signifer (Reitter, 1884)
Male adult
Dzhilikul, tugai forest, Tadzh. 15.5.61 [ex coll. Khnzorian]
L=3.8
Copyright © Kirill Makarov

Cymindis (Cymindis) lineata (Quensel in Schonherr, 1806)
Male adult
S Ukraine, Crimea,
Cape Tarkhankut, Site Djangul, 2-3.VII.2010, Shaporinskiy V.leg.
L=10 mm

Copyright © Ruslan Panin

Glycia spencei (Gistel, 1838)
Male adult

Copyright © Kirill Makarov

Cymindis (Cymindis) angularis Gyllenhal, 1810
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Cymindis (Cymindis) discoidea Dejean, 1824
Male adult
Spain, Sierra Nevada, puerto de la Ragua, 2000m, 15.I.2004, leg. & det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Cymindis (Tarulus) vaporariorum (Linne, 1758)
Male adult

Copyright © Kirill Makarov

Apristus reticulatus Schaum, 1857
Male adult
Азербайджан, окр. п.Аврора, Ханбулакское вдхрн. 1.VI.2018 leg. Б.Саратовских. det. Anichtchenko A., 2018
Copyright © Kirill Makarov

Paradromius (Manodromius) linearis (Olivier, 1795)
Male adult

Copyright © Kirill Makarov

Metablus paracenthesis (Motschulsky, 1839)
Female adult
Azerbajan, Shirvan NP, 39.40N, 49.11E, 23.6.2024 Lason A. leg. (cAA). det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Licinus (Licinus) cassideus Fabricius, 1792: 148 (Carabus)
Female adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr.,near Bogatoe Ushchele vill.04.VIII.2009, Shehovcov A. leg.
female, 17mm
photo & det. Panin R.
Copyright © Ruslan Panin

Licinus (Licinus) silphoides (P. Rossi, 1790)
Female adult
Corse, Col de Bavella, 14.8.1966
Copyright © Alexander Anichtchenko

Notiophilus rufipes Curtis, 1829: pl. 254
Female adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Nebria (Eunebria) jockischi jockischii Sturm, 1815
Male adult
SE France, Alpes-Maritimes, Mt. Col de Turini h=1500m, Graben SW, 18.VIII.2005, Manfred Egger leg.
L= 13 mm

Copyright © Ruslan Panin

Nebria (Tyrrhenia) holtzi K. Daniel, 1903a: 162
Female adult
Greece, N-Peloponnese, Kalentzi vill., Mt. Enimanthos, 11.VI.2006, Manfred Egger leg.
L=13 mm
Copyright © Ruslan Panin

Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843
Male adult
SW Bosnia, Bijakoviči - Križevac Mt.; 8mm
Copyright © Aleš Sedláček

Nebria (Nebria) kratteri pindica Jeanne, 1974
Male adult
C Greece, Thessalia, Kastraki; 10mm
Copyright © Aleš Sedláček

Leistus (Pogonophorus) rufomarginatus (Duftschmid, 1812)
Male adult

Copyright © Kirill Makarov

Paussus (Cochliopaussus) turcicus Frivaldszky von Frivald, 1835
Male adult

Copyright © Kirill Makarov

Paussus (Cochliopaussus) turcicus Frivaldszky von Frivald, 1835: 263
Male adult
Iran, Hamadan env., Ali Sadr 14.5.1999, leg. Orszulik
Copyright © Alexander Anichtchenko

Agonum (Olisares) lugens (Duftschmid, 1812)
Male adult
Volgograd Reg., Pallasovka Distr., N shore of Elton Lake, right bank of Khara River 3.V.2009 leg.K.Makarov
8.8 mm
Copyright © Kirill Makarov

Omphreus (Paromphreus) krueperi Reitter, 1885
Female adult
S Greece, Taygetos Mts.; 28mm
Copyright © Aleš Sedláček

Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763)
Male adult

Copyright © Kirill Makarov

Atranus ruficollis Gautier des Cottes, 1858b
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Agonum (Agonum) antennarium (Duftschmid, 1812)
Male adult
W Ukraine, NE Carpathians, Chornogora Mt Rng, h=1650 m, source of the creek, 6.VI.2011, R. Panin leg. L=8.5 mm
Copyright © Ruslan Panin

Agonum (Agonum) menetriesii Faldermann, 1839
Male adult
S Kazakhstan, 15 km SE Kyzyl Orda, 30.4.2012, A. Ivanov leg.; det. K. Makarov, 2018
Copyright © Alexander Anichtchenko

Olisthopus fuscatus Dejean, 1828
Male adult
Turkey, Kirish distr., Camyuva riv., 28.9.2014, I. Enuschenko leg. det. Anichtchenko A. 2018 (aedeagus studied)
Copyright © Alexander Anichtchenko

Agonum (Olisares) hypocrita (Apfelbeck, 1904)
Male adult
NW-Ukraine, Polesia, Volhynia, Lyubeshiv Distr., Buchyn vill, swamped pinewood, «Selysko», 14.IX-17.X.2017, Kanarsky Yu. leg./det.Panin R.
Copyright © Ruslan Panin

Omphreus (Elladomphreus) eggeri Monzini & Sciaky, 2025 Paratype
Male adult
GR. Peloponnes / Panachaikogeb. / Kalanistra V.2014, leg. Manfred Egger
Copyright © Riccardo Sciaky

Nomius pygmaeus Dejean, 1831A: 512
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Pterostichus (Platysma) niger niger s.str.
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.2007 Panin R. leg.

Copyright © Ruslan Panin

Amara (Amara) saphyrea Dejean, 1828: 463
Female adult
S Slovakia, Štúrovo; 9mm
Copyright © Aleš Sedláček

Amara (Xenocelia) ingenua Duftschmid, 1812: 110 (Carabus)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Stomis (Stomis) pumicatus (Panzer, 1796)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Tapinopterus (Tapinopterus) duponchelii (Dejean, 1831)
Male adult
Mt. Parnone, Kosmas Peloponneso, Greece
Copyright © Vittorino Monzini

Amara (Amara) tibialis (Paykull, 1798)
Male adult
W Ukraina, Podolian Upland, Kasova Gora near Burshtyn vill., steppe site, 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=5 mm
Copyright © Denis Khrapov

Platyderus (Eremoderus) chatzakiae Guéorguiev, Wrase, Assmann, Muilwijk & Machard, 2022 Holotype
Male adult
8422, KALYMNOS Stimenia, end / road st18 9/6/2005, Lg Chatzaki (NHMC)
Copyright © Borislav Gueorguiev

Dodecanopterus laticornis rhodius (Straneo, 1935) Paratype
Male adult
Rhodos
Copyright © Vittorino Monzini

Elasmopterus janisjoplinae Giachino, Allegro & Vailati, 2024 Holotype
Male adult
Grecia, nom. Fthiótida, O. Kallídromo m 900, 9.VI.1994/8.VI.1995, Giachino & Vailati leg. (CGi).
Copyright © Gianni Allegro

Clinidium (Arctoclinidium) canaliculatum O.G.Costa, 1839d
Male adult
photo from www.zin.ru
Copyright © Kirill Makarov

Scarites (Parallelomorphus) laevigatus Fabricius, 1792
Male adult
SW Ukraine, Odessa region, Tatarbunarsk distr., near Primorskoe vill.,in the sand, 8.VII.2004,
A Gontarenko leg.
L=16 mm

Copyright © Ruslan Panin

Clivina (Clivina) fossor fossor (Linnaeus, 1758)
Male adult
L=5,3 mm
RUSSIA, Moscow city, SW distr., Jasenevo, Bitcevskiy les, under tree. 19.V.2000
leg., det., photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Dyschirius (Dyschiriodes) cylindricus (Dejean, 1825)
Male adult

Copyright © Kirill Makarov

Clivina (Clivina) ypsilon Dejean, 1830
Male adult
NE Greece, Nei Pori; 6 mm
Copyright © Aleš Sedláček

Distichus (Distichus) planus (Bonelli, 1813)
Male adult
Turkey, Side, Colakli, leg. J.Scheuern, 24.IV.2000; det. Balkenohl, 2004. Photo: U.Schmidt, 2014. 12,8 mm
Copyright © Udo Schmidt

Dyschiriodes (Eudyschirius) globosus Herbst, 1784: 142
Male adult

Copyright © Kirill Makarov

Scarites (Scallophorites) buparius (Forster, 1771)
Male adult
Italie, Sicile, Selinunte, 04-07-1983
Copyright © Pascal Deschamps

Dyschiriodes (Eudyschirius) rufipes (Dejean, 1825)
Female adult
W Ukraina, Podolian Upland, Opillia, Ivano-Frankivsk reg., Kasova Gora near Burshtyn vill., steppe site, 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=2,9mm
Copyright © Denis Khrapov

Dyschiriodes (Dyschiriodes) agnatus Motschulsky, 1844: 79
Male adult
W Ukraine, NE Carpathians, Lviv reg., Stryi distr., near Rozgirche vill., bank of Stryi river, 18.VI.2021, D.&O. Khrapov leg./det. R. Panin L=5mm
Copyright © Denis Khrapov

Reicheadella (Reicheadella) corcyrea (Reitter, 1884)
Male adult

Copyright © Vittorino Monzini

Siagona europaea Dejean, 1826: 468
Male adult
Uzbekistan, Kashkadarjnsk Reg., env. Mubarek 5-8.VI.2004.
L=10.3 mm
Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) biguttatum (Fabricius, 1779)
Male adult

Copyright © Kirill Makarov

Porotachys bisulcatus (Nicolai, 1822)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Perileptus (Perileptus) areolatus areolatus Creutzer, 1799
Male adult

Copyright © Kirill Makarov

Odontium (Odontium) striatum (Fabricius, 1792)
Female adult
Latvia, Dunava
Copyright © Alexander Anichtchenko

Bembidion (Testedium) laetum Brulle, 1836
Male adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 3.9 mm
Copyright © Kirill Makarov

Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko