Carabus (Carabus) nanosomus wufengensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) vigilax rautenstrauchi Rapuzzi, 2018 Holotype
Male adult
China, Sichuan, Leibo, Shanlinggang environs, VII.2018, Wang legit in collection Ivan Rapuzzi. Image from Rapuzzi, 2018
Copyright ©

Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Female adult
China, S-Sichuan, SE Mianning, Pass btw Mianshanzhen & Yuexi 28°30’N/102°26’E, 22-23.VI.2017, Ch.Reuter leg., L=22 mm; det. B. Brezina, 2020
Copyright © Ruslan Panin

Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Male adult
China, S-Sichuan, road 56 km E of Xichang to Zhaojue, Mt.WenLin, Pass h=3200 m, 27°52’N/102°30E, 5-20.VI.2017, Ch.Reuter leg.; det B. Brezina, 2020
Copyright © Ruslan Panin

Carabus (Acoptopterus) rautenstrauchi alberti Rapuzzi, 2024 Holotype
Male adult
Sichuan, Gu Dui Xiang, Meigu to Shanlenggang; 2250 m asl; Jul. 2022
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Carabus (Apotomopterus) infirmior homodynamus Deuve, 1994 (as jiucaishan) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) guzhangensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) guizhouensis yunwuensis Kleinfeld, 1996 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) pseudoguizhouensis Kleinfeld, 2006 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) arrowi arrowi Hauser, 1913
Female adult
Simianshan, Guizhou, Chine. 31 mm
Copyright © Alain Mollard

Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
Female adult
SC-China, Guangxi Reg., Pingle County, Mt. Shizishan h=1290 m,5-20,VII.2011, local coll. leg. L=43 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) arrowi trilobatus Cavazzuti, 1994
Male adult
Jinsha, Guizhou, CHINA
Copyright © Alain Mollard

Carabus (Apotomopterus) yuae arlequinus Deuve, 1994
Female adult
Yuanbao Shan, Guangxi, CHINA
Copyright © Alain Mollard

Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
Female adult
Fanjing Shan, Guizhou, 1900 m, Leg: C.Reuter. L= 42 mm
Copyright © Alain Mollard

Carabus (Apotomopterus) longeantennatus acorepoides Deuve, 2018 Holotype
Female adult
Chine, Chongqing, Wulingshan Forest Park, 1690 m, 29°33’N-107°36’E (Jatua), in coll. J. Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve

Carabus (Apotomopterus) protenes erlangbaensis Deuve, 2016 Holotype
Male adult
Chine, Guizhou nord occidental, Erlangba, 1200- 1350 m, 28°33’N-106°00’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve

Carabus (Apotomopterus) rugulothorax Deuve, 2016 Holotype
Female adult
Chine, Guizhou nord occidental, Erlangba, 1200-1350 m, 28°33’N-106°00’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve

Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
Female adult
S-China, NE-Guizhou prov., Fanjing Shan, 1800-2000 m, 5-11.VI.14, 27.54°N 108.42°E , L=40 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes protenes Bates, 1889 Lectotype
Female adult
Ichang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec81
Copyright © MNHN

Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Male adult

Copyright © Igor Solodovnikov

Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Female adult

Copyright © Igor Solodovnikov

Carabus (Apotomopterus) elliptipennis Deuve, 1995
Male adult
SE Guizhou, Leishan Co., Leigong Mt. Image from Hackel & Sehnal, 2004
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Carabus (Apotomopterus) breuningianus maolanensis Imura, Zhou & Su, 1999
Male+Female adult
SE Guizhou, Libo Co., Yaogu env. Image from Hackel & Sehnal, 2004
Copyright © - - -

Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
Male adult
C Guangxi, Guigang Pref., Zhenlongshan Mt. Image from Hackel & Sehnal, 2004
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Carabus (Apotomopterus) guibeicus Deuve & Tian, 1999
Female adult
SE Guizhou, Libo Co., Yaogu env. Image from Hackel & Sehnal, 2004
Copyright © - - -

Carabus (Apotomopterus) protenes buycki Hauser, 1924
Male adult
China, Shaanxi
Copyright © - - -

Carabus (Apotomopterus) protenes baoxingensis Deuve & Murzin, 2003 Holotype
Male adult
Lazarev & Murzin, 2026
Copyright © - - -

Carabus (Apotomopterus) yuae yuae Deuve, 1989 (as arlequinus)
Male adult
Mt. Yuanbaoshan, Rongshui, Guangxi, China, Jun–Aug 2025, leg. local collector (coll. Junle YE). 35.5 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male+Female adult
Mt. Julong, Baokang, Hubei Prov., CHINE
25.VII.2007
Copyright © Alexandr Safronov

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
Chine, SW-Hubei, Enshi, VI.2005 Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
Chine, SW-Hubei, Enshi, 06.2005, Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Female adult
Chine, CE Hunan, Changde 07.2004 h=700m Local people
Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Male adult
Chine, CE Hunan, Changde 07.2004 h=700m Local people<br /> Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) augustus pustuliferus Kleinfeld, 1998 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) augustus cristianofonti Deuve & Font, 1998
Female adult
China, Guizhou Prov., Sinan, 14.V.2009. L= 41mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) augustus adriaenssensi (as pseudoignigena Kleinfeld, 1996) Holotype
Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) augustus yangminshanicus Font, 1997
Male adult
Guangxi prov., Gongcheng county, N. Lianhua, CHINA<br /> 9-17.06.2007, L (mail) = 40 mm, L (female) = 45 mm.
Copyright ©

Carabus (Coptolabrus) augustus yangminshanicus Font, 1997
Female adult
Mt. Yangmingshan, Yongzhou City, Hunan, CHINA<br /> 20-28.06.2007
Copyright ©

Carabus (Coptolabrus) augustus ignigena (as subpustulatus Breuuning 1961) Holotype
Male adult
Hunan, China. ignigena subpustulatus (In coll Lassalle).
Copyright © Bernard Lassalle

Carabus (Coptolabrus) augustus adriaenssensi Lassalle & Prunier, 1993 Holotype
Male adult
Chine, Guangxi, Mts Miao Er Shan, 1800m, 5.92 (in coll B. Lassalle) ? pustulifer adriaenssensi Lassalle & Prunier, 1993
Copyright © Bernard Lassalle

Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Female adult
China : Guizhoh, Sinan vi.2006 (Size : 35 mm) Coll. N. Mal
Copyright © Noël Mal

Carabus (Coptolabrus) augustus tenuitarsatus Rapuzzi, 2016 Holotype
Male adult
China, Chongqing province, Pengshui county, Mt. Heimending. Image from Rapuzzi, 2016
Copyright ©

Carabus (Coptolabrus) pustulifer lengchongensis Deuve, 2016 Holotype
Male adult
Chine, Guizhou nord-occidental, Lengchong, 1900 mètres, 27°06’N-105°15’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve

Carabus (Coptolabrus) augustus ignigena Hauser, 1914 (as lautus)
Male+Female adult
Mt.Yingpan Shan (Qiubei Xian, Yunnan) CHINE, VII-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer aeriviolaceus Cavazzuti & Rapuzzi, 2010
Male+Female adult
Meigu (Xichang, Sichuan) CHINE, VII-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer elisabettianus Font, 1997
Male+Female adult
Huanjan Shan (Wuchuan, Guizhou) CHINE, VII-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer inusitatus Cavazzuti, 1997
Male+Female adult
Fanjing Shan (Jiangkou, Guizhou) CHINE, VI-2007
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer mirandior Deuve, 1993
Male+Female adult
Dayao (Yunnan) CHINE, VII-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer qiannaicus Deuve & Tian, 2000
Male+Female adult
Longzjia (Maoping, Guizhou) CHINE, V-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer szechwanensis Hauser, 1931
Male+Female adult
Jinfo Shan (Chongqing) CHINE, V-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer xianfengensis Deuve & Li, 2009
Male+Female adult
Mt.Maoping (Zhaoshi, Hunan/Hubei) CHINE, V-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus chenzhouensis Deuve & Sehnal, 2010
Male+Female adult
Yizhang Xian (Hunan) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus tongrenensis Deuve & Li, 2006
Male adult
Tongren Xian (Jiangkou, Guizhou) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus haiyangicus Deuve, 2012
Male+Female adult
Xing’an (Guilin, Guangxi) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus guizhouicus Deuve, 1994
Male+Female adult
Leigong Shan (Kaili, Guizhou) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus mugepoensis Deuve & Li in Deuve, 2009
Male+Female adult
Mt.Gupo Shan (Zhongshan, Guangxi) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
SC-Chine, NW-Hunan, Longshan County, Mt. Maoping,10-30.VI.2008, L=44 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) pustulifer jiucailingensis Rapuzzi, 2026 Holotype
Female adult
Hunan Province, Jiucai Ling; 1300 m a.s.l.; 25°32’N, 111°22’E; Jun. 2006; V. Siniaev legit
Copyright © - - -

Carabus (Coptolabrus) pustulifer qiannanicus Deuve & Tian, 2000
Female adult
China, Guizhou Province, Longzija, 1800 m, V.2008, I. Rapuzzi collection. 39 mm
Copyright © - - -

Carabus (Coptolabrus) ignigena tenellus Rapuzzi, 2026 Holotype Paratype
Male+Female adult
Hunan Province, Shaoyang city, Chengbu Co., Nanshan Pasture Longtouping; 1400–1500 m a.s.l.; Jul. 2025
Copyright © - - -

Carabus (Coptolabrus) pustulifer adriaenssensi Lassalle & Prunier, 1993
Male adult
China, Guangxi Province, Mt. Maoershan, 1900 m, VII.2022. I. Rapuzzi collection; 45mm
Copyright © - - -

Carabus (Coptolabrus) ignigena rautenstrauchianus Rapuzzi, 2026 Holotype
Male adult
Hunan Province, Huaihua, Xuefeng Shan; 1500–1800 m a.s.l.; 1 May 2025; 40 mm
Copyright © - - -

Carabus (Coptolabrus) evidens Rapuzzi, 2026 Holotype Paratype
Male+Female adult
Hunan Province, Zhangjiajie, Mt. Tianmen Shan; 1400 m a.s.l.; 10 Jun. 2025; local collector legit
Copyright © - - -

Carabus (Coptolabrus) pustulifer adriaenssensi Lassalle & Prunier, 1993 (as pseudoignigena)
Male adult
Mt. Yuanbaoshan, Rongshui, Guangxi, China, Jun–Aug 2025, leg. local collector (coll. Junle YE). 44 mm (left), 47 mm (right)
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894 (as xianfengensis)
Female adult
Banqiao Town, Enshi, Hubei, China, alt. 3300m, 20 Jun 2024, leg. local collector (coll. Junle YE). 45 and 46 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894 (as xianfengensis)
Male adult
2025 Captive Breeding. Zhongshan Town, Wanzhou, Chongqing, 30.512734°N 108.530713°E, (coll. Junle YE). 39.5 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer adriaenssensi Lassalle & Prunier, 1993 (as tenellus)
Female adult
border area of Ziyuan, Guangxi and Chengbu, Hunan, China, Jun–Aug 2025 leg. local collector (coll. Junle YE). 40.5mm and 39 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) augustus adriaenssensi Lassalle & Prunier, 1993
Male adult
42mm; Mt. Mao'ershan, Ziyuan, Guangxi, China, alt. 1700m, Apr 2026, leg. local collector (coll. Junle YE)
Copyright © Lokin Yeh

Carabus (Coptolabrus) augustus adriaenssensi Lassalle & Prunier, 1993
Male adult
41mm; Shuiling Town, Dong'an, Hunan, China, May 2025 (coll. Junle YE)
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer wenshuicus Font, 1998
Male adult
38mm; Mt. Yunlongshan, Dafang, Guizhou, China, alt. 1700-1900m, Apr-May, 2026, leg. local collector (coll. Junle YE)
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
Jianshi, Hubei, China, Jun, 2005 (coll. Junle YE). 45.5 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer qiannanicus Deuve & Tian, 2000
Male adult
35.5mm; Xujiawan, Zhuzang Town, Weng'an, Guizhou, China , alt. 1000-1200m, Apr, 2026, leg. local collector (coll. Junle YE)
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer xianfengensis Deuve & Li, 2009
Male adult
Banqiao Town, Enshi, Hubei, China, May, 2026 alt. 1300m, leg. local collector (coll. Junle YE). 40 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) augustus tenellus Rapuzzi, 2026
Male adult
35mm. Nanshan Pasture, Chengbu, Hunan, China, Jun, 2025, leg. local collector (coll. Junle YE)
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer adriaenssensi Lassalle & Prunier, 1993
Male adult
Nanshan Pasture, Chengbu, Hunan, China, Jun, 2025, leg. local collector (coll. Junle YE). 38.5 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) augustus tenellus Rapuzzi, 2026
Male adult
Mt. Huangzhushan, Wanfoshan Town, Tongdao, Hunan, China, alt. 1700m, Jun, 2026, leg. local collector (coll. Junle YE). 42 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) augustus tenellus Rapuzzi, 2026
Male adult
Nanshan Pasture, Chengbu, Hunan, China, alt. 1600m, May, 2026 (coll. Junle YE). 38 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) augustus tenellus Rapuzzi, 2026
Male adult
Nanshan Pasture, Chengbu, Hunan, China, alt. 1600m, May, 2026 (coll. Junle YE). 39 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer szechwanensis Hauser, 1931
Male adult
Taiping Village, Puchang Town, Suiyang, Guizhou, alt. 900-1100m, May, 2025, leg. local collector (coll. Junle YE). 40.5 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Female adult
Jianshi, Hubei, China, Jul, 2008, alt. 2000m (coll. Junle YE). 46 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer qiannanicus Deuve & Tian, 2000
Male adult
Huoshipo, Zhuzang Town, Weng'an, Guizhou, China , alt. 1000m, Apr, 2026, leg. local collector (coll. Junle YE). 38mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer wenshuicus Font, 1998
Male adult
Shanbao Village, Jiuba Town, Tongzi, Guizhou, alt. 1200-1400m, Apr-May, 2026 leg. local collector (coll. Junle YE). 43 mm
Copyright © Lokin Yeh

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
Shazi Town, Shizhu, Chongqing, alt. 1600m, Sep, 2025, leg. local collector (coll. Junle YE). 40.5 mm
Copyright © Lokin Yeh