Apotomus rufithorax Pecchioli, 1837
Male adult

Copyright © Kirill Makarov

Brachinus (Cnecostolus) bayardi Dejean, 1831
Female adult
E Turkmenistan, Charjev env., V.1996, V.Perepel leg.
12 mm
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sclopeta (Fabricius, 1792)
Male adult
S Bosnia, Mostar; 6mm
Copyright © Aleš Sedláček

Brachinus (Brachynolomus) nigricornis Gebler, 1829
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinus) elegans Chaudoir, 1842
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Cnecostolus) bipustulatus Quensel In Schoenherr, 1806
Male adult
Greece, Etolia Akarnania, Neohori, 1.5.1999. F. Angelini leg.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sichemita Reiche & Saulcy, 1855
Male adult
Greece, Ahaia, Kalavryta, 28-29.4.1999, leg. F. Angelini
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) alexandri F. Battoni, 1984
Male adult
SE Azerbajan, Talysh, Mistan v., 7-10.10.1993. det. A. Anichtchenko
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) explodens Duftschmid, 1812
Female adult
Bulgaria Chernomorets, 24.11.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Female adult
W Ukraine, Lviv city, Sykhiv (Zubra) forest, 49.791889, 24.047417, 13.VII.2024, Panin R. leg.
Copyright © Ruslan Panin

Broscus nobilis Dejean, 1828: 432
Male adult
Greece, Isl. Rhodos, 10 km NNW Lindos 10-20m, 18.4.1994 A Korell leg.
17 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Male adult
Italy, Emilia reg., Modena, Castelnuovo Rangone, 31.XII.1991, Sola C. leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni wiedemanni Menetries, 1836
Male adult
SE-Bulgaria:Burgas env. Tsarevo (Michurin), 10-15.VII.1981, J.Slaba leg.<br /> L=22 mm
Copyright © Ruslan Panin

Carabus (Procrustes) banoni Dejean, 1829
Male adult
Greece,W Crete Island, Mili gorge ,12.V.2009, Manfred Egger leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Procrustes) banoni Dejean, 1829
Male adult
Crete, Agia Irini (Sélinos); 31mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Female adult
SW Romania, Banat; 15mm
Copyright © Aleš Sedláček

Calosoma (Calosoma) sycophanta Linne, 1758
Female adult
Turkey : Antalya, Yukarikaraman 3.iv.2000 - M. Grotz Leg. (Size : 33 mm) Coll. N. Mal
Copyright © Noël Mal

Carabus (Pachystus) hortensis preslii Dejean, 1830
Male adult
S-Greece:, N Peloponnes, Erymanthos Reg., Kalentzi, 26.VI.2007, Manfred Egger leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni (as mussardianus Breuning, 1965)
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Male adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 37 mm
Copyright © Ruslan Panin

Calomera littoralis nemoralis Olivier, 1790
Male adult
UKRAINE,
Kherson Reg.,Genitchesk Distr., near Tehongar st., Kujuktuk isl., on saline 8-9.V.1995 Mishustin R. leg.
16 mm.
Copyright © Ruslan Panin

Calomera aulica aulica s.str.
Male adult
Oman, Dhofar, Jabal al Qamar, 5 km NE Dhalqut, 16°43'22.48’’N 53°16’27.26’’E, 22.9.2011, Richard Ambrus lgt.
11 mm
Copyright © Alexander Anichtchenko

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
CZ, C Moravia, Přerov; 9mm
Copyright © Aleš Sedláček

Calomera concolor concolor Dejean, 1822
Female adult
14,0 mm; Greece, Crete, Nom. Hanion. Georgioupolis, 4 km E, Kavros, leg. J. Scheuern, 4.-12.X.1999; det. J.Schoenfeld, 2000; Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Megacephala (Grammognatha) euphratica euphratica
Unsexed
Spain : Alicante, Salina la Mata, 15.vii.1979 - E. Vives leg. (Size : 24 mm) Coll. N. Mal
Copyright © Noël Mal

Cylindera (Eugrapha) trisignata hellenica (Cassola, 1973)
Male adult
SW Ukraine, Odessa region, Tatarbunary distr., Lebedivka vill., sea beach, 29.VII-6.VIII.2021, Yu. Kanarsky leg. /det. R.Panin, L=10.5 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg., L=12 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Male adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg.
Copyright © Ruslan Panin

Polistichus fasciolatus (P. Rossi, 1790)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr., near Bogatoe Ushchele vill., 04.VIII.2009, Shehovcov A. leg.
L=8mm
photo Panin R.
Copyright © Ruslan Panin

Zuphium olens (P.Rossi, 1790)
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 27-30.VI.2006 K.Makarov leg.
L=8.7 mm
Copyright © Kirill Makarov

Drypta (Drypta) dentata (P.Rossi, 1790)
Male adult

Copyright © Kirill Makarov

Parazuphium (Parazuphium) damascenum (Fairmaire, 1897)
Male adult
Morocco, Tagounite vill area 28.6.2010, Anichtchenko A. leg. & det. (aedeagus studied)
Copyright © Alexander Anichtchenko

Parazuphium (Parazuphium) chevrolatii chevrolatii Laporte, 1833a: 254
Male adult
Iran Gilan Elburz 1000m 5.viii.1968 W.Heinz leg. at light. J. Muilwijk det.
Copyright © Jan Muilwijk

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Acupalpus (Acupalpus) parvulus (Sturm, 1825)
Male adult

Copyright © Kirill Makarov

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Pachycarus (Mystropterus) cyaneus (Dejean, 1830)
Male adult
Grèce, Mycène, 27/05/1992, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Dixus obscurus Dejean 1825
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carterus (Pristocarterus) angustus (Menetries, 1832)
Male adult
Iran, Lorestan prov., near Dorud, 5 km SW Chabdar vill. 33.501061°N 49.003441°E 14-15.V.2017 leg. D.Kasatkin L=8.5 mm
Copyright © Kirill Makarov

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Cryptophonus) melancholicus Dejean, 1829
Male adult
Italy, Patro di Moncalvo (Asti).
Copyright © Gianni Allegro

Dicheirotrichus (Pelagophilus) punicus Bedel, 1899
Female adult
Espagne, Guardias Viejas, Almeria 04-05-1991
Copyright © Pascal Deschamps

Demetrias (Demetrias) atricapillus Linne, 1758: 416
Male adult
length: 4,5-6 mm
Holland, Beek Gld.
leg.det. W.Veldkamp, 1971
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Philorhizus melanocephalus (Dejean, 1825)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Platytarus faminii (Dejean, 1826: 447)
Male adult
Iran, Fars, 20 km W Estahban, 2500m
9-10.V.2007 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Trichis maculata Klug, 1832
Male adult
Ю Таджикистан, нижн. теч. р.Вахш, заповедник Тигровая Балка 26.V.1988 leg. Т.Компанцева
Copyright © Kirill Makarov

Cymindis (Cymindis) angularis Gyllenhal, 1810: 173
Male adult
L=7,5 mm
RUSSIA, Moscow reg., Lukhovitskiy distr., Beloomut settl. env, mixed forest, under bark. 21.IV.2007
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Cymindis (Menas) miliaris (Fabricius, 1801)
Female adult
SE Slovakia, Slovenský kras; 10,5mm
Copyright © Aleš Sedláček

Microlestes luctuosus Holdhaus, 1904b: 330
Male adult
Crymea, N slopu Baba-Dag Plateau, Taban-Dere, h~500 44°35'38"N 33°47'42"E 8.V.2017 leg. K. et O. Makarovs L=2,5 mm
Copyright © Kirill Makarov

Paradromius (Paradromius) suturalis (Motschulsky, 1844)
Female adult

Copyright © Igor Solodovnikov

Paradromius (Manodromius) linearis (Olivier, 1795)
Male adult

Copyright © Kirill Makarov

Metablus paracenthesis (Motschulsky, 1839)
Female adult
Azerbajan, Shirvan NP, 39.40N, 49.11E, 23.6.2024 Lason A. leg. (cAA). det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Licinus (Licinus) cassideus Fabricius, 1792: 148 (Carabus)
Female adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr.,near Bogatoe Ushchele vill.04.VIII.2009, Shehovcov A. leg.
female, 17mm
photo & det. Panin R.
Copyright © Ruslan Panin

Licinus (Licinus) silphoides (P. Rossi, 1790)
Female adult
Corse, Col de Bavella, 14.8.1966
Copyright © Alexander Anichtchenko

Notiophilus rufipes Curtis, 1829: pl. 254
Female adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Nebria (Eunebria) jockischi jockischii Sturm, 1815
Male adult
SE France, Alpes-Maritimes, Mt. Col de Turini h=1500m, Graben SW, 18.VIII.2005, Manfred Egger leg.
L= 13 mm

Copyright © Ruslan Panin

Nebria (Tyrrhenia) holtzi K. Daniel, 1903a: 162
Female adult
Greece, N-Peloponnese, Kalentzi vill., Mt. Enimanthos, 11.VI.2006, Manfred Egger leg.
L=13 mm
Copyright © Ruslan Panin

Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843
Male adult
SW Bosnia, Bijakoviči - Križevac Mt.; 8mm
Copyright © Aleš Sedláček

Nebria (Nebria) kratteri pindica Jeanne, 1974
Male adult
C Greece, Thessalia, Kastraki; 10mm
Copyright © Aleš Sedláček

Leistus (Pogonophorus) rufomarginatus (Duftschmid, 1812)
Male adult

Copyright © Kirill Makarov

Paussus (Cochliopaussus) turcicus Frivaldszky von Frivald, 1835
Male adult

Copyright © Kirill Makarov

Paussus (Cochliopaussus) turcicus Frivaldszky von Frivald, 1835: 263
Male adult
Iran, Hamadan env., Ali Sadr 14.5.1999, leg. Orszulik
Copyright © Alexander Anichtchenko

Agonum (Olisares) lugens (Duftschmid, 1812)
Male adult
Volgograd Reg., Pallasovka Distr., N shore of Elton Lake, right bank of Khara River 3.V.2009 leg.K.Makarov
8.8 mm
Copyright © Kirill Makarov

Omphreus (Paromphreus) krueperi Reitter, 1885
Female adult
S Greece, Taygetos Mts.; 28mm
Copyright © Aleš Sedláček

Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763)
Male adult

Copyright © Kirill Makarov

Atranus ruficollis Gautier des Cottes, 1858b
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Agonum (Agonum) antennarium (Duftschmid, 1812)
Male adult
W Ukraine, NE Carpathians, Chornogora Mt Rng, h=1650 m, source of the creek, 6.VI.2011, R. Panin leg. L=8.5 mm
Copyright © Ruslan Panin

Agonum (Agonum) menetriesii Faldermann, 1839
Male adult
S Kazakhstan, 15 km SE Kyzyl Orda, 30.4.2012, A. Ivanov leg.; det. K. Makarov, 2018
Copyright © Alexander Anichtchenko

Olisthopus fuscatus Dejean, 1828
Male adult
Turkey, Kirish distr., Camyuva riv., 28.9.2014, I. Enuschenko leg. det. Anichtchenko A. 2018 (aedeagus studied)
Copyright © Alexander Anichtchenko

Agonum (Olisares) hypocrita (Apfelbeck, 1904)
Male adult
NW-Ukraine, Polesia, Volhynia, Lyubeshiv Distr., Buchyn vill, swamped pinewood, «Selysko», 14.IX-17.X.2017, Kanarsky Yu. leg./det.Panin R.
Copyright © Ruslan Panin

Omphreus (Elladomphreus) eggeri Monzini & Sciaky, 2025 Paratype
Male adult
GR. Peloponnes / Panachaikogeb. / Kalanistra V.2014, leg. Manfred Egger
Copyright © Riccardo Sciaky

Nomius pygmaeus Dejean, 1831A: 512
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Dolichus halensis Schaller, 1783: 317
Male adult
S Ukraine, Krym Peninsula,
Simpheropil, 15-30.VIII.2009, Shaporinskiy V. leg.
male, 19 mm
Copyright © Ruslan Panin

Zabrus (Pelor) hiekei Anichtchenko & Gueorguiev, 2009 Paratype
Male adult

14 mm

Copyright © Alexander Anichtchenko

Myas (Myas) chalybeus Palliardi, 1825
Male adult
C. Bulgaria, Stara Zagora area, Stara Planina Mts., Shipka Mt. 17-VIII-2005 A. Safronov leg.
Copyright ©

Sphodrus leucophthalmus Linne, 1758
Female adult
W Austria, Hall in Tirol, 11.IV.1952, Richler leg.
L=26 mm

Copyright © Ruslan Panin

Amara (Amara) saphyrea Dejean, 1828: 463
Female adult
S Slovakia, Štúrovo; 9mm
Copyright © Aleš Sedláček

Calathus (Neocalathus) ambiguus ambiguus Paykull, 1790: 130
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) diligens (Sturm, 1824)
Male adult
Russia, Moskow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Argutor) leonisi Apfelbeck, 1904: 257
Female adult

Copyright © Igor Solodovnikov

Dodecanopterus laticornis rhodius (Straneo, 1935) Paratype
Male adult
Rhodos
Copyright © Vittorino Monzini

Caecomolopsis genesti Giachino, Allegro & Vailati, 2024 Holotype
Male adult
Grecia, nom. Fokída, O. Gióna m 1720, vers. N sopra Kaloskopí, 19.V.1989, Giachino legit (CGi) [38°39’24’’N 22°18’36’’E]
Copyright © Gianni Allegro

Clinidium (Arctoclinidium) canaliculatum O.G.Costa, 1839d
Male adult
photo from www.zin.ru
Copyright © Kirill Makarov

Scarites (Parallelomorphus) laevigatus Fabricius, 1792
Male adult
SW Ukraine, Odessa region, Tatarbunarsk distr., near Primorskoe vill.,in the sand, 8.VII.2004,
A Gontarenko leg.
L=16 mm

Copyright © Ruslan Panin

Clivina (Clivina) fossor fossor (Linnaeus, 1758)
Male adult
L=5,3 mm
RUSSIA, Moscow city, SW distr., Jasenevo, Bitcevskiy les, under tree. 19.V.2000
leg., det., photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Dyschirius (Dyschiriodes) cylindricus (Dejean, 1825)
Male adult

Copyright © Kirill Makarov

Clivina (Clivina) ypsilon Dejean, 1830
Male adult
NE Greece, Nei Pori; 6 mm
Copyright © Aleš Sedláček

Distichus (Distichus) planus (Bonelli, 1813)
Male adult
Turkey, Side, Colakli, leg. J.Scheuern, 24.IV.2000; det. Balkenohl, 2004. Photo: U.Schmidt, 2014. 12,8 mm
Copyright © Udo Schmidt

Dyschiriodes (Eudyschirius) globosus Herbst, 1784: 142
Male adult

Copyright © Kirill Makarov

Scarites (Scallophorites) buparius (Forster, 1771)
Male adult
Italie, Sicile, Selinunte, 04-07-1983
Copyright © Pascal Deschamps

Clivina (Leucocara) laevifrons Chaudoir, 1842
Male adult
Italy
Copyright © Alexander Anichtchenko

Dyschiriodes (Eudyschirius) rufipes (Dejean, 1825)
Female adult
W Ukraina, Podolian Upland, Opillia, Ivano-Frankivsk reg., Kasova Gora near Burshtyn vill., steppe site, 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=2,9mm
Copyright © Denis Khrapov

Dyschiriodes (Dyschiriodes) agnatus Motschulsky, 1844: 79
Male adult
W Ukraine, NE Carpathians, Lviv reg., Stryi distr., near Rozgirche vill., bank of Stryi river, 18.VI.2021, D.&O. Khrapov leg./det. R. Panin L=5mm
Copyright © Denis Khrapov

Siagona europaea Dejean, 1826: 468
Male adult
Uzbekistan, Kashkadarjnsk Reg., env. Mubarek 5-8.VI.2004.
L=10.3 mm
Copyright © Kirill Makarov

Bembidion (Testedium) bipunctatum (Linne, 1761)
Male adult
length: 3,6-4,8 mm
Austria, Schladminger Tauern, Mariapfarr
leg.det. U.Schmidt, 1984
Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) biguttatum (Fabricius, 1779)
Male adult

Copyright © Kirill Makarov

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult
4,4 mm
Namibia, Etosha National Park, Okaukuejo, 1400 mб leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012
Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Bembidion (Peryphanes) dalmatinum dalmatinum Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult

Copyright © Kirill Makarov

Bembidion (Euperyphus) combustum combustum Menetries, 1832: 138
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=7.0 mm
Copyright © Kirill Makarov

Bembidion (Peryphanes) cf brunnicorne Dejean, 1831
Male adult
W.Ukraine, Transcarpathian, Polonynian Beskids, Yavirnyk Rng, h=600 m, near Ruskyi Mochar vill.,15.V.2021, V. Khrapova leg./R. Panin det., L=4.7 mm
Copyright © Denis Khrapov

Odontium (Odontium) striatum (Fabricius, 1792)
Female adult
Latvia, Dunava
Copyright © Alexander Anichtchenko

Pogonistes liliputanus (Apfelbeck, 1904)
Male adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 3.8 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko