Daptus vittatus Fischer von Waldheim, 1823: pl.46
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Harpalus (Loboharpalus) rubefactus Bates, 1873
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Harpalus (Harpalus) tarsalis (as hokkaidensis) Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) tarsalis Mannenheim, 1825: 28 Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) tarsalis Mannenheim, 1825: 28 Holotype
Male adult
Onneyu Spa, HOKKAIDO, 24. VI. 1958, K. Baba. COL-134
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes (as ainus) Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Harpalus) laevipes Zetterstedt, 1828: 26 Holotype
Male adult
Sounkyo-Spa, HOKKAIDO, 25. VI. 1958, K. Baba. COL-133
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Male adult
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Male adult
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Endophallus
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Male adult
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Male adult
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Endophallus
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Male adult
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Endophallus
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) tarsalis (as hokkaidensis Habu et Baba, 1963)
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39250
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883: 239
Male adult
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883: 239
Endophallus
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Male adult
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Endophallus
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) chalcentus Bates, 1873a: 263
Male adult
Collection ground Minobu cho, Yamanashi Pref. 35617
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) chalcentus Bates, 1873a: 263
Endophallus
Collection ground Minobu cho, Yamanashi Pref. 35617
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Female adult
Russia
Copyright © Kirill Makarov

Harpalus (Loboharpalus) platynotus Bates, 1873
Male adult
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Loboharpalus) platynotus Bates, 1873
Endophallus
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Male adult
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Endophallus
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883
Male adult
SE Maritime Prov., Lazovsky dist., Lazo env., 17-20.VI.2010, leg. М. Smirnov Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) major (Motschulsky, 1850)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) ussuricus Mlynar, 1979: 97
Male adult
Ю Приморье, Хасанский р-н, 7.5 км Ю п.Барабаш, устье руч. Олений 43°07'01"N 131°29'15"E 27.V.2015 leg. К.Макаров L=8.7 mm
Copyright © Kirill Makarov

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) modestus Dejean, 1829
Female adult
Russia, 13 km from Irkutsk by Aleksandrovskii road, 20.8.1992, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Caloharpalus) chalcentus Bates, 1873
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Nephoharpalus) pallidipennis A. Morawitz, 1862
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Male adult

Copyright © Kirill Makarov

Oxycentrus (Oxycentrus) argutoroides (Bates, 1873)
Male adult
Collection ground Tsushima, Nagasaki pref. 35695<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright ©

Platymetopus flavilabris (Fabricius, 1798)
Male adult
Collection ground Higashisonogi cho, NagasakiPref. 35330
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Platymetopus flavilabris laticeps Dejean, 1829
Male adult
Japan. Image from Kataev, 2023
Copyright © - - -

Platymetopus flavilabris laticeps Dejean, 1829
Male adult
Philippines. Image from Kataev, 2023
Copyright © - - -

Trichotichnus (Trichotichnus) coruscus (as sachalinensis) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Male adult
Collection ground Ashikaga city,Tochigi Pref.Ashikaga～shi 36079
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Collection ground Ashikaga city,Tochigi Pref.Ashikaga～shi 36079
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) noctuabundus Habu, 1954b: 56
Male adult
Collection ground Siota cho, Saga pref, 34869
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) noctuabundus Habu, 1954b: 56
Male adult
Collection ground Siota cho, Saga pref, 34869
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) lucidus (A. Morawitz, 1863)
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) lucidus (A. Morawitz, 1863)
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Male adult
Daegu-si, Republic of Korea. Scale bar: 3 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Scale bar: 0.6 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Male adult
Daegu-si, Republic of Korea. Scale bar: 2 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Trichotichnus (Bottchrus) nanus Habu, 1954c: 233
Male adult
Daegu-si, Republic of Korea. Scale bar: 2 mm.
Copyright © Dooyoung Kim

Trichotichnus (Bottchrus) nanus Habu, 1954c: 233
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) noctuabundus Habu, 1954
Male adult
Vietnam. Image from Kataev et al, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Habitat
Under the leaf litter, Daegu-si, Republic of Korea. (10.III.2021)
Copyright © Dooyoung Kim