Brachinus (Brachynolomus) stenoderus Bates, 1873
Endophallus
Collection ground Hobetsu cho, Hokkaido Pref. 33485
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pheropsophus (Stenaptinus) javanus (Dejean, 1825)
Endophallus

Copyright © Dmitri Fedorenko

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Endophallus
NE-China:Liaoning prov.,Fengcheng City, Miaoyang,
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Endophallus
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan,
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Endophallus
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
Copyright © Ruslan Panin

Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
Endophallus

Copyright © Ruslan Panin

Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Endophallus
Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
Copyright © Igor Solodovnikov

Galerita (Galerita) orientalis Schmidt-Goebel, 1846
Endophallus
Higashihagio, Omuta City, Fukuoka pref., Japan, 21 may 2016, S. Nishimoto leg.
Copyright © Alexander Anichtchenko

Stenolophus (Stenolophus) propinquus A.Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Endophallus
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Endophallus
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883: 239
Endophallus
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Endophallus
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) chalcentus Bates, 1873a: 263
Endophallus
Collection ground Minobu cho, Yamanashi Pref. 35617
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Endophallus
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) punctatipennis A.Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Endophallus
Collection ground Mukawa cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Acupalpus (Palcuapus) inornatus Bates, 1873
Endophallus
Collection ground Kamogawa city, ChibaPref. 34922
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Endophallus
Primorye Territory, Khasan Distr., vicinity of Slavyanka Vill., Cape Busse, 42°52'42"N 131°27'58"E, 13.VI.2023, I. Melnik leg. L=11.0mm
Copyright © Kirill Makarov

Anisodactylus (Pseudanisodactylus) punctatipennis A.Morawitz, 1862
Endophallus
Primorye Territory, Khasan Distr., 12.5 km WNW of Barabash Vill., Ovchinnikovo, 43°13'56"N 131°21'23"E, h~110 m, 10–13.VIII.2020, K. Makarov, A. Matalin leg. L=11.1 mm
Copyright © Kirill Makarov

Cymindis (Menas) daimio Bates, 1873
Endophallus
Collection ground Tomakomai city, Hokkaido Pref. 37172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Lebia (Poecilothais) bifenestrata A. Morawitz, 1862a: 245
Endophallus
Collection ground Misugi machi, Mie pref. 34126
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Diplocheila (Isorembus) zeelandica L. Redtenbacher, 1867: 5
Endophallus
Collection ground Shimamaki mura, Hokkaido Pref. 33049
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Panagaeus (Panagaeus) japonicus Chaudoir, 1861c: 356
Endophallus
Collection ground Kaminokuni cho, Hokkaido Pref. 35988
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Metacolpodes buchanani Hope, 1831: 21
Endophallus
Russia, Evreiskaya AO., Birobidjan 31.07.2013
Copyright © Pavel Budilov

Amara (Bradytus) sinuaticollis A. Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Bradytus) simplicidens A. Morawitz, 1863: 60
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Bradytus) majuscula (Chaudoir, 1850)
Endophallus
Collection ground Monbetsu cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) macronota Solsky, 1875
Endophallus
Collection ground Tomakomai city, Hokkaido Pref. 37172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) hiogoensis (Bates, 1873)
Endophallus
Collection ground Tsushima, NagasakiPref. 35695
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) gigantea (Motschulsky, 1844)
Endophallus
Collection ground Kagoshima city, Kagoshima Pref. 38979
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) obscuripes Bates, 1873a: 294
Endophallus
Collection ground Shibecha cho, Hokkaido Pref. 38493
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) familiaris (Duftschmid, 1812)
Endophallus
Collection ground Kiyosato cho, Hokkaido Pref. pref. 38785
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) congrua A. Morawitz, 1862b: 326
Endophallus
Collection ground Ise city, Mie Pref. 37125
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) chalcites Dejean, 1828: 462
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Eosteropus) japonicus (Motschulsky, 1860)
Endophallus
Collection ground Tomakomai, Hokkaido Pref. 30926
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Endophallus
Collection ground Rebun Is. Hokkaido Pref. 27191
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) communis (Panzer, 1797)
Endophallus
det. K. Makarov
Copyright © Kirill Makarov

Amara (Curtonotus) gigantea (Motschulsky, 1844)
Endophallus
Russia, Primorskii reg.
Copyright © Alexander Anichtchenko

Amara (Bradytus) simplicidens A. Morawitz, 1863: 60
Endophallus
Kunashir Isl., hills S of Stolbchatyi Cape, 44°00'20"N, 145°42'05"E 23.VIII.2003 leg. I. Melnik
Copyright © Kirill Makarov

Amara (Bradytus) pallidula (Motschulsky, 1844)
Endophallus
Primorskii Terr., Khasanskii Distr., southern outskirts of Barabash Vill., 43°10'52"N 131°29'21"E 23.V.2021 leg. K. Makarov
Copyright © Kirill Makarov

Amara (Bradytus) mikae Lafer, 1980
Endophallus
Primorskii Terr., Khasanskii Distr., 4 km NE of Andreevka Vill., Andreevskaya Pad’, 42°39'41"N 131°10'59"E, 26.V.2022 leg. K. Makarov
Copyright © Kirill Makarov

Amara (Bradytus) majuscula (Chaudoir, 1850)
Endophallus
Buryatia, Tunkinskii Distr., environs of Tunka Vill. 3.VIII.2012 leg. A. Prosvirov
Copyright © Kirill Makarov

Amara (Bradytus) fritzhiekei Sundukov, 2013
Endophallus
Primorskii Terr., Lazovskii Distr., Prosyolochnaya River, floodland, 1–5.IX.2001 leg. A. Berzan
Copyright © Kirill Makarov

Amara (Bradytus) fritzhiekei Sundukov, 2013
Endophallus
Primorskii Terr., Terneiskii Distr., environs of Lake Golubichnoe, sediments on seacost, 18.IX.2018 leg. M. Sergeev
Copyright © Kirill Makarov

Amara (Bradytus) apricaria (Paykull, 1790)
Endophallus
Kyrgyzstan, Moldy-Too Mt. Range, Kurtka River, h ~ 2200 m, 20–21.VII.1998 leg. A. Klimenko
Copyright © Kirill Makarov