Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy, Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.; female, 23 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola roccae Born, 1923
Male adult
NW-Italy, Liguria: Savona env., Monte San Giorgio, 30.X.1977, Monauzzi leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) alysidotus Illiger, 1798
Female adult
Italy, Reggio Emilia, Albinea env. h=150 m, VI.2018, A. Cartini leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
Female adult
SE France, Alpes-Maritimes, Col de Braus h=1000 m, 23.V-18.VI.2014, P. Cavazzuti & G. Proscia leg., L= 24 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Female adult
France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Male adult
NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W Ukraine,
Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
male, 21mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
long. 22mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
Female adult
CZ, C Moravia, Dřevohostice - Líšná, 13mm
Copyright © Aleš Sedláček

Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
Female adult
W-Ireland, County Mayo, Letterneevoge (R312),16-30.VI.2013, J.Stypula leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Female adult
NW Ukraine, Polesia, Rovno reg., Rokytne distr., near Bil’sk vill., 4-6.VII.2015, Panin R. leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Carabus) arcensis arcensis Herbst, 1784
Female adult
Lower Austria, Baden, Klausen-Leopoldsdorf, 15.XII.2024, M. Egger leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
Female adult
W- FRANCE,
Bretagne Reg., Finistrere, Foret de Coatloc'h, XI.2006
female, 26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Female adult
NW-FRANCE,
Basse-Normandie Reg., Orne, Forêt de Bellême, 14.XII.2008, Pascal Stefani leg.
female, 26mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Male+Female adult
NW-FRANCE,
Basse-Normandie Reg,, Orne, Foret d' Ecouves, V.2009, Pascal Stefani leg.
male, 25 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826 (as barthei Lapouge, 1912)
Male+Female adult
SC -France,
Midi-Pyrйnйes reg., Ariиge, Foret de Belesta, XI.2008, J.Nicolas leg.
L=31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
Female adult
NW Italy,
Altare, Savona,15.VI.1995, Cartini A.leg.
female, 33mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) splendens Olivier, 1790
Male+Female adult
SE France,
Midi-Pyrйnйes reg., Ariиge, Vaychis, 22.X.2002, A.Laforgue leg.
female, 30mm.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Female adult
SE France, Alpes-Maritimes, Ascros, VI.2011
L=30 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Male adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L= 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri clairi Gehin, 1885 (as devillei Tarrier, 1972)
Male adult
SE-France, Alpes-Maritimes, Beuil env., h=1550m, 14.VII.1986
Copyright © Ruslan Panin

Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin

Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult

Copyright © Kirill Makarov

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Female adult
SE-France, Var, near Le Muy, VIII.1980, Michon leg.
L=31 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
Female adult
SE-France, Var, Fréjus env., 26.II.1997, <br /> L=31 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fiorii Born, 1901
Male+Female adult
NW-Italy,
Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
female, 30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
Male+Female adult
S France,
Midi-Pyrénées Region, Ariège, Col du Chioula h=1500m, V-VI.2005, Pascal Stefani leg.
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
Male adult
W-FRANCE, Brittany , Finistère, Botvarec Forest, 19.XI.2006, Pascal Stefani leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus mixtus (as neomixtus Tarrier, 1975)
Male adult
L=29mm SE-France, Drôme, Foret de Saou, VII.2014, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
Male+Female adult
E-Netherlands, Gelderland, Winterswijk Woold env.,15.II.2015, H.de Haan leg.
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
SW-France,
Foret d'Iraty Pyrenees-Atlantiques, d'Occabe, 15.VIII.1988
male,30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
Male adult
SE–France, Var departament, Foret royale -Saint-Paul-en-Foret vill.,18.VI.2002, Thelot J.P. leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
W-Ireland, County Mayo, Clare Isl., 17-28.VI.2002, S. McCormack leg.<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Female adult
N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg.<br /> L=27mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Male adult
Luxembourg, Bollendorf-Pont env., 3.III.1999 /det.R Panin
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Male adult
NE Spain, Catalonia, Girona Prov., San Hilario Sacalm env., 20.II.1988, A. Mollard leg.<br /> L=25 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
Female adult
W-Hungary, NE-Alps, Mts. Kőszeg, Velem env., Borha-forrás, 26.VII.1981, Gy. Somorijai leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
NC-Spain, Basque Co, Álava, Sierra de Gorbea, VI.2014, L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Male+Female adult
NW France,
Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
female, 28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult

France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov

Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
Female adult
Czech, WS Bohemia,
Plzen Reg., Domazlice Distr., Babylon, h=500m, 30.VIII.2008, S.I. Budka leg.
L=24mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
Male adult
N Italy, Alpi Graie; 20 mm
Copyright © Aleš Sedláček

Carabus (Orinocarabus) fairmairei fenestrellanus Beuthin, 1899
Male adult
NW-Italy, Piemonte prov., Cottian Alps, Mt Albergian h=2450 m.<br /> L=20 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
Male adult
SE France, Alpes-Maritimes, Isola env., Colle Santa-Anna h=2309 m, 44*13',127" N, 07*06',242"E, 31-7.VIII..2012. L=21 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) sylvestris sylvestris (as micklitzi Sokolar, 1910)
Endophallus
W Austria, Styria, Mt Hochwechsel W slope, 47.531667, 15.906000 h=1640 m,
Copyright © Ruslan Panin

Carabus (Orinocarabus) sylvestris sylvestris (as micklitzi Sokolar, 1910)
Male adult
W Austria, Styria, Mt Hochwechsel W slope, 47.531667, 15.906000 h=1640 m, 14.IV.2025, I. Uspenskyi leg.
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Male adult

Copyright © Kirill Makarov

Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
Male adult
NE-Italy,
Friuli Reg., Udine, Carnia, Pierabech h~1100m, I.Rapuzzi leg.
male, 23 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
SE-France: Alpi Marittime, Monte Viso, 14.VII.2010
L=23 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis irregularis (as jurassicus Born, 1911)
Male+Female adult
France, Ain (01), Jura Mts, Grand Colombier, Forêt d' Arvières h~1250 m, 25.X.2014, Ya. Vasseur leg., L=23 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
Copyright © Noël Mal

Carabus (Tachypus) auratus brunieri Barthe, 1921
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg., 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček

Carabus (Tachypus) vagans Olivier, 1795
Female adult
SE-France, Var: Le Muy env., 30.VII.2008., A. Paulian leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male+Female adult
France : Vaucluse, Mt. Ventoux-Mt. Serein, 1400-1800m, vii.1969-N. Mal Leg.-Coll.RBINS, N. Mal - including f.i. ventouxensis and nicolasi
Copyright © Noël Mal

Carabus (Tachypus) cancellatus carinatus (misellus Lapouge, 1898)
Male adult
SC-France, Massif central, Mt.Cantal, Plateau du Limon h=1300m, VIII.2013, Pascal Stefani leg., L=23 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus carinatus (as subcarinatus Kolbe, 1912)
Male adult
E-Netherlands, Gelderland prov., Nijmegen., Berg en Dal vill., 6.XII.2014, H.de Haan leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko