Parena monostigma (Bates, 1873)
Male adult
SE Primorskii reg., Lazo res., seashore, Ta-Chigouz bay,
26.06.04 leg. M. & L. Smirnov
photo from www.zin.ru
Copyright © Maksim Smirnov

Parena perforata (Bates, 1873)
Male adult
S Primorie, Lazovsky Reserve, kordon “Amerika” 43°16'16"N 134°03'01"E Window trap 14-16.VIII.2007 leg. K.Makarov, A.Zaitzev<br /> L=10.5
Copyright © Kirill Makarov

Parena tripunctata (Bates, 1873)
Male adult

Copyright © Kirill Makarov

Parena kurosai Habu, 1967 Holotype
Male adult
near Mt. Takao, TOKYO, 27. VII. 1962, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-179
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Parena kurosai Habu, 1967 Holotype
Male adult
near Mt. Takao, TOKYO, 27. VII. 1962, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-179
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Parena kurosai Habu, 1967 Holotype
Male adult
near Mt. Takao, TOKYO, 27. VII. 1962, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-179
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Parena kurosai Habu, 1967 Holotype
Male adult
near Mt. Takao, TOKYO, 27. VII. 1962, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-179
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Parena kurosai Habu, 1967 (Label) Holotype
Male adult
near Mt. Takao, TOKYO, 27. VII. 1962, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-179
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Parena nigrolineata nipponensis Habu, 1964 Holotype
Male adult
Fuchu C., TOKYO, 03. VII. 1958, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-177<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Parena nigrolineata nipponensis Habu, 1964 Holotype
Male adult
Fuchu C., TOKYO, 03. VII. 1958, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-177<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Parena nigrolineata nipponensis Habu, 1964 Holotype
Male adult
Fuchu C., TOKYO, 03. VII. 1958, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-177<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Parena wrasei Kirschenhofer, 2006 Holotype
Male adult
Japan (Hyogo), Asako gun, Okuno – cho, Tochi-hara, 29.VII.1990, T. Ito/Parena ? perforata Bates Ito det. (cDW)
Copyright © Schillhammer

Parena schillhammeri Kirschenhofer, 2006 Holotype
Male adult
Indonesia, C. Sulawesi, Kab. Donggalia, viilage Toro UTM 51 S (WGS-84) X 9833613, Y 170666. Alt 815 m/from: T.cacao under diverse, planted shade (canopy fogging) 24-XII-2003: MM. BOS leg. 05C241203E. (cDW)
Copyright © Schillhammer

Parena kataevi Kirschenhofer, 2006 Holotype
Female adult
China, W Hubei, Muyuping S. env. 31°45’N 110°45’E, 1300m, 20.–21.VI.2003. (cDW)
Copyright © Schillhammer

Parena yunnana Kirschenhofer, 1994
Male adult

Copyright © Schillhammer

Parena laesipennis (Bates, 1873)
Male adult
det. Hongliang Shi, 2023
Copyright © Schillhammer

Parena formosana Ohkura, 1978
Male adult

Copyright © Schillhammer

Parena perforata (Bates, 1873)
Male adult
det. Kirschenhofer
Copyright © Schillhammer

Parena laesipennis (Bates, 1873)
Male adult
Collection ground Nago city, OkinawaPref. 30065
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Parena latecincta (Bates, 1873)
Male adult
Collection ground Naha city, OkinawaPref. 29833
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Parena nigrolineata (Chaudoir, 1852)
Male adult
Collection ground Yokohama city, KanagawaPref. 34172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Parena tripunctata (Bates, 1873)
Female adult
Russia, SE Primorie
Приморский кр., Лазовский зап., ~10 км SW пос. Сокольчи, окр. кордона Корпадь 43°15' N, 134°7' E, 23-27.06.10, leg. М.Смирнов
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Parena dorsigera (Schaum, 1863)
Male adult
Tonkin
Copyright © Erich Kirschenhofer

Parena sellata (Heller, 1921)
Male adult
Philippines, E Luzon, Sierra Madre, Gabaldon, Nueva Ecija, 2.2018 (DUBC). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Parena nigrolineata (Chaudoir, 1852)
Male adult
India, Assam, Chapar, 26.267N 90.413E 30.12.2013, K. Tomkovich leg. (cAA) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Parena tesari (Jedlicka, 1951)
Male adult
Taiwan, Hualien, Hsinpaiyang, 18.7.2007. det. det. Hongliang Shi, 2023
Copyright © Erich Kirschenhofer

Parena cf circumdata Shibata, 1987
Male adult
Taiwan, Ilan, Fushan Park, 16.9.2007. det. J. Marrero, 2014
Copyright © Jose Marrero

Parena (Crossoglossa) cavipennis (Bates, 1873) Lectotype
Male adult

Copyright © Hongliang Shi

Parena rufotestacea Jedlička, 1934 Holotype
Female adult

Copyright © Hongliang Shi

Parena (Crossoglossa) cavipennis (Bates, 1873)
Male adult
Beijing
Copyright © Hongliang Shi

Parena (Crossoglossa) cavipennis (Bates, 1873)
Male genitalia
A Beijing, B Taiwan. Scale bar = 0.5 mm
Copyright © Hongliang Shi

Parena (Parena) monticola Shibata, 1987
Male adult
China: Sichuan Mt. Emei, 1050 m, 18.VII.1990, L.&M. Bocak lgt.
Copyright © Hongliang Shi

Parena (Parena) monticola Shibata, 1987
Male adult
Jiangxi, Wugongshan, 1300–1900 m, 2008.VII.15-16, Yang Zaihua lgt (IZAS)
Copyright © Hongliang Shi

Parena (Parena) monticola Shibata, 1987
Male genitalia
Jiangxi
Copyright © Hongliang Shi

Parena (Parena) circumdata Shibata, 1987
Female adult
India Darjeeling D., C.J. Rai", "Chairy Cow. 26.VIII.85 800 m
Copyright © Hongliang Shi

Parena (Parena) circumdata Shibata, 1987
Male adult
Taiwan, Hsinchu county, Wufeng, Dalu forest road, 2003.VI.15
Copyright © Hongliang Shi

Parena (Parena) circumdata Shibata, 1987
Male genitalia
Guangdong
Copyright © Hongliang Shi

Parena (Parena) latecincta (Bates, 1873)
Male adult
Filippine –Mindanao, Bukidnon, V.2013". 1 male (CRS), "Filippine Negros oc., Don Salvator, Benedicto II.2018
Copyright © Hongliang Shi

Parena (Parena) latecincta (Bates, 1873)
Female adult
Nepal Kathmandu V. M. Brancucci
Copyright © Hongliang Shi

Parena (Parena) latecincta (Bates, 1873)
Male adult
LAOS-NE, Xieng Khouang prov., 19°37-8'N 103°20-1'E, Phonsavan (30 kn NE): Phou Sane Mt., 1400-1700 m, 10.-30.v.2009, D. Hauck leg
Copyright © Hongliang Shi

Parena (Parena) latecincta (Bates, 1873)
Male adult
Beijing, Haidian, Baiwangshan, 2009.VI.20, Shi Hongliang, Yang Ganyan lgt
Copyright © Hongliang Shi

Crossoglossa viridilineata Jedlička, 1939 Holotype
Female adult

Copyright © Hongliang Shi

Parena (Parena) latecincta (Bates, 1873) Lectotype
Female adult

Copyright © Hongliang Shi

Parena (Parena) latecincta (Bates, 1873)
Male genitalia
A - Chongqing. B - Taiwan. Scale bar = 0.5 mm.
Copyright © Hongliang Shi

Parena (Parena) latecincta (Bates, 1873)
Male genitalia
A - Laos. B - India. C - Philippine. Scale bar = 0.5 mm
Copyright © Hongliang Shi

Parena (Parena) nigrolineata (Chaudoir, 1852)
Male adult
Taiwan
Copyright © Hongliang Shi

Parena (Parena) nigrolineata (Chaudoir, 1852)
Female adult
Guizhou, Chishui, Jinsha valley, 500 m, 2000.IX.22, Liang Hongbin lgt
Copyright © Hongliang Shi

Parena (Parena) nigrolineata (Chaudoir, 1852)
Male adult
Shanghai
Copyright © Hongliang Shi

Parena schillhammeri Kirschenhofer, 2006 Holotype
Male adult

Copyright © Hongliang Shi

Parena (Parena) nigrolineata (Chaudoir, 1852) Lectotype
Male adult

Copyright © Hongliang Shi

Parena (Parena) nigrolineata (Chaudoir, 1852)
Male genitalia
A Taiwan. B Shanghai (elytra without lateral strips). C Philippine. D Java. Scale bar = 0.5 mm
Copyright © Hongliang Shi

Parena (Parena) nigrolineata (Chaudoir, 1852)
Male genitalia
A India. B Holotype of Parena schillhammeri Kirschenhofer [= Parena nigrolineata (Chaudoir)]. Scale bar = 0.5 mm
Copyright © Hongliang Shi

Parena (Parena) fasciata (Chaudoir, 1872)
Female adult
Philippines, M. Halcon, Mindoro VII.2011
Copyright © Hongliang Shi

Parena (Parena) fasciata (Chaudoir, 1872)
Male adult
Philippines - E. Luzon, Isabela, Dindin, VIII.2014
Copyright © Hongliang Shi

Parena (Parena) fasciata (Chaudoir, 1872)
Male adult
PNG, Yanarba, Fgum Is. Meek. ii.95
Copyright © Hongliang Shi

Parena (Parena) fasciata (Chaudoir, 1872)
Male adult
Centr. Java, Nglirip., Mme E. Walsh.
Copyright © Hongliang Shi

Parena (Parena) fasciata var. unicolor Louwerens, 1949 Holotype
Male adult

Copyright © Hongliang Shi

Crossoglossa sellatoides Jedlička Holotype
Male adult

Copyright © Hongliang Shi

Phloeodromus hastatus Heller [= Parena fasciata (Chaudoir)] Holotype
Male adult

Copyright © Hongliang Shi

Phloeodromus sellatus Heller [= Parena fasciata (Chaudoir)] Holotype
Female adult

Copyright © Hongliang Shi

Parena (Parena) fasciata (Chaudoir, 1872) Holotype
Female adult

Copyright © Hongliang Shi

Parena (Parena) fasciata (Chaudoir, 1872)
Male genitalia
Philippine, the "hastata" form [h]. Scale bar = 0.5 mm.
Copyright © Hongliang Shi

Parena (Parena) fasciata (Chaudoir, 1872)
Male genitalia
A - Taiwan, the "sellata" form [s]. B - Tonkin, the rhombic form [r]. C - Java, the "unicolor" form [u]. D - Papua New Guinea, the melanic form [m]. E - Queensland, the "fasciata" form [f]. F - Philippine, the "sellata" form [s]. Scale bar = 0.5 mm.
Copyright © Hongliang Shi

Parena (Parena) rubripicta Andrewes, 1928
Female adult

Copyright © Hongliang Shi

Parena (Parena) rubripicta Andrewes, 1928: 17
Female adult
Yunnan
Copyright © Hongliang Shi

Parena (Parena) rubripicta Andrewes, 1928: 17
Female adult
Guangxi
Copyright © Hongliang Shi

Parena (Parena) rubripicta Andrewes, 1928 Holotype
Male adult

Copyright © Hongliang Shi

Parena (Parena) rubripicta Andrewes, 1928
Male genitalia
Xizang (Medog). Scale bar = 0.5 mm.
Copyright © Hongliang Shi

Parena (Bothynoptera) triguttata Shi & Liang, 2023 Paratype
Female adult
CHINA, Sichuan, Ebian / county, Heizhugou forestry / park, mixed forest / N29.03432 E102.98993", "1800 m, 2012.VI.17, / on vegetation. / HUANG Hao lgt", "IOZ(E)1700239", "PARATYPE / Parena (Bothynoptera) / triguttata sp. nov. / des. Shi H.L. 2022" [red label]
Copyright © Hongliang Shi

Parena (Bothynoptera) triguttata Shi & Liang, 2023 Holotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) triguttata Shi & Liang, 2023
Male genitalia

Copyright © Hongliang Shi

Parena (Bothynoptera) gonggaica Shi & Liang, 2023 Holotype
Male adult
Hailuogou, Luding, Sichuan / 27-VII-2007 / alt.1940 m Liu, Zhang, Zhou, & Bi lgt." [in Chinese], "HOLOTYPE / Parena (Bothynoptera) / gonggaica sp. nov. / des. Shi H.L. 2022" [red label]
Copyright © Hongliang Shi

Parena (Bothynoptera) gonggaica Shi & Liang, 2023
Male genitalia

Copyright © Hongliang Shi

Parena (Bothynoptera) emarginata Shi & Liang, 2023 Holotype
Female adult

Copyright © Hongliang Shi

Parena (Bothynoptera) emarginata Shi & Liang, 2023 Paratype
Female adult
Xizang, Cona county, Lai xiang / 2500 m, Yang Xiaodong leg. / 2013.VI.20" [in Chinese], "IOZ(E)1700238", "PARATYPE / Parena (Bothynoptera) / emarginata sp. nov. / des. Shi H.L. 2022" [red label]
Copyright © Hongliang Shi

Parena (Bothynoptera) quadrisignata Mateu, 1977: 164
Male adult
Yunnan
Copyright © Hongliang Shi

Parena phongsalyensis Kirschenhofer, 2011 Holotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) quadrisignata Mateu, 1977: 164 Holotype
Female adult

Copyright © Hongliang Shi

Parena (Bothynoptera) quadrisignata Mateu, 1977: 164
Male genitalia
A - Yunnan. B - Holotype of Parena phongsalyensis Kirschenhofer [= Parena quadrisignata Mateu]. Scale bar = 0.5 mm
Copyright © Hongliang Shi

Parena (Bothynoptera) malaisei (Andrewes, 1947)
Male adult
Yunnan
Copyright © Hongliang Shi

Parena (Bothynoptera) malaisei (Andrewes, 1947)
Male adult
Mure Num, 2000-1550 m, 4-7.VI.1983", "E-Nepal, Arun v., M. Brancucci
Copyright © Hongliang Shi

Parena (Bothynoptera) malaisei (Andrewes, 1947)
Male genitalia
A - Guangxi. B - Taiwan. C - Nepal. Scale bars = 0.5 mm
Copyright © Hongliang Shi

Parena (Bothynoptera) malaisei (Andrewes, 1947) Holotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) monostigma (Bates, 1873)
Male adult
Zhejiang
Copyright © Hongliang Shi

Parena koreana Kirschenhofer, 1994 Holotype
Female adult

Copyright © Hongliang Shi

Parena japoincia Jedlička, 1946 Holotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) monostigma (Bates, 1873) Lectotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) monostigma (Bates, 1873)
Male genitalia
A - Zhejiang. B - Japan. Scale bar = 0.5 mm.
Copyright © Hongliang Shi

Parena (Bothynoptera) shapingensis Xie & Yu, 1993b: 192
Male adult
Yunnan
Copyright © Hongliang Shi

Parena (Bothynoptera) shapingensis Xie & Yu, 1993b: 192 Holotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) shapingensis Xie & Yu, 1993b: 192 Holotype
Male genitalia

Copyright © Hongliang Shi

Parena (Bothynoptera) kurosai Habu, 1967
Male adult
India, Uttarakhand Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Parena (Bothynoptera) tripunctata (Bates, 1873)
Male adult
Shaanxi
Copyright © Hongliang Shi

Crossoglossa piceola Chaudoir [=Parena tripunctata (Bates)] Lectotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) tripunctata (Bates, 1873) Lectotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) tripunctata (Bates, 1873)
Male genitalia
A - Shaanxi. B - Lectotype of Parena tripunctata (Bates). C - Lectotype of Crossoglossa piceola Chaudoir [=Parena tripunctata (Bates)]. Scale bars = 0.5 mm
Copyright © Hongliang Shi

Parena (Bothynoptera) taiwana Hua, 2002
Male adult
Guangxi, Wuming, Damingshan mt., 518 m, 2014.VI.08, Lu Yanquan leg. On flower
Copyright © Hongliang Shi

Parena (Bothynoptera) taiwana Hua, 2002
Female adult
Taiwan, Nantou, Ren'ai, N. Dongyanshan, 1800 m, 2010.IV.22, Chou Wen-I leg.
Copyright © Hongliang Shi

Parena (Bothynoptera) taiwana Hua, 2002
Male adult
Taiwan, Nantou, Ren'ai, N. Dongyanshan, 1800 m, 2010.IV.22, Chou Wen-I leg.
Copyright © Hongliang Shi

Parena (Bothynoptera) taiwana Hua, 2002
Male genitalia
Taiwan
Copyright © Hongliang Shi

Parena (Bothynoptera) tesari (Jedlicka, 1951)
Female adult
Yunnan
Copyright © Hongliang Shi

Parena (Bothynoptera) tesari (Jedlicka, 1951)
Male adult
Taiwan
Copyright © Hongliang Shi

Parena kataevi Kirschenhofer, 2006 Holotype
Female adult

Copyright © Hongliang Shi

Parena (Bothynoptera) tesari (Jedlicka, 1951) Holotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) tesari (Jedlicka, 1951)
Male genitalia
A Yunnan. B Taiwan. C Zhejiang. Scale bar = 0.5 mm
Copyright © Hongliang Shi

Parena (Bothynoptera) kurosai Habu, 1967
Male adult
Guangdong
Copyright © Hongliang Shi

Parena (Bothynoptera) kurosai Habu, 1967
Male adult
Nepal
Copyright © Hongliang Shi

Parena (Bothynoptera) dorsigera (Schaum, 1863)
Male adult
Zhejiang
Copyright © Hongliang Shi

Parena (Bothynoptera) dorsigera (Schaum, 1863)
Male adult
Taiwan
Copyright © Hongliang Shi

Parena (Bothynoptera) dorsigera (Schaum, 1863)
Male adult
Nepal
Copyright © Hongliang Shi

Parena (Bothynoptera) dorsigera (Schaum, 1863)
Female adult
Yunnan (Kunming)
Copyright © Hongliang Shi

Bothynoptera perforata Bates Lectotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) dorsigera (Schaum, 1863) Holotype
Male adult

Copyright © Hongliang Shi

Parena (Bothynoptera) kurosai Habu, 1967
Male genitalia
Japan
Copyright © Hongliang Shi

Parena (Crossoglossa) laesipennis (Bates, 1873)
Male adult
Taiwan
Copyright © Hongliang Shi

Parena (Crossoglossa) laesipennis (Bates, 1873)
Female adult
Yunnan
Copyright © Hongliang Shi

Parena (Crossoglossa) laesipennis (Bates, 1873)
Male genitalia
right-lateral (r), ventral (v), left lateral (l), and dorsal (d) views. A Yunnan. B Yunnan. Scale bar = 0.5 mm
Copyright © Hongliang Shi

Parena (Crossoglossa) laesipennis (Bates, 1873)
Male genitalia
A Laos. B Taiwan. C Nepal. Scale bar = 0.5 mm
Copyright © Hongliang Shi

Parena (Crossoglossa) testacea (Chaudoir, 1872)
Male adult
Guandong
Copyright © Hongliang Shi

Parena (Crossoglossa) testacea (Chaudoir, 1872) Holotype
Male adult

Copyright © Hongliang Shi