Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov

Anillinus langdoni Sokolov & Carlton, 2004
Male adult
USA-TN, Sevier Co, GSMNP, secondary site, 490 m, ex leaf et log litter, 35°43.157' N, 83°23.309' W, 19.04.2011, J.-S. Park det. I. Sokolov, 2013
Copyright © Roman Dudko

Anillinus affabilis (Brues, 1902) Lectotype
Male adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm.
Copyright © Igor Sokolov

Anillinus sinuatus (Jeannel, 1963) Holotype
Female adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm. (Deposited in NMNH)
Copyright © Igor Sokolov

Anillinus depressus (Jeannel, 1963) Holotype
Female adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm. (NMNH)
Copyright © Igor Sokolov

Anillinus forthoodensis Sokolov & Reddell, 2014 Holotype
Male adult
TX: Bell Co., Talking Crows Cave, Fort Hood, 4.V.2006, J. Fant, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #45.781 \ Holotype Anillinus forthoodensis Sokolov & Reddell 2014 [red label] \ CAS Type No. 18872
Copyright © Igor Sokolov

Anillinus affabilis (Brues, 1902)
Male adult
TEXAS, Travis County, Tooth Cave
Copyright © Igor Sokolov

Anillinus acutipennis Sokolov & Reddell, 2014 Paratype
Female adult
TX: Bell Co., Hidden Pit Cave, Fort Hood, 27.X.2007, J. Reddell, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #60.107
Copyright © Igor Sokolov

Anillinus comalensis Sokolov & Kavanaugh, 2014 Paratype
Female adult
TX: Comal Co., 7mi W New Braunfels, 27.I.1995, A. G. Grubbs
Copyright © Igor Sokolov

Anillinus wisemanensis Sokolov & Kavanaugh, 2014 Holotype
Male adult
TX: Hays Co., Wiseman Sink, 28.IV.1995, A. G. Grubbs \ Texas Memorial Museum Invertebrate Zool Coll #27.149 \ Holotype Anillinus wisemanensis Sokolov & Kavanaugh 2014 [red label] \ CAS Type No. 18873
Copyright © Igor Sokolov

Anillinus sinuatus (Jeannel, 1963)
Male adult
TEXAS, Bexar County
Copyright © Igor Sokolov

Anillinus felicianus Sokolov, 2021 Holotype
Male adult
(NMNH): \ USA-LA: West Feliciana Par., ~4mi SW Jackson at 30.794°N, 91.254°W, mixed pine-hardwood forest, soil washing/berlese Sokolov I.M. 19–25 Apr 2018
Copyright © Igor Sokolov

Anillinus relictus Sokolov, 2021 Holotype
Male adult
(CMNH): ALABAMA: Blount Co., Tidwell Hollow Nature Trail east of Oneonta. T. N. King April 1 1972 \ 4/1/72 o [handwritten] \ THOMAS C. BARR COLLECTION 2011 Acc. No. 38,014
Copyright © Igor Sokolov

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Anillinus bexarensis Sokolov, 2022 Paratype
Female adult
Up the Creek Cave, Bexar County, Texas ; 1.7 mm
Copyright © Igor Sokolov

Anillinus reddelli Sokolov, 2022 Paratype
Female adult
Beck Horse Cave, Williamson County, Texas
Copyright © Igor Sokolov

Medusapyga hehalis LaBonte, 2023 Paratype
Female adult
WASHINGTON, Thurston Co., 11.3 km NW Littlerock, elev. ~400 m, D.J. Cox and J.R. LaBonte, 6 May 2007, 46.9600°N 123.13450°W. Image from LaBonte & Maddison, 2023
Copyright © - - -

Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
Copyright © - - -

Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
Copyright © - - -

Anillinus hirsutus Sokolov, 2012 Paratype
Male adult
Paratype male, CMNH
Copyright © Curt Harden

Anillinus alleni Sokolov & Carlton, 2017
Male adult
USA: Arkansas, Stone Co., Ozark National Forest, Blanchard Springs Rec. Area. 27 September 2024. C.W. Harden & K.D. Karns leg. Under rock during rain. / Det. C.W. Harden 2024.
Copyright © Curt Harden

Bembidion (Trepanedoris) concretum Casey, 1918
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Plataphus) simplex Hayward, 1897
Male adult
Great Smoky Mountains National Park. photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Plataphus) gebleri (Gebler, 1833)
Male adult
Far East, Lazovskij Nat. Res., Formozova spring 43°00'06"N 134°05'13"E 10.VII.2005 K.Makarov leg.
L=4.5 mm
Copyright © Kirill Makarov

Bembidion (Peryphus) petrosum Gebler, 1833: 275
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphanes) lacunarium (Zimmermann, 1869)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Odontium) confusum Hayward, 1897
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Ochthedromus) bifossulatum (Leconte, 1852)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Notaphus) patruele Dejean, 1831
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Furcacampa) versicolor (Leconte, 1848)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Furcacampa) impotens Casey, 1918
Female adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Furcacampa) affine Say, 1823
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Eurytrachelus) nitidum (Kirby, 1837)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Amerizus (Amerizus) wingatei (Bland, 1863) (Bembidium)
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Asioperyphus) postremum Say, 1830
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Bracteon) foveum Motschulsky, 1844: 271
Male adult
Khabarovsk Reg., bank of Amur Rin. in Khabarovsk city 48°28'33"N 135°02'39"E 7.VIII.2003 I.Melnik leg.
L=6.0 mm
Copyright © Kirill Makarov

Bembidion (Bracteon) punctatostriatum Say, 1823
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Eupetedromus) graciliforme Hayward, 1897
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Colorado: Huerfano Co., Huerfano River at Badito, 37.7285°N 105.0167°W.
Scale bar is 1 mm.
Photo David R. Maddison & Alexander P. Swanson
Copyright © ---

Bembidion (Trichoplataphus) ozarkense Maddison & Hildebrandt, 2011 Paratype
Male adult
USA: Missouri: Carter Co., Current River at Van Buren, voucher DRMV100311
Copyright © - - -

Bembidion (Trichoplataphus) rolandi Fall, 1922
Male adult
USA: Virginia: Rockbridge Co., Maury River at Glasgow, voucher DRMV100193
Copyright © ---

Bembidion (Hydrium) laevigatum Say, 1830
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Thompson Island, 7.5.2008, Massachusetts
Images J. Rawlins
Copyright © ---

Asaphidion curtum curtum (Heyden, 1870)
Male adult
USA
image of Henri Goulet
Copyright © Henri Goulet

Bembidion (Peryphanes) grapii Gyllenhal, 1827
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov <br /> Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Peryphanes) yukonum Fall, 1926: 131
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) bruxellense Wesmael, 1835
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) obscurellum obscurellum (Motschulsky, 1845)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Ochthedromus) americanum Dejean, 1831
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Bembidion (Eupetedromus) immaturum Lindroth, 1954 Paratype
Male adult
W Newfoundland, Deer Lake, 30.V.51, Lindroth. 4254:2 (MZLU)
Copyright © Lund University

Bembidion (Peryphus) tetracolum tetracolum Say, 1823: 89
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Lionepha) disjunctum Lindroth, 1963 Holotype
Male adult

Copyright © - - -

Bembidion (Peryphus) transversale Dejean, 1831
Male adult
Images from David R. Maddison & Alexander P. Swanson, 2010
Copyright © - - -

Bembidion (Cillenus) palosverdes Kavanaugh and Erwin, 1992
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Notaphus) contractum Say, 1823
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Peryphodes) ephippigerum (Leconte, 1852)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Notaphus) indistinctum Dejean, 1831
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Lymnaeum) laticeps (Leconte, 1858)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Peryphus) sejunctum Casey, 1918
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Leuchydrium) tigrinum Leconte, 1879
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Bracteon) zephyrum Fall, 1910
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Amerizus (Amerizus) oblongulus (Mannerheim, 1852)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Asaphidion yukonense Wickham, 1919
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Lionepha) australerasa Maddison, 2020 Holotype
Male adult
USA: California: Amador Co., Oyster Lake, Silver Lake Campground
Copyright © David Maddison

Bembidion (Lionepha) tuulukwa Maddison, 2020
Male adult
USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls
Copyright © David Maddison

Bembidion (Lionepha) lindrothi Maddison & Sproul, 2020 Holotype
Male adult
USA: California: Tulare Co., outlet of Emerald Lake
Copyright © David Maddison

Bembidion (Lionepha) kavanaughi Maddison, 2020 Holotype
Male adult
USA: Oregon: Wallowa Co., Lostine River Valley
Copyright © David Maddison

Bembidion (Lionepha) probata (Casey, 1918)
Male adult
USA: Oregon: Klamath Co., Odell Creek near Davis Lake
Copyright © David Maddison

Bembidion (Lionepha) pseudoerasum Lindroth, 1963
Male adult
USA: California: El Dorado Co., Strawberry Creek at Sciots Camp
Copyright © David Maddison

Bembidion (Lionepha) osculans Casey, 1918
Male adult
USA: California: Tehama Co., Nanny Creek
Copyright © David Maddison

Bembidion (Lionepha) erasa Leconte, 1859
Male adult
USA: Oregon: Benton Co., Marys Peak
Copyright © David Maddison

Bembidion (Lionepha) casta Casey, 1918
Male adult
USA: Oregon: Benton Co., Marys Peak, Alder Creek Falls
Copyright © David Maddison

Bembidion (Lionepha) sequoiae Lindroth, 1963
Male adult
USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir
Copyright © David Maddison

Bembidion (Liocosmius) darlingtonielum Cooper & Maddison, 2014
Male adult

Copyright © David Maddison

Bembidion (Neobembidion) constricticolle Hayward, 1897
Male adult

Copyright © David Maddison

Bembidion (Notaphus) flohri Bates, 1878
Male adult

Copyright © David Maddison

Bembidion (Hydrium) laevigatum Say, 1830
Male adult
USA: Utah: Grand Co., Moab, Colorado River, 1190 m, 38.5739°N, 109.5769°W
Copyright © David Maddison

Bembidion (Hydrium) mimbres Maddison, 2020 Paratype
Male adult
USA: New Mexico: Grant Co., Gila River, Billings Vista, 1320 m, 32.8163°N, 108.6032°W, 11.viii.2005. DRM 05.043. D.R. & J.H.A. Maddison
Copyright © David Maddison

Bembidion (Ocydromus) corgenoma Maddison, 2020 Paratype
Male adult
USA: Oregon: Benton Co., Corvallis, Willamette River, 44.5491°N, 123.2449°W, 60 m
Copyright © David Maddison

Bembidion (Ocydromus) transversale Dejean, 1831
Male adult
Canada: Alberta: Burbank, junction of Red Deer and Blindman Rivers, 52.3542°N, 113.7556°W (voucher V101454)
Copyright © David Maddison

Bembidion (Ocydromus) erosum (Motschulsky, 1850)
Male adult
USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051°N, 124.1005°W (V101453)
Copyright © David Maddison

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Arizona: Cochise Co., San Pedro R at Charleston, 31.6239°N, 110.1722°W (voucher V101461)
Copyright © David Maddison

Bembidium haplogonum Chaudoir, 1868 (NEOTYPE)
Male adult
USA: California: Lake Co., North Branch Cache Creek at hwy 20, 305 m 38.9881°N, 122.5400°W
Copyright © David Maddison

Bembidion (Ocydromus) sarpedon Casey, 1918
Male adult
USA: Colorado: Las Animas Co., Cokedale, Reilly Canyon, 37.1346°N, 104.6114°W (V101459)
Copyright © David Maddison

Bembidion (Notaphus) brownorum Maddison, Sproul & Will, 2023 Holotype
Male adult
EMEC: 39.13841/−122.34621 USA: California: Colusa Co. Antelope Valley, Freshwater Creek uv light pan trap 133 m. 1.vii.2021 K.Will
Copyright © David Maddison

Bembidion (Notaphus) mormon Hayward, 1897
Male adult

Copyright © David Maddison

Bembidion (Notaphus) callens Casey, 1918 Paralectotypes
Female adult
Tuçson [Pima County], Arizona.
Copyright © David Maddison

Bembidion (Notaphus) obtusangulum Leconte, 1863
Male adult

Copyright © David Maddison

Bembidion (Liocosmius) cachagua Maddison, 2023 Holotype
Male adult
CAS: “U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln.”, “C.A.S. Accession 29-V-48 Cachagua Creek”, “David R. Maddison DNA3842 DNA Voucher”
Copyright © David Maddison

Odontium (Bracteon) lapponicum Zetterstedt, 1828
Female adult
Irkutsk reg., Usolskii distr., Malaya Belaya riv., Mutovka vill., 12-15.7.1996, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Peryphanes) stephensii Crotch, 1866
Female adult
CZECH REPUBLIC - Moravian-Silesian reg., Opava dist., Závada, 25.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Peryphus) femoratum femoratum Sturm, 1825
Female adult
Latvia, Jekabpils distr, Dunava, 23.7.2008 (DUBC)
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) petrosum petrosum
Male adult
E Siberia, Lake Baikal, Baikalo-Lenskii NP., Bereg Buryh Medvedei, Pokoiniki, 27.6.1995, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Horologion hubbardi Harden & Davidson, 2024 Paratype
Female adult
CWH-452 (VMNH).
Copyright © Curt Harden

Elaphropus (Tachyura) parvula (Dejean, 1831)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Tachyura) parvula (Dejean, 1831)
Male adult
length: 1,7-2,4 mm
Germania, Oberbayern, Eching
leg.det. A.Skale, 2001 (as Elaphropus)
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Tachys (Tachys) halophilus Lindroth, 1966
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Tachys (Polyderis) laevis (Say, 1823)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Tachys (Paratachys) proximus Say, 1823
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Pericompsus (Pericompsus) ephippiatus (Say, 1834)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Barytachys) tripunctatus (Say, 1830)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Tachyta (Tachyta) angulata Casey, 1918
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Tachys (Paratachys) oblitus Casey, 1918
Male adult
Copyright © Ben Smith
Copyright © - - -

Nothoderis rufotestacea Hayward, 1900
Male adult
USA, Arizona, Cochise Co.
Copyright © Terry Erwin

Nothoderis rufotestacea Hayward, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Paratachys) scitulus Leconte, 1848
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Pericompsus (Pericompsus) laetulus LeConte, 1851
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Polyderis) laeva (Say, 1823)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachys (Tachys) corax Leconte, 1852
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Barytachys) ferrugineus (Dejean, 1831)
Female adult
Alabama, Blount Co., Highland Lake. 9.iv.2019. T. King leg. Soil berlese
Copyright © Curt Harden

Tachys (Paratachys) austinicus Casey, 1918
Male adult
USA: Alabama, Blount Co., Highland Lake, 11.ix.2020, T. King leg. Window trap
Copyright © Curt Harden

Tachys (Paratachys) albipes Leconte, 1863
Male adult
"USA: Florida, Columbia Co., Osceola National Forest, Range Rd., 1.ii.2023, C.W. Harden leg., sifted from litter." ca. 3mm
Copyright © Curt Harden

Tachys (Paratachys) sagax Casey, 1918
Male adult
"USA: Alabama, Blount Co., Highland Lake, 24.viii.2020, T. King leg., window trap"
Copyright © Curt Harden

Tachys (Paratachys) ventricosus Leconte, 1863
Female adult
"USA: Virginia, Virginia Beach City, First Landing State Park, 18.iv.2018, C.W. Harden leg., sifting litter"
Copyright © Curt Harden

Mioptachys flavicauda (Say, 1823)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Mioptachys flavicauda (Say, 1823)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Mioptachys flavicauda (Say, 1823)
Male adult
Ukraine, Vic. Odessa, forest Lusanovskiy (Liski) 29.4.2019 Gontarenko A.V. leg
Copyright © Alexander Anichtchenko