Amerizus (Amerizus) wingatei (Bland, 1863) (Bembidium)
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Amerizus (Amerizus) oblongulus (Mannerheim, 1852)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Asaphidion curtum curtum (Heyden, 1870)
Male adult
USA
image of Henri Goulet
Copyright © Henri Goulet

Asaphidion yukonense Wickham, 1919
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Trepanedoris) concretum Casey, 1918
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Plataphus) simplex Hayward, 1897
Male adult
Great Smoky Mountains National Park. photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Plataphus) gebleri (Gebler, 1833)
Male adult
Far East, Lazovskij Nat. Res., Formozova spring 43°00'06"N 134°05'13"E 10.VII.2005 K.Makarov leg.
L=4.5 mm
Copyright © Kirill Makarov

Bembidion (Peryphus) petrosum Gebler, 1833: 275
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphanes) lacunarium (Zimmermann, 1869)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Notaphus) patruele Dejean, 1831
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Furcacampa) versicolor (Leconte, 1848)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Furcacampa) impotens Casey, 1918
Female adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Furcacampa) affine Say, 1823
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Eurytrachelus) nitidum (Kirby, 1837)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Asioperyphus) postremum Say, 1830
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Eupetedromus) graciliforme Hayward, 1897
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Colorado: Huerfano Co., Huerfano River at Badito, 37.7285°N 105.0167°W.
Scale bar is 1 mm.
Photo David R. Maddison & Alexander P. Swanson
Copyright © ---

Bembidion (Trichoplataphus) ozarkense Maddison & Hildebrandt, 2011 Paratype
Male adult
USA: Missouri: Carter Co., Current River at Van Buren, voucher DRMV100311
Copyright © - - -

Bembidion (Trichoplataphus) rolandi Fall, 1922
Male adult
USA: Virginia: Rockbridge Co., Maury River at Glasgow, voucher DRMV100193
Copyright © ---

Bembidion (Hydrium) laevigatum Say, 1830
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Thompson Island, 7.5.2008, Massachusetts
Images J. Rawlins
Copyright © ---

Bembidion (Peryphanes) grapii Gyllenhal, 1827
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov <br /> Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Peryphanes) yukonum Fall, 1926: 131
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) bruxellense Wesmael, 1835
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) obscurellum obscurellum (Motschulsky, 1845)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Eupetedromus) immaturum Lindroth, 1954 Paratype
Male adult
W Newfoundland, Deer Lake, 30.V.51, Lindroth. 4254:2 (MZLU)
Copyright © Lund University

Bembidion (Peryphus) tetracolum tetracolum Say, 1823: 89
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) transversale Dejean, 1831
Male adult
Images from David R. Maddison & Alexander P. Swanson, 2010
Copyright © - - -

Bembidion (Cillenus) palosverdes Kavanaugh and Erwin, 1992
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Notaphus) contractum Say, 1823
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Peryphodes) ephippigerum (Leconte, 1852)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Notaphus) indistinctum Dejean, 1831
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Lymnaeum) laticeps (Leconte, 1858)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Peryphus) sejunctum Casey, 1918
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Leuchydrium) tigrinum Leconte, 1879
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Liocosmius) darlingtonielum Cooper & Maddison, 2014
Male adult

Copyright © David Maddison

Bembidion (Neobembidion) constricticolle Hayward, 1897
Male adult

Copyright © David Maddison

Bembidion (Notaphus) flohri Bates, 1878
Male adult

Copyright © David Maddison

Bembidion (Hydrium) laevigatum Say, 1830
Male adult
USA: Utah: Grand Co., Moab, Colorado River, 1190 m, 38.5739°N, 109.5769°W
Copyright © David Maddison

Bembidion (Hydrium) mimbres Maddison, 2020 Paratype
Male adult
USA: New Mexico: Grant Co., Gila River, Billings Vista, 1320 m, 32.8163°N, 108.6032°W, 11.viii.2005. DRM 05.043. D.R. & J.H.A. Maddison
Copyright © David Maddison

Bembidion (Ocydromus) corgenoma Maddison, 2020 Paratype
Male adult
USA: Oregon: Benton Co., Corvallis, Willamette River, 44.5491°N, 123.2449°W, 60 m
Copyright © David Maddison

Bembidion (Ocydromus) transversale Dejean, 1831
Male adult
Canada: Alberta: Burbank, junction of Red Deer and Blindman Rivers, 52.3542°N, 113.7556°W (voucher V101454)
Copyright © David Maddison

Bembidion (Ocydromus) erosum (Motschulsky, 1850)
Male adult
USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051°N, 124.1005°W (V101453)
Copyright © David Maddison

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Arizona: Cochise Co., San Pedro R at Charleston, 31.6239°N, 110.1722°W (voucher V101461)
Copyright © David Maddison

Bembidium haplogonum Chaudoir, 1868 (NEOTYPE)
Male adult
USA: California: Lake Co., North Branch Cache Creek at hwy 20, 305 m 38.9881°N, 122.5400°W
Copyright © David Maddison

Bembidion (Ocydromus) sarpedon Casey, 1918
Male adult
USA: Colorado: Las Animas Co., Cokedale, Reilly Canyon, 37.1346°N, 104.6114°W (V101459)
Copyright © David Maddison

Bembidion (Notaphus) brownorum Maddison, Sproul & Will, 2023 Holotype
Male adult
EMEC: 39.13841/−122.34621 USA: California: Colusa Co. Antelope Valley, Freshwater Creek uv light pan trap 133 m. 1.vii.2021 K.Will
Copyright © David Maddison

Bembidion (Notaphus) mormon Hayward, 1897
Male adult

Copyright © David Maddison

Bembidion (Notaphus) callens Casey, 1918 Paralectotypes
Female adult
Tuçson [Pima County], Arizona.
Copyright © David Maddison

Bembidion (Notaphus) obtusangulum Leconte, 1863
Male adult

Copyright © David Maddison

Bembidion (Liocosmius) cachagua Maddison, 2023 Holotype
Male adult
CAS: “U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln.”, “C.A.S. Accession 29-V-48 Cachagua Creek”, “David R. Maddison DNA3842 DNA Voucher”
Copyright © David Maddison

Bembidion (Peryphanes) stephensii Crotch, 1866
Female adult
CZECH REPUBLIC - Moravian-Silesian reg., Opava dist., Závada, 25.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Peryphus) femoratum femoratum Sturm, 1825
Female adult
Latvia, Jekabpils distr, Dunava, 23.7.2008 (DUBC)
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) petrosum petrosum
Male adult
E Siberia, Lake Baikal, Baikalo-Lenskii NP., Bereg Buryh Medvedei, Pokoiniki, 27.6.1995, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) sordidum (Kirby, 1837)
Male adult
Edmonton, Alta. VIII.5.56 GE & KE Ball Bembidion sordidum Kby det. George E. Ball L=5.9 mm ex coll. ZISP
Copyright © Kirill Makarov

Bembidion (Asioperyphus) sordidum (Kirby, 1837)
Male genitalia
Edmonton, Alta. VIII.5.56 GE & KE Ball Bembidion sordidum Kby det. George E. Ball ex coll. ZISP
Copyright © Kirill Makarov

Bembidion (Trepanedoris) kieranae Maddison & Sproul, 2026 Paratype
Male adult
“USA: Oregon: Lane Co., SW Eugene, 288m 44.0031°N 123.1299°W, 5.vi.2015. DRM 15.069. Maddison, Kieran, Mendez, Zeleznik”, “David R. Maddison DNA4576 DNA Voucher” (OSAC)
Copyright © David Maddison

Bembidion (Trepanedoris) kieranae Maddison & Sproul, 2026 Paratype
Male genitalia
c: (USA: Oregon: Lane Co., SW Eugene; voucher DNA4507). d: (USA: Oregon: Lane Co., SW Eugene; voucher DNA4577).
Copyright © David Maddison

Bembidion (Trepanedoris) clemens clemens Casey, 1918
Male adult
USA: Arizona: Yavapai Co., Prescott)
Copyright © David Maddison

Bembidion (Trepanedoris) clemens Casey, 1918
Male genitalia
e: B. clemens clemens (USA: New Mexico: Taos Co., Rio Grande; voucher DNA4345). f: B. clemens disparile (USA: California: Kern Co., Davis Camp, Sequoia NF; voucher DNA4590).
Copyright © David Maddison

Bembidion (Trepanedoris) endeca Maddison & Sproul, 2026 Paratype
Male adult
Dairy Creek, Harney Co., Oregon
Copyright © David Maddison

Bembidion (Trepanedoris) endeca Maddison & Sproul, 2026 Paratype
Male genitalia
c: B. endeca (USA: Oregon: Harney Co., Steens Mountain Loop Road, Malheur NWR; voucher DNA4546). d: B. endeca (USA: Oregon: Klamath Co., Klamath Marsh NWR; voucher DNA4445).
Copyright © David Maddison

Bembidion (Trepanedoris) ampliceps Casey, 1918
Male genitalia
a: Bembidion ampliceps (USA: California: Kern Co., Lake Isabella at mouth of French Gulch; voucher DNA4570). b: B. ampliceps (USA: California: Kern Co., Stable Creek at Sawmill Rd.; voucher DNA4329).
Copyright © David Maddison

Bembidion (Trepanedoris) connivens (Leconte, 1852)
Male genitalia
g: B. connivens, southern form (USA: California: Tehama Co., Red Bluff; voucher DNA4343). h: B. connivens, northern form (USA: Oregon: Lincoln Co., Siletz River E of Kernville; voucher DNA1759). Scale bars 0.1 mm.
Copyright © David Maddison

Bembidion (Trepanedoris) connivens (Leconte, 1852)
Male adult
USA: California: Tehama Co., Red Bluff
Copyright © David Maddison

Bembidion (Trepanedoris) frontale (Leconte, 1848)
Male adult
Canada: Ontario: Dwight
Copyright © David Maddison

Bembidion (Trepanedoris) frontale (Leconte, 1848)
Male genitalia
a: Bembidion frontale (Canada: Ontario: Arrowhead Provincial Park; DNA1335). b: B. frontale (USA: Vermont: Chittenden Co., mouth of Winooski River). d: B. frontale (USA: New Hampshire: Carroll Co., Saco River at North Conway).
Copyright © David Maddison

Bembidion (Trepanedoris) acutifrons Leconte, 1879
Male adult
USA: Oregon: Harney Co., Marshall Pond, Malheur NWR
Copyright © David Maddison

Bembidion (Trepanedoris) altipaludis Maddison, 2026 Paratype
Female adult
USA: Oregon: Deschutes Co., Three Creek Meadow 1925m 44.1153°N 121.6263°W
Copyright © David Maddison

Bembidion (Trepanedoris) ampliceps Casey, 1918
Male adult
USA: California: Kern Co., Stable Creek
Copyright © David Maddison

Bembidion (Trepanedoris) anacalypsi Mendez & Maddison, 2026 Paratype
Male adult
USA: Oregon: Harney Co., Dairy Creek
Copyright © David Maddison

Bembidion (Trepanedoris) canadianum Casey, 1924
Female adult
USA: Montana: Beaverhead Co., Melrose
Copyright © David Maddison

Bembidion (Trepanedoris) microreticulatum Hatch, 1950
Male adult
USA: Washington: Lewis Co., Centralia
Copyright © David Maddison

Bembidion (Trepanedoris) siticum Casey, 1918
Female adult
USA: Oregon: Harney Co., Malheur NWR
Copyright © David Maddison

Bembidion (Lionepha) disjunctum Lindroth, 1963 Holotype
Male adult

Copyright © - - -

Bembidion (Lionepha) australerasa Maddison, 2020 Holotype
Male adult
USA: California: Amador Co., Oyster Lake, Silver Lake Campground
Copyright © David Maddison

Bembidion (Lionepha) tuulukwa Maddison, 2020
Male adult
USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls
Copyright © David Maddison

Bembidion (Lionepha) lindrothi Maddison & Sproul, 2020 Holotype
Male adult
USA: California: Tulare Co., outlet of Emerald Lake
Copyright © David Maddison

Bembidion (Lionepha) kavanaughi Maddison, 2020 Holotype
Male adult
USA: Oregon: Wallowa Co., Lostine River Valley
Copyright © David Maddison

Bembidion (Lionepha) probata (Casey, 1918)
Male adult
USA: Oregon: Klamath Co., Odell Creek near Davis Lake
Copyright © David Maddison

Bembidion (Lionepha) pseudoerasum Lindroth, 1963
Male adult
USA: California: El Dorado Co., Strawberry Creek at Sciots Camp
Copyright © David Maddison

Bembidion (Lionepha) osculans Casey, 1918
Male adult
USA: California: Tehama Co., Nanny Creek
Copyright © David Maddison

Bembidion (Lionepha) erasa Leconte, 1859
Male adult
USA: Oregon: Benton Co., Marys Peak
Copyright © David Maddison

Bembidion (Lionepha) casta Casey, 1918
Male adult
USA: Oregon: Benton Co., Marys Peak, Alder Creek Falls
Copyright © David Maddison

Bembidion (Lionepha) sequoiae Lindroth, 1963
Male adult
USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir
Copyright © David Maddison

Bembidion (Odontium) confusum Hayward, 1897
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Ochthedromus) bifossulatum (Leconte, 1852)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Bracteon) foveum Motschulsky, 1844: 271
Male adult
Khabarovsk Reg., bank of Amur Rin. in Khabarovsk city 48°28'33"N 135°02'39"E 7.VIII.2003 I.Melnik leg.
L=6.0 mm
Copyright © Kirill Makarov

Bembidion (Bracteon) punctatostriatum Say, 1823
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Ochthedromus) americanum Dejean, 1831
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Bembidion (Bracteon) zephyrum Fall, 1910
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Odontium (Bracteon) lapponicum Zetterstedt, 1828
Female adult
Irkutsk reg., Usolskii distr., Malaya Belaya riv., Mutovka vill., 12-15.7.1996, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko