Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour.
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male blue colour
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
Endophallus
endophallus ulrichi (as podolicus Semenov, 1898)
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Endophallus

Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Endophallus
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill.,
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Endophallus
W Ukraine, NW Podolian, Lviv reg., Javoriv distr.,near Stradch vill.,
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Endophallus
Czech, S Moravia, Pouzdrany
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis hortensis Linne, 1758
Endophallus
W-Ukraine, Male Polissya, near Kamenka-Buzka vill.
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Endophallus
Luxembourg, Bollendorf-Pont env., 3.III.1999
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Endophallus
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh.
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei linnaei Panzer, 1812
Endophallus
W.UKRAINE, Lviv Reg., NE Carpathians, Skolivski Beskidy Mts. Zelemyanka Mt. Rng
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Endophallus
Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) N-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) nitens Linne, 1758
Endophallus
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov

Carabus (Orinocarabus) sylvestris sylvestris (as micklitzi Sokolar, 1910)
Endophallus
W Austria, Styria, Mt Hochwechsel W slope, 47.531667, 15.906000 h=1640 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri Panzer, 1799
Endophallus
E-Austria, Wienerwald, Hochstraß am Lengbachl, h=575 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri (as floriani Penecke, 1905)
Endophallus
SE-Austria, Styria, Rein bei Graz VI.1970, R.Kenyery leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri (as floriani Penecke, 1905)
Endophallus
SE-Austria, Styria, Gratwein Kehrerwaldh=580 m, 47.130800, 15.270800
Copyright © Ruslan Panin

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Endophallus
Collection ground Mukawa cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Dromius (Dromius) agilis (Fabricius, 1787)
Endophallus
Karachay-Cherkessia, gorge. Dukka, h~2100 3-5.VII.2020 leg. M. Nabozhenko
Copyright © Kirill Makarov

Calathus (Neocalathus) micropterus (Dufischmid, 1812)
Endophallus
Sakhalin isl.
Copyright © Alexander Anichtchenko

Calathus (Neocalathus) melanocephalus melanocephalus
Endophallus
Left - Spain, Montes Cantabricos; right – paratype of C. (N.) anistschenkoi syn.
Copyright © Alexander Anichtchenko

Amara (Zezea) plebeja (Gyllenhal, 1810)
Endophallus
Collection ground Chitose city, Hokkaido Pref. pref. 34882
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Bradytus) majuscula (Chaudoir, 1850)
Endophallus
Collection ground Monbetsu cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) familiaris (Duftschmid, 1812)
Endophallus
Collection ground Kiyosato cho, Hokkaido Pref. pref. 38785
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) communis (Panzer, 1797)
Endophallus
det. K. Makarov
Copyright © Kirill Makarov

Poecilus (Poecilus) cupreus (Linne, 1758)
Endophallus
Turkije : Ardahan; Ardahan 7KM E Karst 13-7-1992 1900m. Heinz W. leg.
Copyright © Jan Muilwijk

Amara (Amara) pulpani Kult, 1949
Endophallus
W Ukraine, NE Carpathians, Lviv reg., Skole Beskids Mts, Grabnyk Range h=670 m,
Copyright © Denis Khrapov

Amara (Bradytus) majuscula (Chaudoir, 1850)
Endophallus
Buryatia, Tunkinskii Distr., environs of Tunka Vill. 3.VIII.2012 leg. A. Prosvirov
Copyright © Kirill Makarov

Amara (Bradytus) fulva (O.F. Muller, 1776)
Endophallus
Altai Terr., Smolenskii Distr., 5 km S of Smolenskoe Vill., Peschanaya River bank, 31.VII–2.VIII.2000 leg. A. Matalin
Copyright © Kirill Makarov

Amara (Bradytus) crenata Dejean, 1828: 507
Endophallus
Stavropol Terr., Izobil’nenskii Distr., Moskovskoe Vill. 4(17).III.1922 (collector unknown)
Copyright © Kirill Makarov

Amara (Bradytus) consularis (Duftschmid, 1812)
Endophallus
Armenia, mountains W of Lake Arpi, h ~ 2000 m, 8.VI.2002 leg. A. Rubenyan
Copyright © Kirill Makarov

Amara (Bradytus) apricaria (Paykull, 1790)
Endophallus
Kyrgyzstan, Moldy-Too Mt. Range, Kurtka River, h ~ 2200 m, 20–21.VII.1998 leg. A. Klimenko
Copyright © Kirill Makarov