Apotomus rufus (P. Rossi, 1790)
Male adult
S Spain,
8.IV.2010 Anichtchenko A. leg., det. & coll.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sclopeta (Fabricius, 1792)
Male adult
S Bosnia, Mostar; 6mm
Copyright © Aleš Sedláček

Aptinus (Aptinus) acutangulus Chaudoir, 1876
Female adult
SW-Montenegro, Dinaric Alps, Herceg Novi, Vrbanje, Mt Orjen h=1800m, 13-19.VII.2012, Manfred Egger leg.
L=13 mm

Copyright © Ruslan Panin

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinus) elegans Chaudoir, 1842
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachynolomus) immaculicornis Dejean, 1826
Male adult
Italy, Calabria, Asprom, San Luca (RC), 200m, 28.4.2002, leg. F. Angelini.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) explodens Duftschmid, 1812
Female adult
Bulgaria Chernomorets, 24.11.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Female adult
W Ukraine, Lviv city, Sykhiv (Zubra) forest, 49.791889, 24.047417, 13.VII.2024, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Male+Female adult
W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Male+Female adult
W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
Copyright © Ruslan Panin

Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
SW Slovenia, Ilirska Bistrica, Sviščaki, Snežnik,17-28.VI.2009, Manfred Egger leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus germarii Sturm, 1815
Male adult
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m, 3-13.VI.2010, Manfred Egger leg.
29 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) semigranosus (as montanus Winkler, 1939)
Female adult
W Ukraina, Chernivtsi Reg., Hertsa Distr., 23-25.VI.2017, Kanarsky Yu. leg. L=19 mm
Copyright © Ruslan Panin

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
Italy: Abruzzo, Pallano, h = 1000m, 18.06.1993, leg. Facovone
Copyright © Igor Solodovnikov

Calomera littoralis nemoralis Olivier, 1790
Male adult
UKRAINE,
Kherson Reg.,Genitchesk Distr., near Tehongar st., Kujuktuk isl., on saline 8-9.V.1995 Mishustin R. leg.
16 mm.
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
CZ, C Moravia, Přerov; 9mm
Copyright © Aleš Sedláček

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg., L=12 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Male adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg.
Copyright © Ruslan Panin

Cicindela (Cicindela) sylvicola Dejean, 1822
Female adult
W Ukraine, Podolian Upland, Gologory range, Mt Khom h=410 m, 49.681278, 24.279111, 16.VI.2025, Ludmyla Panina leg.
Copyright © Ruslan Panin

Polistichus connexus (Geoffroy, 1785)
Male adult
length: 7-9 mm
Korsika, Ghisonacia
leg. U.Eitschberger, 1987; det. U.Schmidt, 1994
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Polistichus fasciolatus (P. Rossi, 1790)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr., near Bogatoe Ushchele vill., 04.VIII.2009, Shehovcov A. leg.
L=8mm
photo Panin R.
Copyright © Ruslan Panin

Zuphium olens (P.Rossi, 1790)
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 27-30.VI.2006 K.Makarov leg.
L=8.7 mm
Copyright © Kirill Makarov

Polistichus connexus (Geoffroy, 1785)
Male adult

Copyright © Kirill Makarov

Drypta (Drypta) dentata (P.Rossi, 1790)
Male adult

Copyright © Kirill Makarov

Parazuphium (Parazuphium) damascenum (Fairmaire, 1897)
Male adult
Morocco, Tagounite vill area 28.6.2010, Anichtchenko A. leg. & det. (aedeagus studied)
Copyright © Alexander Anichtchenko

Parazuphium (Parazuphium) chevrolatii chevrolatii Laporte, 1833a: 254
Male adult
Iran Gilan Elburz 1000m 5.viii.1968 W.Heinz leg. at light. J. Muilwijk det.
Copyright © Jan Muilwijk

Elaphrus (Elaphroterus) aureus aureus P.W.J. Muller, 1821
Male adult
Germany, Westfalen, Greven-Schmedehausen 14.5.2006 leg. C Kerkering (DUBC)
Copyright © Alexander Anichtchenko

Acupalpus (Acupalpus) flavicollis (Sturm, 1825)
Male adult
L=3,6 mm
RUSSIA, Moscow reg., Orekhovo-Zuevskiy reg., 3 km to S from Anciferovo settl., grassland. 16.V.2001
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Acinopus (Acinopus) picipes (Olivier, 1795)
Female adult

Copyright © Kirill Makarov

Ophonus (Hesperophonus) azureus (Fabricius, 1775)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Ophonus) stictus Stephens, 1828
Female adult
W Ukraine, NW Podolian Upland, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.V-15.VI.2017, Panin R. & Kanarsky Yu. leg. L=15 mm
Copyright © Ruslan Panin

Ophonus (Incisophonus) incisus (Dejean, 1829)
Female adult
France Gard dep., Nimes 3.11.2013 (cAA) 14 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) laticollis Mannerheim, 1825
Male adult
Russia, NW Caucasus, Novorossiisk distr., Markoth Mts., 23.5.2005 (DUBC) det. Anichtchenko A. 2021; 10.5mm
Copyright © Alexander Anichtchenko

Bradycellus (Bradycellus) verbasci (Duftschmid, 1812)
Male adult
W Ukraine, Lviv city, Holosko Cemetery, 3.IX.2014, Panin R. leg. L=5.1 mm
Copyright © Denis Khrapov

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest, on the sands, 17.V-25.VI.2024, R.Panin leg., L=13 mm
Copyright © Ruslan Panin

Dromius (Dromius) fenestratus (Fabricius, 1794)
Male adult
length: 5-6,5 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2005
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Perigona (Trechicus) nigriceps (Dejean, 1831)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Odacantha (Odacantha) melanura Linne, 1767
Male adult
Moskow Reg., Istra Distr., env. Polevstschina Vill. 15.VI.2006 Konoshenko leg.
L=7.0 mm
Copyright © Kirill Makarov

Microlestes corticalis (L. Dufour, 1820)
Male adult
Spain, Algeciras vic., Arroyo de la Cava, 36.145849, -5.481230 12-13.05.2014 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Perigona (Trechicus) nigriceps (Dejean, 1831)
Female adult
Belarus, Brest reg.
Copyright © Igor Solodovnikov

Licinus (Licinus) cassideus Fabricius, 1792: 148 (Carabus)
Female adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr.,near Bogatoe Ushchele vill.04.VIII.2009, Shehovcov A. leg.
female, 17mm
photo & det. Panin R.
Copyright © Ruslan Panin

Licinus (Neorescius) hoffmannseggii (Panzer, 1803)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Licinus (Licinus) depressus Paykull, 1790: 34
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Licinus (Licinus) silphoides (P. Rossi, 1790)
Female adult
Corse, Col de Bavella, 14.8.1966
Copyright © Alexander Anichtchenko

Loricera (Loricera) pilicornis pilicornis Fabricius, 1775: 293
Female adult
L=7,4 mm
RUSSIA, Moscow city, East distr., 9/2 Khimushina str., frontgarden, under stone. 17.II.2002
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Loricera (Loricera) pilicornis (Fabricius, 1775)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Leistus (Leistus) ferrugineus Linne, 1758
Male adult
Moskow Reg., Istra Distr., E shore of Istrinskoe Reserv., Novorakovo Vill. 16.IX.2007 K.Makarov leg.
L=6.8 mm
Copyright © Kirill Makarov

Notiophilus biguttatus Fabricius, 1779: 222 (Elaphrus)
Female adult
L=5,4 mm
RUSSIA, Moscow city, East distr., National park «Losiniy Ostrov», in leaf litter. 9.VI.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Notiophilus rufipes Curtis, 1829: pl. 254
Female adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Leistus (Leistus) piceus piceus Frolich, 1799
Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 9mm
Copyright © Aleš Sedláček

Leistus (Leistus) nitidus (Duftschmid, 1812)
Male adult
SW Romania, Munţii Retezat; 8mm
Copyright © Aleš Sedláček

Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843
Male adult
SW Bosnia, Bijakoviči - Križevac Mt.; 8mm
Copyright © Aleš Sedláček

Nebria (Nebria) dahlii dahilii Duftschmid, 1812: 50
Male adult
NW Slovenia, Trnovski Gozd Mts.; 8mm
Copyright © Aleš Sedláček

Nebria (Eunebria) psammodes (P. Rossi, 1792)
Male adult
C-Italy, Toscana, Pistoia (PT), Pracchia env.,16.IX.1971, Campadelli leg. L= 13 mm
Copyright © Ruslan Panin

Leistus (Pogonophorus) rufomarginatus (Duftschmid, 1812)
Male adult

Copyright © Kirill Makarov

Nebria (Eunebria) psammodes (P. Rossi, 1792)
Male adult

Copyright © Igor Solodovnikov

Chlaenius (Dinodes) decipiens L. Dufour, 1820b: 326
Female adult
S Ukraine, Crimea, Bakhchisaraysk distr., near Bogatoe Ushchele vill. 04.VIII.2009, Shehovcov A. leg. L=13mm photo Panin R.
Copyright © Ruslan Panin

Chlaenius (Epomis) circumscriptus Duftschmid, 1812: 166
Male adult
Italy, Puglia, S.Cataldo, Oasi WWF "Le Cesine" (LE),
29.VII.1995 F.Angelini leg.
male, 20 mm
photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Oodes helopioides helopioides (Fabricius, 1792)
Male adult
W. UKRAINE,
Ternopol Reg., Pidvolochisk distr., forest Bogdanivskyi, 24.V-15.VI.08 Panin R. leg.
long =9.5 mm
Copyright © Ruslan Panin

Chlaenius (Chlaeniellus) vestitus (Paykull, 1790)
Female adult
L=11,9 mm
MOLDOVA, Sakharna settl. Env. 30.VII.2007
leg. & det. Zubov A.
photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Callistus lunatus lunatus Fabricius, 1775: 247 (Carabus)
Female adult
L=6,2 mm
RUSSIA, Stavropol reg., Mineralovodskiy distr., 2,4 km to SW from sovkhoz Zheleznovodskiy, in ravine. h=654 m. N44*08.186’ E042*57.648’ 15-19.V.2006
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Chlaenius (Trichochlaenius) aeneocephalus aeneocephalus Dejean, 1826
Male adult
Azerbaidjan, Talysh, Avrora 18-05-1999
Copyright © Pascal Deschamps

Chlaenius (Chlaeniellus) flavipes Menetries, 1832
Male adult
Ю Азербайджан, Талыш Лерикский р-н, окр. п. Пештатюк 38°46'56"N 48°32'47"E 21-23.V.2012 h~400 leg. Д.Касаткин. det. K. Makarov
Copyright © Kirill Makarov

Chlaenius (Chlaenius) festivus velutinus Duftschmid, 1812: 168
Unsexed
Morocco : Oued Fourat, 13.xi.1960, 16 mm, Coll. N. Mal
Copyright © Noël Mal

Chlaenius (Chlaeniellus) nigricornis Fabricius, 1787: 202
Female adult

Copyright © Kirill Makarov

Chlaenius (Epomis) circumscriptus Duftschmid, 1812: 166
Male adult
Russia: Astrakhan reg.: B.Bolkhuny (vall. riv. Akkhtuba)
Copyright © Igor Solodovnikov

Agonum (Olisares) lugens (Duftschmid, 1812)
Male adult
Volgograd Reg., Pallasovka Distr., N shore of Elton Lake, right bank of Khara River 3.V.2009 leg.K.Makarov
8.8 mm
Copyright © Kirill Makarov

Paranchus albipes (Fabricius, 1796)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Paranchus albipes (Fabricius, 1796)
Male adult
W. UKRAINE
Lviv city, park Znesinnja 16.IV.2008, leg. Panin R.
8mm
Copyright © Ruslan Panin

Olisthopus rotundatus rotundatus (Paykull, 1790)
Male adult
L=7,8 mm<br /> RUSSIA, Tver reg., Likhoslavlskiy distr., 400 m to N from Domanikha village, road border. H=161 m. N57*13.755’ E035*45.435’ 4-8.IX.2006 <br /> leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763)
Male adult

Copyright © Kirill Makarov

Agonum (Olisares) sexpunctatum (Linne, 1758)
Male adult

Copyright © Kirill Makarov

Olisthopus fuscatus Dejean, 1828
Male adult
Turkey, Kirish distr., Camyuva riv., 28.9.2014, I. Enuschenko leg. det. Anichtchenko A. 2018 (aedeagus studied)
Copyright © Alexander Anichtchenko

Agonum (Olisares) hypocrita (Apfelbeck, 1904)
Male adult
NW-Ukraine, Polesia, Volhynia, Lyubeshiv Distr., Buchyn vill, swamped pinewood, «Selysko», 14.IX-17.X.2017, Kanarsky Yu. leg./det.Panin R.
Copyright © Ruslan Panin

Pterostichus (Platysma) niger niger s.str.
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.2007 Panin R. leg.

Copyright © Ruslan Panin

Amara (Amara) eurynota Panzer, 1796
Female adult

    W-UKRAINE
    NW Podolian, Lviv Reg., Brody Distr., near Jaseniv vill. Panin R. leg. 30.IV.08
    long 12mm

Copyright © Ruslan Panin

Amara (Curtonotus) gebleri Dejean, 1831
Male adult

Copyright © Kirill Makarov

Laemostenus (Antisphodrus) schreibersii (Kuster, 1846)
Female adult
NE Italy, Belluno reg., Cansiglio, Val Di Piera, 12.V-22.IX. 2007, Manfed Egger leg.
L=14.5 mm
Copyright © Ruslan Panin

Calathus (Neocalathus) ambiguus ambiguus Paykull, 1790: 130
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Melanius) aterrimus aterrimus (Herbst, 1784)
Male adult
W Ukraine, Lviv, Holosko, bank of Natural lake, 24.V-8.VII.2016, Panin R. leg. L=15 mm.
Copyright © Ruslan Panin

Laemostenus (Pristonychus) terricola punctatus Dejean, 1828
Male adult
Hungary, Budapest, Télényi, 10-12.VII.2004, I.Retezár leg. L=19 mm
Copyright © Ruslan Panin

Pedius inquinatus (Sturm, 1824)
Female adult
Italia, Basilicata (Matera), Foce Fiume Sinni, II.1979
Copyright © Riccardo Sciaky

Amara (Amara) ovata (Fabricius, 1792)
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. & det. (cAA). 8.5mm
Copyright © Alexander Anichtchenko

Amara (Amara) pulpani Kult, 1949
Male adult
W Ukraine, NE Carpathians, Lviv reg., Skole Beskids Mts, Grabnyk Range h=670 m, 5.V-9.VI.2021, D.Khrapov leg. /det. R. Panin, L= 7,5мм
Copyright © Denis Khrapov

Clinidium (Arctoclinidium) canaliculatum O.G.Costa, 1839d
Male adult
photo from www.zin.ru
Copyright © Kirill Makarov

Clivina (Clivina) fossor fossor (Linnaeus, 1758)
Male adult
L=5,3 mm
RUSSIA, Moscow city, SW distr., Jasenevo, Bitcevskiy les, under tree. 19.V.2000
leg., det., photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Dyschiriodes (Eudyschirius) globosus Herbst, 1784: 142
Male adult

Copyright © Kirill Makarov

Reicheadella (Chaetomargoreicheia) radei Bulirsch & Magrini, 2021 Holotype
Female adult
[Croatia] Velika pećina / Neorić, Sutina, Bazin / 3.v.2005; leg. P. Rade, (CNHM).
Copyright © Petr Bulirsch

Dyschiriodes (Eudyschirius) rufipes (Dejean, 1825)
Female adult
W Ukraina, Podolian Upland, Opillia, Ivano-Frankivsk reg., Kasova Gora near Burshtyn vill., steppe site, 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=2,9mm
Copyright © Denis Khrapov

Dyschirius (Dyschiriodes) auriculatus Wollaston, 1867
Male adult
Senegal, M`Bour, 14 km south, Club Aldiana, leg. U.Schmidt, 8.-20.X.1992; det. P.Bulirsch, 2010; 3.5 mm
Copyright © Udo Schmidt

Reicheidius frondicola (Reitter, 1881)
Female adult
Croatia, Dalmatian archipel., Brač isl., Vidova gora, 27.Х.2024, 567 м., D. Strelnikoff leg. (cAA).
Copyright © Alexander Anichtchenko

Siagona europaea Dejean, 1826: 468
Male adult
Uzbekistan, Kashkadarjnsk Reg., env. Mubarek 5-8.VI.2004.
L=10.3 mm
Copyright © Kirill Makarov

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Bembidion (Diplocampa) fumigatum (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Chlorodium) pygmaeum ab. bilunulatum Bielz, 1852
Male adult
N Moravia, Týn nad Bečvou; 2,5mm. det. J. Coulon, 2015
Copyright © Aleš Sedláček

Tachys (Paratachys) micros (Fischer von Waldheim, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum dalmatinum Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Dalmataphaenops chiarae Monguzzi, 1993: 235
Male adult

Copyright © Vittorino Monzini

Neolovricia ozimeci Lakota, Jazlic & Moravec, 2009 Holotype
Female adult
Spilja u Radinovcima, Dedici, Rastovac, Biokovo, 25.10.2006 Coleoptera (1) Leg: Roman Ozimec. Deposited in the Croatian Natural History Museum, Zagreb. Image from Lakota, Jazlic & Moravec, 2009
Copyright © - - -

Typhlotrechus velebiticus (Ganglbauer, 1904c)
Male adult

Copyright © Vittorino Monzini

Velebitaphaenops giganteus Casale & Jalzic, 2012
Female adult
Image from cavebiology.net
Copyright © Creative Commons

Tachyura (Tachyura) thoracicus (Kolenati, 1845)
Female adult
India, Uttarakhand, 14 km E Uttarkashi, 1500m, 10-12.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr as octostriata (Netolitzky, 1929)
Copyright © Alexander Anichtchenko