Apotomus rufus (P. Rossi, 1790)
Male adult
S Spain,
8.IV.2010 Anichtchenko A. leg., det. & coll.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sclopeta (Fabricius, 1792)
Male adult
S Bosnia, Mostar; 6mm
Copyright © Aleš Sedláček

Aptinus (Aptinus) acutangulus Chaudoir, 1876
Female adult
SW-Montenegro, Dinaric Alps, Herceg Novi, Vrbanje, Mt Orjen h=1800m, 13-19.VII.2012, Manfred Egger leg.
L=13 mm

Copyright © Ruslan Panin

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinus) elegans Chaudoir, 1842
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachynolomus) immaculicornis Dejean, 1826
Male adult
Italy, Calabria, Asprom, San Luca (RC), 200m, 28.4.2002, leg. F. Angelini.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) explodens Duftschmid, 1812
Female adult
Bulgaria Chernomorets, 24.11.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Female adult
W Ukraine, Lviv city, Sykhiv (Zubra) forest, 49.791889, 24.047417, 13.VII.2024, Panin R. leg.
Copyright © Ruslan Panin

Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
Italia, Lombardia (BG), Arera mt., 2100m<br /> Zambla, 19.6.2003 A. Degiovanni leg.<br /> det. V. Monzini, 2014 male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT) Paratype
Male+Female adult
Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env., VIII.1998, I.Rapuzzi leg.
photo & koll. Panin R.
female, 32 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
CZ; S Moravia, Břeclav; 26mm
Copyright © Aleš Sedláček

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
Italy: Abruzzo, Pallano, h = 1000m, 18.06.1993, leg. Facovone
Copyright © Igor Solodovnikov

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909
Female adult
NW Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Endophallus
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m,
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=22.5 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Male adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=23.5 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Dalmatia, Isl. Rab, 16-30.V.2019, G. Proscia leg.
Copyright © Ruslan Panin

Calomera littoralis nemoralis Olivier, 1790
Male adult
UKRAINE,
Kherson Reg.,Genitchesk Distr., near Tehongar st., Kujuktuk isl., on saline 8-9.V.1995 Mishustin R. leg.
16 mm.
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
CZ, C Moravia, Přerov; 9mm
Copyright © Aleš Sedláček

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg., L=12 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Male adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg.
Copyright © Ruslan Panin

Polistichus connexus (Geoffroy, 1785)
Male adult
length: 7-9 mm
Korsika, Ghisonacia
leg. U.Eitschberger, 1987; det. U.Schmidt, 1994
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Polistichus fasciolatus (P. Rossi, 1790)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr., near Bogatoe Ushchele vill., 04.VIII.2009, Shehovcov A. leg.
L=8mm
photo Panin R.
Copyright © Ruslan Panin

Zuphium olens (P.Rossi, 1790)
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 27-30.VI.2006 K.Makarov leg.
L=8.7 mm
Copyright © Kirill Makarov

Polistichus connexus (Geoffroy, 1785)
Male adult

Copyright © Kirill Makarov

Drypta (Drypta) dentata (P.Rossi, 1790)
Male adult

Copyright © Kirill Makarov

Parazuphium (Parazuphium) damascenum (Fairmaire, 1897)
Male adult
Morocco, Tagounite vill area 28.6.2010, Anichtchenko A. leg. & det. (aedeagus studied)
Copyright © Alexander Anichtchenko

Parazuphium (Parazuphium) chevrolatii chevrolatii Laporte, 1833a: 254
Male adult
Iran Gilan Elburz 1000m 5.viii.1968 W.Heinz leg. at light. J. Muilwijk det.
Copyright © Jan Muilwijk

Drypta (Drypta) dentata (P.Rossi, 1790)
Female adult
Republic of Belarus, Brest region, south of vill. Tomashevka, southern outflow of the Okhryovsky reservoir, under the pine branches, h = 160 m, 51.538967° N, 23.604839° E, 16.10.2019, leg. V.A. Kuznetzov
Copyright © Igor Solodovnikov

Ditomus tricuspidatus, Fabricius 1792
Male adult
length: 10,5-15 mm
Spanien, Totana
leg. Mager & Mühle, 1975
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Acupalpus (Acupalpus) flavicollis (Sturm, 1825)
Male adult
L=3,6 mm
RUSSIA, Moscow reg., Orekhovo-Zuevskiy reg., 3 km to S from Anciferovo settl., grassland. 16.V.2001
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Acupalpus (Acupalpus) parvulus (Sturm, 1825)
Male adult

Copyright © Kirill Makarov

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Acupalpus (Acupalpus) meridianus (Linnaeus, 1761)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
Moravia, Kobylí; 11 mm
Copyright © Aleš Sedláček

Ophonus (Hesperophonus) azureus (Fabricius, 1775)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) solitaris Dejean, 1829a: 337
Female adult
W-Ukraine, Transcarpathian reg., Chornagora Massif, 8 km NE Bohdan vill., h=800m, 9-29.VI..2014, Panin R. leg. L=10.5 mm
Copyright © Ruslan Panin

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest, on the sands, 17.V-25.VI.2024, R.Panin leg., L=13 mm
Copyright © Ruslan Panin

Dromius (Dromius) fenestratus (Fabricius, 1794)
Male adult
length: 5-6,5 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2005
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Perigona (Trechicus) nigriceps (Dejean, 1831)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Odacantha (Odacantha) melanura Linne, 1767
Male adult
Moskow Reg., Istra Distr., env. Polevstschina Vill. 15.VI.2006 Konoshenko leg.
L=7.0 mm
Copyright © Kirill Makarov

Microlestes corticalis (L. Dufour, 1820)
Male adult
Spain, Algeciras vic., Arroyo de la Cava, 36.145849, -5.481230 12-13.05.2014 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Perigona (Trechicus) nigriceps (Dejean, 1831)
Female adult
Belarus, Brest reg.
Copyright © Igor Solodovnikov

Chlaenius (Chlaenius) festivus (Panzer, 1796)
Female adult

Copyright © Kirill Makarov

Chlaenius (Epomis) circumscriptus Duftschmid, 1812: 166
Male adult
Italy, Puglia, S.Cataldo, Oasi WWF "Le Cesine" (LE),
29.VII.1995 F.Angelini leg.
male, 20 mm
photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Oodes helopioides helopioides (Fabricius, 1792)
Male adult
W. UKRAINE,
Ternopol Reg., Pidvolochisk distr., forest Bogdanivskyi, 24.V-15.VI.08 Panin R. leg.
long =9.5 mm
Copyright © Ruslan Panin

Licinus (Licinus) cassideus Fabricius, 1792: 148 (Carabus)
Female adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr.,near Bogatoe Ushchele vill.04.VIII.2009, Shehovcov A. leg.
female, 17mm
photo & det. Panin R.
Copyright © Ruslan Panin

Callistus lunatus lunatus Fabricius, 1775: 247 (Carabus)
Female adult
L=6,2 mm
RUSSIA, Stavropol reg., Mineralovodskiy distr., 2,4 km to SW from sovkhoz Zheleznovodskiy, in ravine. h=654 m. N44*08.186’ E042*57.648’ 15-19.V.2006
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Chlaenius (Chlaeniellus) nitidulus Schrank, 1781: 213
Male adult
W Ukraine, Lviv Region, Zhydahciv distr., Borodchytsi vill., 49°22'44"N, 24°13'37"E,
the right bank of Dniester river, 11-16.VI.2012, Panin.R leg.
L=12 mm
Copyright © Ruslan Panin

Chlaenius (Trichochlaenius) aeneocephalus aeneocephalus Dejean, 1826
Male adult
Azerbaidjan, Talysh, Avrora 18-05-1999
Copyright © Pascal Deschamps

Chlaenius (Chlaeniellus) flavipes Menetries, 1832
Male adult
Ю Азербайджан, Талыш Лерикский р-н, окр. п. Пештатюк 38°46'56"N 48°32'47"E 21-23.V.2012 h~400 leg. Д.Касаткин. det. K. Makarov
Copyright © Kirill Makarov

Licinus (Neorescius) hoffmannseggii (Panzer, 1803)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Chlaenius (Chlaenius) festivus velutinus Duftschmid, 1812: 168
Unsexed
Morocco : Oued Fourat, 13.xi.1960, 16 mm, Coll. N. Mal
Copyright © Noël Mal

Loricera (Loricera) pilicornis pilicornis Fabricius, 1775: 293
Female adult
L=7,4 mm
RUSSIA, Moscow city, East distr., 9/2 Khimushina str., frontgarden, under stone. 17.II.2002
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Loricera (Loricera) pilicornis (Fabricius, 1775)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Leistus (Leistus) ferrugineus Linne, 1758
Male adult
Moskow Reg., Istra Distr., E shore of Istrinskoe Reserv., Novorakovo Vill. 16.IX.2007 K.Makarov leg.
L=6.8 mm
Copyright © Kirill Makarov

Notiophilus biguttatus Fabricius, 1779: 222 (Elaphrus)
Female adult
L=5,4 mm
RUSSIA, Moscow city, East distr., National park «Losiniy Ostrov», in leaf litter. 9.VI.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Notiophilus rufipes Curtis, 1829: pl. 254
Female adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Leistus (Leistus) piceus piceus Frolich, 1799
Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 9mm
Copyright © Aleš Sedláček

Leistus (Leistus) nitidus (Duftschmid, 1812)
Male adult
SW Romania, Munţii Retezat; 8mm
Copyright © Aleš Sedláček

Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843
Male adult
SW Bosnia, Bijakoviči - Križevac Mt.; 8mm
Copyright © Aleš Sedláček

Nebria (Nebria) dahlii dahilii Duftschmid, 1812: 50
Male adult
NW Slovenia, Trnovski Gozd Mts.; 8mm
Copyright © Aleš Sedláček

Nebria (Eunebria) psammodes (P. Rossi, 1792)
Male adult
C-Italy, Toscana, Pistoia (PT), Pracchia env.,16.IX.1971, Campadelli leg. L= 13 mm
Copyright © Ruslan Panin

Leistus (Pogonophorus) rufomarginatus (Duftschmid, 1812)
Male adult

Copyright © Kirill Makarov

Nebria (Eunebria) psammodes (P. Rossi, 1792)
Male adult

Copyright © Igor Solodovnikov

Agonum (Olisares) lugens (Duftschmid, 1812)
Male adult
Volgograd Reg., Pallasovka Distr., N shore of Elton Lake, right bank of Khara River 3.V.2009 leg.K.Makarov
8.8 mm
Copyright © Kirill Makarov

Laemostenus (Laemostenus) venustus Dejean, 1828: 60 (Pristonychus)
Male adult
S Ukraine,
Crimea Peninsula, Angarskyi Pass, 26.XII.2010, Shaporinskiy V. leg./det. Panin R.
L=15 mm
Copyright © Ruslan Panin

Olisthopus rotundatus rotundatus (Paykull, 1790)
Male adult
L=7,8 mm<br /> RUSSIA, Tver reg., Likhoslavlskiy distr., 400 m to N from Domanikha village, road border. H=161 m. N57*13.755’ E035*45.435’ 4-8.IX.2006 <br /> leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Sphodrus leucophthalmus Linne, 1758
Female adult
W Austria, Hall in Tirol, 11.IV.1952, Richler leg.
L=26 mm

Copyright © Ruslan Panin

Laemostenus (Antisphodrus) elongatus elongatus (Dejean, 1828)
Female adult
SW Slovenia, Obrov, Poljane, 5-19.IX.2009, Manfed Egger leg.
L=16 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) schreibersii (Kuster, 1846)
Female adult
NE Italy, Belluno reg., Cansiglio, Val Di Piera, 12.V-22.IX. 2007, Manfed Egger leg.
L=14.5 mm
Copyright © Ruslan Panin

Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763)
Male adult

Copyright © Kirill Makarov

Agonum (Olisares) sexpunctatum (Linne, 1758)
Male adult

Copyright © Kirill Makarov

Laemostenus (Laemostenus) venustus Dejean, 1828: 60 (Pristonychus)
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Tiachiv distr., near Mala Ugolka vill., h=700m, 15.V-27.VI.2015, Panin R. & Kanarsky Yu. leg. L=16mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) cavicola cavicola Schaum, 1858b: 382 (Sphodrus)
Male adult
Slovenia, Poljane, Obrov, Mts Ćićarija, Cave Polina Peč (Cad. No. 938), II.2016, L=18 mm
Copyright © Ruslan Panin

Pterostichus (Platysma) niger niger s.str.
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.2007 Panin R. leg.

Copyright © Ruslan Panin

Zabrus (Zabrus) tenebrioides Goeze, 1777
Male adult

length: 12-16 mm
Austria, Neusiedler See
leg.det. R.Budberg, 1959

Copyright © U.Schmidt

Poecilus (Ancholeus) puncticollis (Dejean, 1828)
Male adult

Copyright © Kirill Makarov

Poecilus (Poecilus) striatopunctatus (Duftschmid, 1812)
Male adult
SE Slovakia, Malé Trakany; 8mm
Copyright © Aleš Sedláček

Pterostichus (Cophosus) cylindricus (Herbst, 1784)
Female adult
CZ; S Moravia, Břeclav; 20mm
Copyright © Aleš Sedláček

Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pedius inquinatus (Sturm, 1824)
Female adult
Italia, Basilicata (Matera), Foce Fiume Sinni, II.1979
Copyright © Riccardo Sciaky

Amara (Amara) tibialis (Paykull, 1798)
Male adult
W Ukraina, Podolian Upland, Kasova Gora near Burshtyn vill., steppe site, 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=5 mm
Copyright © Denis Khrapov

Amara (Zezea) fulvipes (Audinet-Serville, 1821)
Male adult
NW Greece, Ioannina env., Kalpaki, 2.6.2008 (DUBC). det. Anichtchenko A. 2022. 11 mm
Copyright © Alexander Anichtchenko

Pterostichus (Melanius) elongatus (Duftschmid, 1812)
Female adult
Marruecos, Gada de Debdou, 9.6.1993 (cAA)
Copyright © Alexander Anichtchenko

Clinidium (Arctoclinidium) canaliculatum O.G.Costa, 1839d
Male adult
photo from www.zin.ru
Copyright © Kirill Makarov

Clivina (Clivina) fossor fossor (Linnaeus, 1758)
Male adult
L=5,3 mm
RUSSIA, Moscow city, SW distr., Jasenevo, Bitcevskiy les, under tree. 19.V.2000
leg., det., photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Dyschiriodes (Eudyschirius) globosus Herbst, 1784: 142
Male adult

Copyright © Kirill Makarov

Reicheadella (Chaetomargoreicheia) radei Bulirsch & Magrini, 2021 Holotype
Female adult
[Croatia] Velika pećina / Neorić, Sutina, Bazin / 3.v.2005; leg. P. Rade, (CNHM).
Copyright © Petr Bulirsch

Dyschiriodes (Eudyschirius) rufipes (Dejean, 1825)
Female adult
W Ukraina, Podolian Upland, Opillia, Ivano-Frankivsk reg., Kasova Gora near Burshtyn vill., steppe site, 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=2,9mm
Copyright © Denis Khrapov

Dyschirius (Dyschiriodes) auriculatus Wollaston, 1867
Male adult
Senegal, M`Bour, 14 km south, Club Aldiana, leg. U.Schmidt, 8.-20.X.1992; det. P.Bulirsch, 2010; 3.5 mm
Copyright © Udo Schmidt

Reicheidius frondicola (Reitter, 1881)
Female adult
Croatia, Dalmatian archipel., Brač isl., Vidova gora, 27.Х.2024, 567 м., D. Strelnikoff leg. (cAA).
Copyright © Alexander Anichtchenko

Siagona europaea Dejean, 1826: 468
Male adult
Uzbekistan, Kashkadarjnsk Reg., env. Mubarek 5-8.VI.2004.
L=10.3 mm
Copyright © Kirill Makarov

Bembidion (Talanes) subfasciatum Chaudoir, 1850b: 187
Female adult
Krasnodar Reg., Taman Peninsula, Tuzlo Lake 6.V.2009
2.7 mm
Copyright © Kirill Makarov

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Thompson Island, 7.5.2008, Massachusetts
Images J. Rawlins
Copyright © ---

Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) inoptatum Schaum, 1857a: 149
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Dalmataphaenops chiarae Monguzzi, 1993: 235
Male adult

Copyright © Vittorino Monzini

Derossiella lukici Lohaj & Delić, 2019
Male adult
Image from Lohaj & Delić, 2019 (photo courtesy of Dušan Beňo).
Copyright © - - -

Typhlotrechus velebiticus (Ganglbauer, 1904c)
Male adult

Copyright © Vittorino Monzini

Tachys (Paratachys) fulvicollis (Dejean, 1831)
Female adult
NW-Ukraine, Volhynia, Turiysk env ., into the light, 6.VII.2015, Voytko P. leg./ R.Panin det., L=3,3 мм
Copyright © Denis Khrapov

Velebitaphaenops giganteus Casale & Jalzic, 2012
Female adult
Image from cavebiology.net
Copyright © Creative Commons

Bembidion (Princidium) punctulatum Drapiez, 1820
Male adult
Pechoro-Ilychskii N.P. (Печеро-Илычский зап)
Copyright © Kirill Makarov