Apotomus rufus (P. Rossi, 1790)
Male adult
S Spain,
8.IV.2010 Anichtchenko A. leg., det. & coll.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sclopeta (Fabricius, 1792)
Male adult
S Bosnia, Mostar; 6mm
Copyright © Aleš Sedláček

Aptinus (Aptinus) acutangulus Chaudoir, 1876
Female adult
SW-Montenegro, Dinaric Alps, Herceg Novi, Vrbanje, Mt Orjen h=1800m, 13-19.VII.2012, Manfred Egger leg.
L=13 mm

Copyright © Ruslan Panin

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinus) elegans Chaudoir, 1842
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachynolomus) immaculicornis Dejean, 1826
Male adult
Italy, Calabria, Asprom, San Luca (RC), 200m, 28.4.2002, leg. F. Angelini.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) explodens Duftschmid, 1812
Female adult
Bulgaria Chernomorets, 24.11.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Female adult
W Ukraine, Lviv city, Sykhiv (Zubra) forest, 49.791889, 24.047417, 13.VII.2024, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Male+Female adult
W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus germarii Sturm, 1815
Female adult
Croatia, Ivanščica Mts , Ivanec, 6-13.VI.2003, Krause Rene leg.
L=36 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
Male adult
CW-Serbia, Zlatar Mts, Nova Varoš, Zlatar Lake env., 1-12.VII.2007, P.Sima leg. L=30 mm
Copyright © Ruslan Panin

Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
NC-Italy, Bergamo, Mt. Presolana h=1500m, Colere env., 4-15.VI.2010, Manfred Egger leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček

Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Female adult
Bosnia, Mostar; 40 mm
Copyright © Aleš Sedláček

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Female adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 38 mm
Copyright © Ruslan Panin

Calomera littoralis nemoralis Olivier, 1790
Male adult
UKRAINE,
Kherson Reg.,Genitchesk Distr., near Tehongar st., Kujuktuk isl., on saline 8-9.V.1995 Mishustin R. leg.
16 mm.
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
CZ, C Moravia, Přerov; 9mm
Copyright © Aleš Sedláček

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg., L=12 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Male adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg.
Copyright © Ruslan Panin

Cicindela (Cicindela) sylvicola Dejean, 1822
Female adult
W Ukraine, Podolian Upland, Gologory range, Mt Khom h=410 m, 49.681278, 24.279111, 16.VI.2025, Ludmyla Panina leg.
Copyright © Ruslan Panin

Polistichus connexus (Geoffroy, 1785)
Male adult
length: 7-9 mm
Korsika, Ghisonacia
leg. U.Eitschberger, 1987; det. U.Schmidt, 1994
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Polistichus fasciolatus (P. Rossi, 1790)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr., near Bogatoe Ushchele vill., 04.VIII.2009, Shehovcov A. leg.
L=8mm
photo Panin R.
Copyright © Ruslan Panin

Zuphium olens (P.Rossi, 1790)
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 27-30.VI.2006 K.Makarov leg.
L=8.7 mm
Copyright © Kirill Makarov

Polistichus connexus (Geoffroy, 1785)
Male adult

Copyright © Kirill Makarov

Drypta (Drypta) dentata (P.Rossi, 1790)
Male adult

Copyright © Kirill Makarov

Parazuphium (Parazuphium) damascenum (Fairmaire, 1897)
Male adult
Morocco, Tagounite vill area 28.6.2010, Anichtchenko A. leg. & det. (aedeagus studied)
Copyright © Alexander Anichtchenko

Parazuphium (Parazuphium) chevrolatii chevrolatii Laporte, 1833a: 254
Male adult
Iran Gilan Elburz 1000m 5.viii.1968 W.Heinz leg. at light. J. Muilwijk det.
Copyright © Jan Muilwijk

Elaphrus (Elaphroterus) aureus aureus P.W.J. Muller, 1821
Male adult
Germany, Westfalen, Greven-Schmedehausen 14.5.2006 leg. C Kerkering (DUBC)
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
NW Podolian Upland, W UKRAINE,<br /> Lviv Reg., Zolochiv distr., near Gologirki vill., 30.04-19.05.07 Panin R.leg.<br /> L= 15mm
Copyright © Ruslan Panin

Bradycellus (Bradycellus) caucasicus Chaudoir, 1846: 187 (Acupalpus)
Male adult
L=4,0 mm
RUSSIA, Tver reg., Likhoslavlskiy distr., Domanikha village, in the garden, in soil. H=160 m. N57*13.629’ E035*45.675’ 21.VII.2005
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Acupalpus (Acupalpus) parvulus (Sturm, 1825)
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Male adult
NW Caucas, Krasnodar reg., SE Goryachii Kluch, леса между пос. Асфальтовая Гора и Садовый (нежилой) 21.V.2012 leg. М. Смирнов
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) laticollis Mannerheim, 1825
Male adult
Russia, NW Caucasus, Novorossiisk distr., Markoth Mts., 23.5.2005 (DUBC) det. Anichtchenko A. 2021; 10.5mm
Copyright © Alexander Anichtchenko

Bradycellus (Bradycellus) verbasci (Duftschmid, 1812)
Male adult
W Ukraine, Lviv city, Holosko Cemetery, 3.IX.2014, Panin R. leg. L=5.1 mm
Copyright © Denis Khrapov

Bradycellus (Bradycellus) harpalinus (Audinet-Serville, 1821)
Male adult
W Ukraine, Lviv city, Holosko, bank of Natural lake, R.Panin leg., 3.VI.2017, L=4,2 mm
Copyright © Denis Khrapov

Dromius (Dromius) fenestratus (Fabricius, 1794)
Male adult
length: 5-6,5 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2005
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Perigona (Trechicus) nigriceps (Dejean, 1831)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Odacantha (Odacantha) melanura Linne, 1767
Male adult
Moskow Reg., Istra Distr., env. Polevstschina Vill. 15.VI.2006 Konoshenko leg.
L=7.0 mm
Copyright © Kirill Makarov

Microlestes corticalis (L. Dufour, 1820)
Male adult
Spain, Algeciras vic., Arroyo de la Cava, 36.145849, -5.481230 12-13.05.2014 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Perigona (Trechicus) nigriceps (Dejean, 1831)
Female adult
Belarus, Brest reg.
Copyright © Igor Solodovnikov

Chlaenius (Dinodes) decipiens L. Dufour, 1820b: 326
Female adult
S Ukraine, Crimea, Bakhchisaraysk distr., near Bogatoe Ushchele vill. 04.VIII.2009, Shehovcov A. leg. L=13mm photo Panin R.
Copyright © Ruslan Panin

Chlaenius (Epomis) circumscriptus Duftschmid, 1812: 166
Male adult
Italy, Puglia, S.Cataldo, Oasi WWF "Le Cesine" (LE),
29.VII.1995 F.Angelini leg.
male, 20 mm
photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Licinus (Licinus) cassideus Fabricius, 1792: 148 (Carabus)
Female adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr.,near Bogatoe Ushchele vill.04.VIII.2009, Shehovcov A. leg.
female, 17mm
photo & det. Panin R.
Copyright © Ruslan Panin

Callistus lunatus lunatus Fabricius, 1775: 247 (Carabus)
Female adult
L=6,2 mm
RUSSIA, Stavropol reg., Mineralovodskiy distr., 2,4 km to SW from sovkhoz Zheleznovodskiy, in ravine. h=654 m. N44*08.186’ E042*57.648’ 15-19.V.2006
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Chlaenius (Trichochlaenius) aeneocephalus aeneocephalus Dejean, 1826
Male adult
Azerbaidjan, Talysh, Avrora 18-05-1999
Copyright © Pascal Deschamps

Licinus (Licinus) depressus Paykull, 1790: 34
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Chlaenius (Chlaenius) festivus velutinus Duftschmid, 1812: 168
Unsexed
Morocco : Oued Fourat, 13.xi.1960, 16 mm, Coll. N. Mal
Copyright © Noël Mal

Chlaenius (Chlaeniellus) nigricornis Fabricius, 1787: 202
Female adult

Copyright © Kirill Makarov

Chlaenius (Epomis) circumscriptus Duftschmid, 1812: 166
Male adult
Russia: Astrakhan reg.: B.Bolkhuny (vall. riv. Akkhtuba)
Copyright © Igor Solodovnikov

Licinus (Licinus) silphoides (P. Rossi, 1790)
Female adult
Corse, Col de Bavella, 14.8.1966
Copyright © Alexander Anichtchenko

Loricera (Loricera) pilicornis pilicornis Fabricius, 1775: 293
Female adult
L=7,4 mm
RUSSIA, Moscow city, East distr., 9/2 Khimushina str., frontgarden, under stone. 17.II.2002
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Loricera (Loricera) pilicornis (Fabricius, 1775)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Leistus (Leistus) ferrugineus Linne, 1758
Male adult
Moskow Reg., Istra Distr., E shore of Istrinskoe Reserv., Novorakovo Vill. 16.IX.2007 K.Makarov leg.
L=6.8 mm
Copyright © Kirill Makarov

Notiophilus biguttatus Fabricius, 1779: 222 (Elaphrus)
Female adult
L=5,4 mm
RUSSIA, Moscow city, East distr., National park «Losiniy Ostrov», in leaf litter. 9.VI.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Notiophilus rufipes Curtis, 1829: pl. 254
Female adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Leistus (Leistus) piceus piceus Frolich, 1799
Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 9mm
Copyright © Aleš Sedláček

Leistus (Leistus) nitidus (Duftschmid, 1812)
Male adult
SW Romania, Munţii Retezat; 8mm
Copyright © Aleš Sedláček

Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843
Male adult
SW Bosnia, Bijakoviči - Križevac Mt.; 8mm
Copyright © Aleš Sedláček

Nebria (Nebria) dahlii dahilii Duftschmid, 1812: 50
Male adult
NW Slovenia, Trnovski Gozd Mts.; 8mm
Copyright © Aleš Sedláček

Nebria (Eunebria) psammodes (P. Rossi, 1792)
Male adult
C-Italy, Toscana, Pistoia (PT), Pracchia env.,16.IX.1971, Campadelli leg. L= 13 mm
Copyright © Ruslan Panin

Leistus (Pogonophorus) rufomarginatus (Duftschmid, 1812)
Male adult

Copyright © Kirill Makarov

Nebria (Eunebria) psammodes (P. Rossi, 1792)
Male adult

Copyright © Igor Solodovnikov

Paranchus albipes (Fabricius, 1796)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Dolichus halensis Schaller, 1783: 317
Male adult
S Ukraine, Krym Peninsula,
Simpheropil, 15-30.VIII.2009, Shaporinskiy V. leg.
male, 19 mm
Copyright © Ruslan Panin

Platyderus (Platyderus) rufus Duftschmid, 1812: 105
Male adult
W Ukraine, NW Podolian,
Lviv reg., Zolochiv distr., near Gologirky vill., steppe sites, 26.V- 29.VI.2011, Panin R. leg.
L=7 mm

Copyright © Ruslan Panin

Olisthopus rotundatus rotundatus (Paykull, 1790)
Male adult
L=7,8 mm<br /> RUSSIA, Tver reg., Likhoslavlskiy distr., 400 m to N from Domanikha village, road border. H=161 m. N57*13.755’ E035*45.435’ 4-8.IX.2006 <br /> leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Laemostenus (Antisphodrus) schreibersii (Kuster, 1846)
Female adult
NE Italy, Belluno reg., Cansiglio, Val Di Piera, 12.V-22.IX. 2007, Manfed Egger leg.
L=14.5 mm
Copyright © Ruslan Panin

Anchomenus (Anchomenus) dorsalis (Pontoppidan, 1763)
Male adult

Copyright © Kirill Makarov

Calathus (Neocalathus) ambiguus ambiguus Paykull, 1790: 130
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Laemostenus (Pristonychus) terricola punctatus Dejean, 1828
Male adult
Hungary, Budapest, Télényi, 10-12.VII.2004, I.Retezár leg. L=19 mm
Copyright © Ruslan Panin

Laemostenus (Pristonychus) terricola terricola (Herbst, 1784)
Female adult
Belarus, Vitebsk reg., Vitebsk distr., vill. Novoselki
Copyright © Igor Solodovnikov

Agonum (Olisares) hypocrita (Apfelbeck, 1904)
Male adult
NW-Ukraine, Polesia, Volhynia, Lyubeshiv Distr., Buchyn vill, swamped pinewood, «Selysko», 14.IX-17.X.2017, Kanarsky Yu. leg./det.Panin R.
Copyright © Ruslan Panin

Poecilus (Poecilus) cupreus Linne, 1758
Female adult
W Ukraine, Lviv Reg., Javorov distr. near Ivano-Frankovo vill., Panin R. leg., 20.IV-4.V.2008 L=13mm
Copyright © Ruslan Panin

Pterostichus (Cheporus) burmeisteri Heer, 1838
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m 9.VI-25.VII.2007 Panin R. leg.
male,15 mm.

Copyright © Ruslan Panin

Amara (Amara) aenea DeGeer, 1774
Male adult

Copyright © Kirill Makarov

Amara (Curtonotus) gebleri Dejean, 1831
Male adult

Copyright © Kirill Makarov

Amara (Parapercosia) taurica Motschulsky, 1844
Male adult

Volgograd Area, Olkhovka Distr., env. Mikhailovka Vill., Markova Ravine 49°77956 N 44°40194 E h~103 4-5.VI.2008 Leg. I.Melnik
L=11.0 mm

Copyright © Kirill Makarov

Poecilus (Poecilus) koyi Germar, 1824: 16
Male adult
BiH; S Hercegovina, Ljubuski, Kravica slap; 14mm
Copyright © Aleš Sedláček

Pterostichus (Cophosus) cylindricus (Herbst, 1784)
Female adult
CZ; S Moravia, Břeclav; 20mm
Copyright © Aleš Sedláček

Pterostichus (Phonias) diligens (Sturm, 1824)
Male adult
Russia, Moskow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult

Copyright © Kirill Makarov

Pterostichus (Adelosia) macer (Marsham, 1802)
Male adult
Latvia, Kuldiga Co. 2019
Copyright © Alexander Anichtchenko

Clinidium (Arctoclinidium) canaliculatum O.G.Costa, 1839d
Male adult
photo from www.zin.ru
Copyright © Kirill Makarov

Clivina (Clivina) fossor fossor (Linnaeus, 1758)
Male adult
L=5,3 mm
RUSSIA, Moscow city, SW distr., Jasenevo, Bitcevskiy les, under tree. 19.V.2000
leg., det., photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Dyschiriodes (Eudyschirius) globosus Herbst, 1784: 142
Male adult

Copyright © Kirill Makarov

Reicheadella (Chaetomargoreicheia) radei Bulirsch & Magrini, 2021 Holotype
Female adult
[Croatia] Velika pećina / Neorić, Sutina, Bazin / 3.v.2005; leg. P. Rade, (CNHM).
Copyright © Petr Bulirsch

Dyschiriodes (Eudyschirius) rufipes (Dejean, 1825)
Female adult
W Ukraina, Podolian Upland, Opillia, Ivano-Frankivsk reg., Kasova Gora near Burshtyn vill., steppe site, 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=2,9mm
Copyright © Denis Khrapov

Dyschirius (Dyschiriodes) auriculatus Wollaston, 1867
Male adult
Senegal, M`Bour, 14 km south, Club Aldiana, leg. U.Schmidt, 8.-20.X.1992; det. P.Bulirsch, 2010; 3.5 mm
Copyright © Udo Schmidt

Reicheidius frondicola (Reitter, 1881)
Female adult
Croatia, Dalmatian archipel., Brač isl., Vidova gora, 27.Х.2024, 567 м., D. Strelnikoff leg. (cAA).
Copyright © Alexander Anichtchenko

Siagona europaea Dejean, 1826: 468
Male adult
Uzbekistan, Kashkadarjnsk Reg., env. Mubarek 5-8.VI.2004.
L=10.3 mm
Copyright © Kirill Makarov

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Thompson Island, 7.5.2008, Massachusetts
Images J. Rawlins
Copyright © ---

Bembidion (Diplocampa) fumigatum (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Porotachys bisulcatus (Nicolai, 1822)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ocys harpaloides (Audinet-Serville, 1821)
Male adult
Spain
Copyright © Kirill Makarov

Bembidion (Ocydromus) transsylvanicum Bielz, 1852
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Marmarosh Mts., near Lug vill., bank of Kuziy (Kizi) stream, 400m, 26.IV.2014, Panin R. & Kanarsky Yu. leg. L= 5 mm
Copyright © Ruslan Panin

Derossiella nonveilleri Queinnec, 2008
Male adult
Image from Lohaj & Delić, 2019 (photo courtesy of Dušan Beňo).
Copyright © - - -

Typhlotrechus velebiticus (Ganglbauer, 1904c)
Male adult

Copyright © Vittorino Monzini

Tachys (Paratachys) fulvicollis (Dejean, 1831)
Female adult
NW-Ukraine, Volhynia, Turiysk env ., into the light, 6.VII.2015, Voytko P. leg./ R.Panin det., L=3,3 мм
Copyright © Denis Khrapov

Odontium (Odontium) striatum (Fabricius, 1792)
Female adult
Latvia, Dunava
Copyright © Alexander Anichtchenko