Apotomus rufus (P. Rossi, 1790)
Male adult
S Spain,
8.IV.2010 Anichtchenko A. leg., det. & coll.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sclopeta (Fabricius, 1792)
Male adult
S Bosnia, Mostar; 6mm
Copyright © Aleš Sedláček

Aptinus (Aptinus) acutangulus Chaudoir, 1876
Female adult
SW-Montenegro, Dinaric Alps, Herceg Novi, Vrbanje, Mt Orjen h=1800m, 13-19.VII.2012, Manfred Egger leg.
L=13 mm

Copyright © Ruslan Panin

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinus) elegans Chaudoir, 1842
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachynolomus) immaculicornis Dejean, 1826
Male adult
Italy, Calabria, Asprom, San Luca (RC), 200m, 28.4.2002, leg. F. Angelini.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) explodens Duftschmid, 1812
Female adult
Bulgaria Chernomorets, 24.11.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Female adult
W Ukraine, Lviv city, Sykhiv (Zubra) forest, 49.791889, 24.047417, 13.VII.2024, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Male+Female adult
W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Female adult
NW Croatia, Bosanci, 1-12,VI.2004, Baros Slovak leg.<br /> L=33 mm
Copyright © Ruslan Panin

Cychrus caraboides Linnaeus, 1758
Male adult
L=17,0 mm
RUSSIA, Moscow city, East distr., National park «Losiniy Ostrov», spruce forest. 9.IX.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Cychrus attenuatus attenuatus Fabricius, 1792
Male+Female adult
W.UKRAINE, NE Carpathians,
Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt.Range h=700m, 26.V -1.VII.2010, Panin R. leg.

Copyright © Ruslan Panin

Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Female adult
Bosnia, Mostar; 40 mm
Copyright © Aleš Sedláček

Carabus (Pachystus) glabratus gibbosus Heyden, 1866
Male adult
N Croatia, Ivanščica Mts, Ivanec, 4-18.VI.2011, Krause Rene leg., L=32 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=22.5 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Male adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=23.5 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Male adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 37 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Female adult
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI. 2018, L=32 mm
Copyright © Ruslan Panin

Calomera littoralis nemoralis Olivier, 1790
Male adult
UKRAINE,
Kherson Reg.,Genitchesk Distr., near Tehongar st., Kujuktuk isl., on saline 8-9.V.1995 Mishustin R. leg.
16 mm.
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
CZ, C Moravia, Přerov; 9mm
Copyright © Aleš Sedláček

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Female adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg., L=12 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) arenaria viennensis (Schrank, 1781)
Male adult
W Ukraine, Lviv city, Holosko cemetery, on sands, 24.VII.2023, R.Panin leg.
Copyright © Ruslan Panin

Cicindela (Cicindela) sylvicola Dejean, 1822
Female adult
W Ukraine, Podolian Upland, Gologory range, Mt Khom h=410 m, 49.681278, 24.279111, 16.VI.2025, Ludmyla Panina leg.
Copyright © Ruslan Panin

Polistichus connexus (Geoffroy, 1785)
Male adult
length: 7-9 mm
Korsika, Ghisonacia
leg. U.Eitschberger, 1987; det. U.Schmidt, 1994
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Polistichus fasciolatus (P. Rossi, 1790)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr., near Bogatoe Ushchele vill., 04.VIII.2009, Shehovcov A. leg.
L=8mm
photo Panin R.
Copyright © Ruslan Panin

Zuphium olens (P.Rossi, 1790)
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 27-30.VI.2006 K.Makarov leg.
L=8.7 mm
Copyright © Kirill Makarov

Polistichus connexus (Geoffroy, 1785)
Male adult

Copyright © Kirill Makarov

Drypta (Drypta) dentata (P.Rossi, 1790)
Male adult

Copyright © Kirill Makarov

Parazuphium (Parazuphium) damascenum (Fairmaire, 1897)
Male adult
Morocco, Tagounite vill area 28.6.2010, Anichtchenko A. leg. & det. (aedeagus studied)
Copyright © Alexander Anichtchenko

Parazuphium (Parazuphium) chevrolatii chevrolatii Laporte, 1833a: 254
Male adult
Iran Gilan Elburz 1000m 5.viii.1968 W.Heinz leg. at light. J. Muilwijk det.
Copyright © Jan Muilwijk

Elaphrus (Elaphroterus) aureus aureus P.W.J. Muller, 1821
Male adult
Germany, Westfalen, Greven-Schmedehausen 14.5.2006 leg. C Kerkering (DUBC)
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Male adult
NW Caucas, Krasnodar reg., SE Goryachii Kluch, леса между пос. Асфальтовая Гора и Садовый (нежилой) 21.V.2012 leg. М. Смирнов
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Stenolophus (Stenolophus) steveni Krynicki, 1832
Female adult
SE Slovakia, Slovenský kras; 6,5mm
Copyright © Aleš Sedláček

Stenolophus (Egadroma) marginatus Dejean, 1829
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)
Male adult
S Tadzhikistan, Kurgan-Tube reg., 20km S Dzhilikul, 11.5.1999 (DUBC) 8.5mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) laticollis Mannerheim, 1825
Male adult
Russia, NW Caucasus, Novorossiisk distr., Markoth Mts., 23.5.2005 (DUBC) det. Anichtchenko A. 2021; 10.5mm
Copyright © Alexander Anichtchenko

Bradycellus (Bradycellus) verbasci (Duftschmid, 1812)
Male adult
W Ukraine, Lviv city, Holosko Cemetery, 3.IX.2014, Panin R. leg. L=5.1 mm
Copyright © Denis Khrapov

Bradycellus (Bradycellus) harpalinus (Audinet-Serville, 1821)
Male adult
W Ukraine, Lviv city, Holosko, bank of Natural lake, R.Panin leg., 3.VI.2017, L=4,2 mm
Copyright © Denis Khrapov

Dromius (Dromius) fenestratus (Fabricius, 1794)
Male adult
length: 5-6,5 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2005
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Perigona (Trechicus) nigriceps (Dejean, 1831)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Odacantha (Odacantha) melanura Linne, 1767
Male adult
Moskow Reg., Istra Distr., env. Polevstschina Vill. 15.VI.2006 Konoshenko leg.
L=7.0 mm
Copyright © Kirill Makarov

Microlestes corticalis (L. Dufour, 1820)
Male adult
Spain, Algeciras vic., Arroyo de la Cava, 36.145849, -5.481230 12-13.05.2014 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Perigona (Trechicus) nigriceps (Dejean, 1831)
Female adult
Belarus, Brest reg.
Copyright © Igor Solodovnikov

Chlaenius (Chlaenius) festivus (Panzer, 1796)
Female adult

Copyright © Kirill Makarov

Chlaenius (Dinodes) decipiens L. Dufour, 1820b: 326
Female adult
S Ukraine, Crimea, Bakhchisaraysk distr., near Bogatoe Ushchele vill. 04.VIII.2009, Shehovcov A. leg. L=13mm photo Panin R.
Copyright © Ruslan Panin

Chlaenius (Epomis) circumscriptus Duftschmid, 1812: 166
Male adult
Italy, Puglia, S.Cataldo, Oasi WWF "Le Cesine" (LE),
29.VII.1995 F.Angelini leg.
male, 20 mm
photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Oodes helopioides helopioides (Fabricius, 1792)
Male adult
W. UKRAINE,
Ternopol Reg., Pidvolochisk distr., forest Bogdanivskyi, 24.V-15.VI.08 Panin R. leg.
long =9.5 mm
Copyright © Ruslan Panin

Chlaenius (Chlaeniellus) vestitus (Paykull, 1790)
Female adult
L=11,9 mm
MOLDOVA, Sakharna settl. Env. 30.VII.2007
leg. & det. Zubov A.
photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Callistus lunatus lunatus Fabricius, 1775: 247 (Carabus)
Female adult
L=6,2 mm
RUSSIA, Stavropol reg., Mineralovodskiy distr., 2,4 km to SW from sovkhoz Zheleznovodskiy, in ravine. h=654 m. N44*08.186’ E042*57.648’ 15-19.V.2006
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Chlaenius (Trichochlaenius) aeneocephalus aeneocephalus Dejean, 1826
Male adult
Azerbaidjan, Talysh, Avrora 18-05-1999
Copyright © Pascal Deschamps

Licinus (Licinus) depressus Paykull, 1790: 34
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Chlaenius (Chlaenius) festivus velutinus Duftschmid, 1812: 168
Unsexed
Morocco : Oued Fourat, 13.xi.1960, 16 mm, Coll. N. Mal
Copyright © Noël Mal

Chlaenius (Epomis) circumscriptus Duftschmid, 1812: 166
Male adult
Russia: Astrakhan reg.: B.Bolkhuny (vall. riv. Akkhtuba)
Copyright © Igor Solodovnikov

Loricera (Loricera) pilicornis pilicornis Fabricius, 1775: 293
Female adult
L=7,4 mm
RUSSIA, Moscow city, East distr., 9/2 Khimushina str., frontgarden, under stone. 17.II.2002
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Loricera (Loricera) pilicornis (Fabricius, 1775)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Leistus (Leistus) ferrugineus Linne, 1758
Male adult
Moskow Reg., Istra Distr., E shore of Istrinskoe Reserv., Novorakovo Vill. 16.IX.2007 K.Makarov leg.
L=6.8 mm
Copyright © Kirill Makarov

Notiophilus biguttatus Fabricius, 1779: 222 (Elaphrus)
Female adult
L=5,4 mm
RUSSIA, Moscow city, East distr., National park «Losiniy Ostrov», in leaf litter. 9.VI.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Notiophilus rufipes Curtis, 1829: pl. 254
Female adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Leistus (Leistus) piceus piceus Frolich, 1799
Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 9mm
Copyright © Aleš Sedláček

Leistus (Leistus) nitidus (Duftschmid, 1812)
Male adult
SW Romania, Munţii Retezat; 8mm
Copyright © Aleš Sedláček

Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843
Male adult
SW Bosnia, Bijakoviči - Križevac Mt.; 8mm
Copyright © Aleš Sedláček

Nebria (Nebria) dahlii dahilii Duftschmid, 1812: 50
Male adult
NW Slovenia, Trnovski Gozd Mts.; 8mm
Copyright © Aleš Sedláček

Nebria (Eunebria) psammodes (P. Rossi, 1792)
Male adult
C-Italy, Toscana, Pistoia (PT), Pracchia env.,16.IX.1971, Campadelli leg. L= 13 mm
Copyright © Ruslan Panin

Leistus (Pogonophorus) rufomarginatus (Duftschmid, 1812)
Male adult

Copyright © Kirill Makarov

Nebria (Eunebria) psammodes (P. Rossi, 1792)
Male adult

Copyright © Igor Solodovnikov

Agonum (Olisares) lugens (Duftschmid, 1812)
Male adult
Volgograd Reg., Pallasovka Distr., N shore of Elton Lake, right bank of Khara River 3.V.2009 leg.K.Makarov
8.8 mm
Copyright © Kirill Makarov

Paranchus albipes (Fabricius, 1796)
Male adult
W. UKRAINE
Lviv city, park Znesinnja 16.IV.2008, leg. Panin R.
8mm
Copyright © Ruslan Panin

Olisthopus rotundatus rotundatus (Paykull, 1790)
Male adult
L=7,8 mm<br /> RUSSIA, Tver reg., Likhoslavlskiy distr., 400 m to N from Domanikha village, road border. H=161 m. N57*13.755’ E035*45.435’ 4-8.IX.2006 <br /> leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Sphodrus leucophthalmus Linne, 1758
Female adult
W Austria, Hall in Tirol, 11.IV.1952, Richler leg.
L=26 mm

Copyright © Ruslan Panin

Calathus (Neocalathus) ambiguus ambiguus Paykull, 1790: 130
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Agonum (Olisares) sexpunctatum (Linne, 1758)
Male adult

Copyright © Kirill Makarov

Laemostenus (Pristonychus) terricola terricola (Herbst, 1784)
Female adult
Belarus, Vitebsk reg., Vitebsk distr., vill. Novoselki
Copyright © Igor Solodovnikov

Calathus (Bedelinus) circumseptus Germar, 1824
Female adult
Italy, Toscana, Frienze, Scandicci, 24.11.2024 (DUBC). 12 mm
Copyright © Alexander Anichtchenko

Laemostenus (Antisphodrus) elongatus (Dejean, 1828)
Male adult
Bosnia Hercegovina, Mostar prov., Mt Cabuljia, 1100m, 20.6.2012 (DUBC). 14 mm
Copyright © Alexander Anichtchenko

Laemostenus (Laemostenus) venustus (Dejean, 1828)
Male adult
Italy, Toscana, Firenze, Parco della Misericordia, Borgo San Lorenzo 29.11.2024 (DUBC). 15 mm
Copyright © Alexander Anichtchenko

Amara (Amara) eurynota Panzer, 1796
Female adult

    W-UKRAINE
    NW Podolian, Lviv Reg., Brody Distr., near Jaseniv vill. Panin R. leg. 30.IV.08
    long 12mm

Copyright © Ruslan Panin

Zabrus (Zabrus) tenebrioides Goeze, 1777
Male adult

length: 12-16 mm
Austria, Neusiedler See
leg.det. R.Budberg, 1959

Copyright © U.Schmidt

Poecilus (Poecilus) koyi Germar, 1824: 16
Male adult
BiH; S Hercegovina, Ljubuski, Kravica slap; 14mm
Copyright © Aleš Sedláček

Poecilus (Poecilus) striatopunctatus (Duftschmid, 1812)
Male adult
SE Slovakia, Malé Trakany; 8mm
Copyright © Aleš Sedláček

Pterostichus (Cophosus) cylindricus (Herbst, 1784)
Female adult
CZ; S Moravia, Břeclav; 20mm
Copyright © Aleš Sedláček

Pedius inquinatus (Sturm, 1824)
Female adult
Italia, Basilicata (Matera), Foce Fiume Sinni, II.1979
Copyright © Riccardo Sciaky

Amara (Amara) ovata (Fabricius, 1792)
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. & det. (cAA). 8.5mm
Copyright © Alexander Anichtchenko

Amara (Celia) praetermissa (C.R. Sahlberg, 1827)
Female adult
W Ukraine, NE Carpathians, Skole Beskids Mts, Grabnyk Rng h=670 m, 16.VII-3.IX.2021, Khrapov D. leg./ Panin R. det., L=8.1 mm
Copyright © Denis Khrapov

Pterostichus (Adelosia) macer (Marsham, 1802)
Male adult
Latvia, Kuldiga Co. 2019
Copyright © Alexander Anichtchenko

Amara (Bradytus) fulva (O.F. Muller, 1776)
Male adult
Krasnodar Terr., Adlerskii Distr., 5.5 km NE of Krasnaya Polyana Vill., lower course of Achipse River, h ~ 600 m, 43°43'10"N, 40°15'25"E 19.VIII.2014 leg. K. Makarov, A. Matalin
Copyright © Kirill Makarov

Clinidium (Arctoclinidium) canaliculatum O.G.Costa, 1839d
Male adult
photo from www.zin.ru
Copyright © Kirill Makarov

Clivina (Clivina) fossor fossor (Linnaeus, 1758)
Male adult
L=5,3 mm
RUSSIA, Moscow city, SW distr., Jasenevo, Bitcevskiy les, under tree. 19.V.2000
leg., det., photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Dyschiriodes (Eudyschirius) globosus Herbst, 1784: 142
Male adult

Copyright © Kirill Makarov

Reicheadella (Chaetomargoreicheia) radei Bulirsch & Magrini, 2021 Holotype
Female adult
[Croatia] Velika pećina / Neorić, Sutina, Bazin / 3.v.2005; leg. P. Rade, (CNHM).
Copyright © Petr Bulirsch

Dyschiriodes (Eudyschirius) rufipes (Dejean, 1825)
Female adult
W Ukraina, Podolian Upland, Opillia, Ivano-Frankivsk reg., Kasova Gora near Burshtyn vill., steppe site, 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=2,9mm
Copyright © Denis Khrapov

Dyschirius (Dyschiriodes) auriculatus Wollaston, 1867
Male adult
Senegal, M`Bour, 14 km south, Club Aldiana, leg. U.Schmidt, 8.-20.X.1992; det. P.Bulirsch, 2010; 3.5 mm
Copyright © Udo Schmidt

Reicheidius frondicola (Reitter, 1881)
Female adult
Croatia, Dalmatian archipel., Brač isl., Vidova gora, 27.Х.2024, 567 м., D. Strelnikoff leg. (cAA).
Copyright © Alexander Anichtchenko

Siagona europaea Dejean, 1826: 468
Male adult
Uzbekistan, Kashkadarjnsk Reg., env. Mubarek 5-8.VI.2004.
L=10.3 mm
Copyright © Kirill Makarov

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Thompson Island, 7.5.2008, Massachusetts
Images J. Rawlins
Copyright © ---

Bembidion (Diplocampa) fumigatum (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) inoptatum Schaum, 1857a: 149
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Tachys (Paratachys) micros (Fischer von Waldheim, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Trepanes) octomaculatum (Goeze, 1777)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Dalmataphaenops chiarae Monguzzi, 1993: 235
Male adult

Copyright © Vittorino Monzini

Typhlotrechus velebiticus (Ganglbauer, 1904c)
Male adult

Copyright © Vittorino Monzini

Tachys (Paratachys) fulvicollis (Dejean, 1831)
Female adult
NW-Ukraine, Volhynia, Turiysk env ., into the light, 6.VII.2015, Voytko P. leg./ R.Panin det., L=3,3 мм
Copyright © Denis Khrapov

Velebitaphaenops giganteus Casale & Jalzic, 2012
Female adult
Image from cavebiology.net
Copyright © Creative Commons

Bembidion (Princidium) punctulatum Drapiez, 1820
Male adult
Pechoro-Ilychskii N.P. (Печеро-Илычский зап)
Copyright © Kirill Makarov