Loading...
Carabidae Carabidae of the World

Subfamily Carabinae Latreille, 1802

Photos
Main (random 10)  |  Main all  |  Adult  |  Immature stages  |  Endophallus  |  Labels  |  Biotope
Carabus (Morphocarabus) ??? as beheimi Niedl, 1957
Carabus (Morphocarabus) ??? as beheimi Niedl, 1957
Male adult
W Ukraine,
Transcarpathian Reg., Mukachevo, 22.V.2010
L=28 mm
photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
L=15.0 mm
Copyright © Evgeniy Zinoviev
Aplothorax burchelli Waterhouse, 1841
Aplothorax burchelli Waterhouse, 1841
Male adult
St. Helena island
Copyright © Boleslav Brezina
Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
Female adult
Armenia, Zangezur range, Meghri suburbs., Vahravar vill., Rivulet vall., h=1400 m Yuriy Shevnin leg.; 23.05.2007; 21 mm.
Copyright © Anton Menstschikov
Calosoma (Calosoma) schayeri schayeri Erichson, 1842
Calosoma (Calosoma) schayeri schayeri Erichson, 1842
Male adult
Australia N.T. Alice Springs, Airport
19.5.2000 M. Langer leg.
male, 21 mm
Copyright © Alexander Anichtchenko
Calosoma (Callisphaena) reticulatum reticulatum s.str.
Calosoma (Callisphaena) reticulatum reticulatum s.str.
Female adult
Germania, or. DDR, Bezirk Frankfurt, 24.V.1989
female, 25 mm
photo & det. Anichtchenko A.
Copyright © Alexander Anichtchenko
Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
Male adult
SE Canada, Nueva Escocia
19.5.1970
male, 17 mm
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) anthrax Semenov, 1900
Calosoma (Callisthenes) anthrax Semenov, 1900 Holotype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
Male adult
SE Kazakhstan Alma-Ata reg., 30 km<br /> N Sary-Ozek 28.4.1998 Klimenko leg.<br /> male, 22 mm
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) elegans amethystinus Semenov &  Redikorzev, 1928
Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
SE Kazakhstan, E Zailiyski mt. rng. Chilik riv.
Malybai vill. 28.4.1999 Klimenko A. leg.
male, 25 mm
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) elegans amethystinus Semenov &  Redikorzev, 1928
Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
26 mm.
Zailiyski Alatay.
06.1997 Chibilov E. leg.
Copyright © Sunvas
Calosoma (Callisthenes) elegans elegans s.str.
Calosoma (Callisthenes) elegans elegans s.str.
Male adult
SE Kazakhstan, Tchu-Iliyskiy Mts., Kordai pass, h=1500; A. Klimenko leg.; 20.04.1997;
24 mm.
photo A. Menstschicov
Copyright © Anton Menstschikov
Calosoma (Callisthenes) elegans manderstjernae Ballion, 1870
Calosoma (Callisthenes) elegans manderstjernae Ballion, 1870
Female adult
Kirgizia, S Kirgizskii mt. rng., 2200m
Kichi-Kimin riv., Ak-Tuz vill. 05.1998 Klimenko leg.
female, 32 mm
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Female adult
SE Kazakhstan, Zailijsky Mt. rng.,
Talgar env., 1500 m, 21.5.2000
female, 27 mm
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) eversmanni Chaudoir, 1850
Calosoma (Callisthenes) eversmanni Chaudoir, 1850
Male adult
Syria, Antilibanon
1.3.1991
male, 22 mm
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) fischeri Fischer, 1842
Calosoma (Callisthenes) fischeri Fischer, 1842
Male adult
Mongolia, Mandalgob, 20 km
N Amardalai vill. 11-12.6.2003 Anichtchenko A. leg
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) grumi Semenov, 1900
Calosoma (Callisthenes) grumi Semenov, 1900 Holotype
Female adult
collection ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) karelini Fischer-Waldheim, 1846
Calosoma (Callisthenes) karelini Fischer-Waldheim, 1846
Female adult
E Kazakhstan, S Alakol lake, Koktuma vill
1-10.5.1999 Nekroshevich S. leg.
female, 22 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) declivis rostislavi Semenov, 1906
Calosoma (Callisthenes) declivis rostislavi Semenov, 1906
Male adult
SE Kazakhstan, near Sary Ozek vill., A. Klimenko leg; 22.04.1997; 22 mm.;
photo, det. & coll. A. Menstschicov
Copyright © Anton Menstschikov
Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Male adult
Karzhantau mt. rng., Susungen riv.,
2300 m, 7.6.1998 Tretjakov leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) mirificus Casale, 1979
Calosoma (Callisthenes) mirificus Casale, 1979
Female adult
Afganistan, S slope of Paropamiz Mt. rng., near Sabzan pass, 2400m, 22.VI.1967 M.Stoljarov leg.
Collection of ZIN
det. D. Obydov 1999
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Calosoma (Callisthenes) panderi Fischer, 1822
Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr.,
near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov
male, 20 mm
Photo from www.zin.ru
Copyright © Maksim Smirnov
Calosoma (Callisthenes) regelianus A. Morawitz, 1886
Calosoma (Callisthenes) regelianus A. Morawitz, 1886
Female adult
Tadzikistan, Darvaz Ezgan 2.5.1971
female, 25 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Calosoma (Callitropa) externum Say, 1823: 149
Calosoma (Callitropa) externum Say, 1823: 149
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Calosoma (Calosoma) maximowiczi A. Morawitz, 1863: 20
Calosoma (Calosoma) maximowiczi A. Morawitz, 1863: 20
Female adult
China, Mt. Wangfoshan (1940 m.), Baokang Countri, Hubei; 20-28.06.2008, 30 mm.
Copyright © Anton Menstschikov
Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
Male adult
INDONESIA, Tanimbar Island, III.2009
photo & coll. N. Belonogov
Copyright © Nil Belonogov
Calosoma (Caminara) denticolle Gebler, 1833: 274
Calosoma (Caminara) denticolle Gebler, 1833: 274
Female adult
Burjatia, Selenga riv.,
Deben vill. 12.7.2003
Anichtchenko A. leg.
female, 25 mm
Copyright © Alexander Anichtchenko
Calosoma (Carabophanus) arrowi Breuning, 1928
Calosoma (Carabophanus) arrowi Breuning, 1928
Male adult
Ethiopia, Oromia reg., Arsi highlands, Wahoro env., Tiliku Lensho river valley10.-12.II.2010, J.Štastny lgt
photo Jaroslav Stastny
Copyright © Jaroslav Štastny
Calosoma (Castrida) alternans granulatum Perty, 1830
Calosoma (Castrida) alternans granulatum Perty, 1830
Female adult
Argentina, Entre Rios,
Liebig, nov.1997
long. 32 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko
Calosoma (Castrida) vagans Dejean,
Calosoma (Castrida) vagans Dejean,
Male adult
Chile, Santiago de Chile, Quilicura
19.9.1991 G. Aceituno leg.
male, 21 mm
Copyright © Alexander Anichtchenko
Calosoma (Charmosta) investigator Illiger, 1798
Calosoma (Charmosta) investigator Illiger, 1798
Male adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov
Calosoma (Charmosta) lugens Chaudoir, 1869f 372
Calosoma (Charmosta) lugens Chaudoir, 1869f 372
Male adult
Wen-dong, North Fengcheng City, Liaoning, China; 1-5.VII.2006
Copyright © Denis Potanin
Calosoma (Chrysostigma) calidum Fabricius, 1775
Calosoma (Chrysostigma) calidum Fabricius, 1775
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Calosoma (Ctenosta) bastardi Alluaud, 1925Calosoma (Ctenosta) senegalense Dejean, 1831
Calosoma (Ctenosta) senegalense Dejean, 1831
Female adult
length: 22-27 mm
Kenia Tsavo National Parc
leg. U.Schmidt, 1991; det. W.Lorenz, 2000
Foto: U.Schmidt, 2006
Copyright © U.Schmidt
Calosoma (Carabops) abyssinicum Gestro, 1880
Calosoma (Carabops) abyssinicum Gestro, 1880
Male adult
Ethiopia, Bale prov., Str. ZW. Sasheme u. Dodola, Mt. Mascall 3100-3300 m
21.VII.2005 Puchner leg.
male, 18 mm
det. Puchner F. 2005
photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Ceroglossus buqueti deuvei Jiroux, 1996
Ceroglossus buqueti deuvei Jiroux, 1996
Male adult
Chile, VIII Reg., 1000-1250 m, P.N. Nahuelbuta, (Piedra del Aguila) 7-12.XII.1999 Heinz leg.
male, 22 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko
Ceroglossus buqueti andestus Kraatz-Koschlau, 1887
Ceroglossus buqueti andestus Kraatz-Koschlau, 1887
Male adult
CHILE, Liquine; 05/12.02.2005; C.Saavedra leg.; photo A. Menstschicov
Copyright © Anton Menstschikov
Ceroglossus buqueti subnitens Kraatz-Koschlau, 1885
Ceroglossus buqueti subnitens Kraatz-Koschlau, 1885
Male adult
Chile, IX reg., Cherquenco Vn. Llaima, 800-1000m, 29-30.I.1999 Heinz leg.
male, 21 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko
Ceroglossus buqueti sybarita Gerstaecker, 1858
Ceroglossus buqueti sybarita Gerstaecker, 1858
Female adult
Chile, Chiloe, 160m, 8 km N Butalcura 22-24.XII.1998 det. Heinz
female, 24 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Ceroglossus chilensis cyanicollis Kraatz, 1887
Ceroglossus chilensis cyanicollis Kraatz, 1887
Male adult
Chile (VIII reg), Umg. Lag. de la Laja
950 m, 19-20.12.1998 Heinz leg.
26 mm
Copyright © Alexander Anichtchenko
Ceroglossus chilensis fallaciosus Kraatz, 1880
Ceroglossus chilensis fallaciosus Kraatz, 1880
Male adult
CHILE, Talca prov., Constitucion tn. env.- II.2008
photo Belonogov N.
Copyright © Nil Belonogov
Ceroglossus chilensis fisheti Jiroux, 1996
Ceroglossus chilensis fisheti Jiroux, 1996
Male adult
Chile (VIII reg), P.N. Nahuelbuta
(Piedra del Aguila) 700 m, 7-12.12.1999
Heinz leg.
28 mm
Copyright © Alexander Anichtchenko
Ceroglossus chilensis gloriosus Gerstaecker, 1858
Ceroglossus chilensis gloriosus Gerstaecker, 1858
Male adult
Chile (IX reg.), Malalcahuello
800 m, 9-10.12.1999 Heinz leg.
25 mm
Copyright © Alexander Anichtchenko
Ceroglossus chilensis latemarginatus Kraatz-Koschlau, 1889
Ceroglossus chilensis latemarginatus Kraatz-Koschlau, 1889
Male adult
Chile (VIII. Reg.) Pata de Gallina
650m, b.Contulmo 7-12.12.1999 Heinz leg.
27 mm
Copyright © Alexander Anichtchenko
Ceroglossus chilensis mochae Reed, 1874
Ceroglossus chilensis mochae Reed, 1874
Female adult
CHILE, Arauco prov., Isla Mocha. - II.2008
photo Belonogov N.
Copyright © Nil Belonogov
Ceroglossus chilensis resplendens Jaffrezic & Rataj, 2006Ceroglossus chilensis seladonicus Kraatz-Koschlau, 1887
Ceroglossus chilensis seladonicus Kraatz-Koschlau, 1887
Female adult
Chile, Malleco prov., Madalcahuello
enero 2003 G. Aceituno leg.
28 mm
Copyright © Alexander Anichtchenko
Ceroglossus chilensis solieri Roeschke, 1900
Ceroglossus chilensis solieri Roeschke, 1900
Female adult
Chile, Chiloe, 160m, 8 km N Butalcura 22-24.XII.1998 det. Heinz
female, 28 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Ceroglossus darwini darwini s.str.
Ceroglossus darwini darwini s.str.
Male adult
Chile, Chiloe, ca. 25 km NW Queilen, 350 m, (n. Castro), 26-27.XII.1999 Leg. Heinz
male, 22 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko
Ceroglossus magellanicus caburgansis Deuve, 1989
Ceroglossus magellanicus caburgansis Deuve, 1989
Male adult
CHILE. X Region T. de Carranco; 01/05 February. 2007; C.Saavedra leg.; photo A. Menstschicov
Copyright © Anton Menstschikov
Ceroglossus speciosus Gerstaecker, 1858
Ceroglossus speciosus Gerstaecker, 1858
Female adult
Chile, Chiloe, S. Pedro de Piruquiña 350 m, (n. Castro), 24-25.XII.1999 Heinz leg.
female, 23 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko
Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933
Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933 Paratype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Cychrus sehnali Haeckel, 2007
Cychrus sehnali Haeckel, 2007 Holotype
Male adult

Copyright © Boleslav Brezina
Cychropsis (Sinocychropsis) tryznai Haeckel & Sehnal, 2007
Cychropsis (Sinocychropsis) tryznai Haeckel & Sehnal, 2007 Holotype
Male adult
China - W Sichuan, 5.6.2006 West of Zhier (Zier) 4118 M
alpine forest N 28.20.507. E 101.28.014 leg. R Sehnal & M. Tryzna
Holotype
Copyright © Boleslav Brezina
Cychrus angustatus Hoppe & Hornschuch, 1825
Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
Italia, Lombardia (BG), Arera mt., 2100m<br /> Zambla, 19.6.2003 A. Degiovanni leg.<br /> det. V. Monzini, 2014 male, 20 mm
Copyright © Alexander Anichtchenko
Cychrus belousovi O.Berlov, 1997
Cychrus belousovi O.Berlov, 1997 Holotype
Male adult

Copyright © Oleg Berlov
Cychrus tuberculatus Harris, 1839
Cychrus tuberculatus Harris, 1839
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Cychropsis (Shuocychropsis) paramontana Sehnal & Hackel, 2006 HolotypeScaphinotus (Scaphinotus) viduus Dejean, 1826
Scaphinotus (Scaphinotus) viduus Dejean, 1826
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Scaphinotus (Steniridia) andrewsii amplicollis (Casey, 1920)
Scaphinotus (Steniridia) andrewsii amplicollis (Casey, 1920)
Male adult
USA Virginia, Smoth Co.,
Whitetop Mt., 10.7.1995 Trejbal leg.
male, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Scaphinotus (Steniridia) guyoti LeConte, 1866
Scaphinotus (Steniridia) guyoti LeConte, 1866
Male adult
USA Georgia, Brasstown, Towns Co.,
Bald Mt., 16.7.2000 Sehnal leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Scaphinotus (Steniridia) tricarinatus (Casey, 1914)
Scaphinotus (Steniridia) tricarinatus (Casey, 1914)
Male adult
USA N.C., BL. R. PKW - 430
MI. 8.7.1995 Sehnal leg.
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Scaphinotus (Steniridia) valentinei Barr,
Scaphinotus (Steniridia) valentinei Barr,
Male adult
USA W, Virginia, Spruce Knob Mt.,
4.7.1996 leg. Sehnal
male, 20 mm
Copyright © Alexander Anichtchenko
Scaphinotus (Nomaretus) bilobus Say, 1825
Scaphinotus (Nomaretus) bilobus Say, 1825
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Sphaeroderus canadensis Chaudoir, 1861
Sphaeroderus canadensis Chaudoir, 1861
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Sphaeroderus stenostomus lecontei Dejean, 1826
Sphaeroderus stenostomus lecontei Dejean, 1826
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Sphaeroderus nitidicollis brevoorti LeConte, 1848
Sphaeroderus nitidicollis brevoorti LeConte, 1848
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Pamborus alternans Latreille, 1817
Pamborus alternans Latreille, 1817
Female adult
Australia, NSW. Kerewong S.F.
21.12.1978
30 mm
Copyright © Alexander Anichtchenko
Pamborus brisbanensis Castelnau, 1867
Pamborus brisbanensis Castelnau, 1867
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko
Pamborus guerini Gory, 1830
Pamborus guerini Gory, 1830
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko
Pamborus opacus Gehin, 1885
Pamborus opacus Gehin, 1885
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko
Pamborus pradieri darlingtoni Banninger, 1940
Pamborus pradieri darlingtoni Banninger, 1940
Female adult
Australia, N.S.W. New England rng.,
Moonan Flats, 18.12.2000 T.Pollak leg.
21 mm
Copyright © Alexander Anichtchenko
Pamborus pradieri pradieri Chaudoir, 1869
Pamborus pradieri pradieri Chaudoir, 1869
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko
Pamborus punctatus Darlington, 1961 Paratype
Pamborus punctatus Darlington, 1961 Paratype Paratype
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko
Pamborus tropicus Darlington, 1961 Paratype
Pamborus tropicus Darlington, 1961 Paratype Paratype
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin<br /> Deposited in Zoolog. Museum St. Petersburg.<br /> male, long. 22 mm<br /> photo B. Brezina
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887Carabus (Acathaicus) interruptocostatus Semenov, 1887 HolotypeCarabus (Acoptolabrus) lopatini Morawitz, 1886
Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin
Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov
Carabus (Acrocarabus) guerini aldarkose Kabak, 1992Carabus (Acrocarabus) guerini guerini Fischer, 1842
Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov
Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 ParatypeCarabus (Alipaster) infantulus infantulus Morawitz, 1886Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype
Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina
Carabus (Apotomopterus) anxiensis Kleinfeld, 1998 Holotype & ParatypeCarabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype & ParatypeCarabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 HolotypeCarabus (Apotomopterus) casaleianus jigongshanicus Kleinfeld, 1998 Holotype & ParatypeCarabus (Apotomopterus) datianshanicus Kleinfeld, 1997 Holotype & ParatypeCarabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype
Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype Holotype
Female adult
as pervarius metallaicus
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) davidis neodavidis Kleinfeld & Schutze, 2001 Holotype & ParatypeCarabus (Apotomopterus) davidis viridithoracis Kleinfeld & Schutze, 2001 Holotype & ParatypeCarabus (Apotomopterus) davidis yizhangensis Kleinfeld, 2002Carabus (Apotomopterus) delavayi s.str. (as medea Kleinfeld, 1999)Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Female adult
Lijiang, Yunnan, Chine
female
photo Daniel Prunier
Copyright © Daniel Prunier
Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
Male+Female adult
Serbia, Debelo Brdo, 5.VI.2007, S. Sapaly leg. L=32 mm photo Panin R.
Copyright © Ruslan Panin
Carabus (Archicarabus) monticola fontanae Born, 1906
Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy,
Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.
female, 23 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822
Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male+Female adult
W.Ukraine.
Transcarpathian region,Vinogradiv distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.2008. Panin R. leg.
female 33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
W.Ukraine.
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male, 28 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Female adult
France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
female, 20 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Male adult
NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) olympiae Sella, 1855
Carabus (Chrysocarabus) olympiae Sella, 1855
Male+Female adult
N-Italy,
Piemonte, Bielmonte, 14-15.VI.1981, Pascal Stefani leg.
female, 34 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,
Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.
male, 23 mm

Copyright © Ruslan Panin
Carabus (Eupachys) glyptopterus Fischer, 1827
Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT)
Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT) Paratype
Male+Female adult
C China, Shaanxi,
Liupan Mts., Guan Shan Mt. h=800 m, , 34:50N/106:48 E, Team V.Siniaev leg.
female, 38 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
NE-FRANCE,
Lorraine Reg., Moselle, 15.II.2010, Pascal Stefani leg.
female, 26 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
Austria,
Weissenbach stt., III.1979, Kretschmer leg.
male, 24mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld
Carabus (Tribax) titan titan Zolotarev, 1913
Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) steveni steveni Menetries, 1832
Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
Karachaevo-Cherkessia,
Hudes riv., near Tsarskie vorota,
V.2009 Anichtchenko A. leg.
male, 25mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Tribax) puschkini puschkini Adams, 1817
Carabus (Tribax) puschkini puschkini Adams, 1817
Male adult
Carabus (Tribax) puschkini puschkini Adams, 1817
Armenia,
Shirak Province, Ashotsk, 18.VI.2001, P.V.Kazaryan leg.
male 25mm
Copyright © Ruslan Panin
Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
Male+Female adult
W-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
female, 25mm
Copyright © Ruslan Panin
Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 29 mm/male, 26mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) circassicus abagonensis Starck, 1894
Carabus (Tribax) circassicus abagonensis Starck, 1894
Male+Female adult
S Russia, NW Caucasus,
Adygea Rep., Mt.Abago, 26.VI.1999
L=24mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
Male adult
S Russia, Karacaj-Cherkessija,
near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009, Fominykh D. leg.
male, 28mm
Copyright © Ruslan Panin
Carabus (Tribax) apschuanus apschuanus Rost, 1893
Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male+Female adult
Carabus (Tribax) apschuanus apschuanus Rost, 1893    
Russia, Caucasus, Krasnodar reg.,
h=1200 m, 29.VII.2000
A. Zamotajlov leg.
Copyright © Safronov
Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult
NW-Caucasus,
Krasnodar reg., Mostovskoj distr., Skirda Mts. N. slope Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008, D.Fominyh leg.
male, 23mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine,
Ternopol reg., Pidvolohcisk distr., Supranivka vill.
3.VII.1908. Leg. Stobetskii. S.
Photo R. Panin
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male adult
W Ukraine,
Ternopil reg., Pidvolohcisk distr., Supranivka vill.
male, 26mm
The end of XIX the beginning of XX century
photo Panin R.
coll. SMNH Ukraine, Lviv
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus fossulatus Dejean, 1826
Carabus (Trachycarabus) sibiricus fossulatus Dejean, 1826
Male+Female adult
E.UKRAINE,
Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg.
female, 26mm/male, 24mm.
photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,
NW Podolian, Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010
leg. Panin R.
female, 22mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) perrini planus Gehin, 1885
Carabus (Trachycarabus) perrini planus Gehin, 1885
Male adult
S Russia,
Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg.
male, 26mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) perrini perrini Dejean, 1831
Carabus (Trachycarabus) perrini perrini Dejean, 1831
Male+Female adult
S Ukraine, Krym Peninsula,
near Perevalnoe vill., in steppe, 15.IV.2009, Shaporinskiy V. leg.
female, 28 mm
photo/det.Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) besseri Fischer, 1822
Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tomocarabus) marginalis Fabricius, 1794
Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) decolor Fischer, 1823
Carabus (Tomocarabus) decolor Fischer, 1823
Male adult
S Russia, NW-Caucasus,
Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
Fominykh D., Bondarenko A.S. leg.
male,17mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Male+Female adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
female, 24mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male+Female adult
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg. female, 32mm/male, 30mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
Copyright © Maksim Smirnov
Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male+Female adult
SE Siberia, Russia
Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominyh leg.
male, 24mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
Male adult
W Sayan Mts., Batenevskii Mt.rng., 12 km E Pervomaiskoe vill.,
Enisey riv., mountain steppe, 400m, 5.VII.2000 A. & N. Menschikovy leg.
female, 26mm

Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
S.Bohemia,
Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
male, 27mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
Male adult
W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., W Ukraine, near Gologirki vill., steppe sites, 19.V.-7.VI.2007, Panin R. leg. 25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Carabus (Morphocarabus) excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Male adult
Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian, Ternopil reg., Pidvilochisk distr., near Rosokhuvatets & Kamyanki vill., steppe sites, 24.V-15.VI.2008, Panin R. leg.
male, 24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour-26mm male blue colour-28mm
Copyright © Ruslan Panin
Carabus (Morphocarabus)  rothi incompsus (as mendax Csiki,1906)
Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
Male+Female adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
female, 32 mm
photo endophallus Solodovnikov I.A
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
Male+Female adult
female, 25 mm
SW-Romania, Banat, Resita - Valiug, 3-8.VI.2007, Krause R. leg.
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
Male+Female adult
female,28 mm
SE Hungary, Csongrбd, Makу env., 1-8.VIII.2004, Krause R., Slovak, Pittner leg
Copyright © Ruslan Panin
Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male+Female adult
NW-Caucasus, Karachay-Cherkessia, Teberda: Dzhamagat gorge, the left bank of the riv. Dzhamagat, 10.VII-28.VIII.2010 S.Pavluchuk leg.
female, 27 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus mingens Quensel, 1806
Carabus (Pachystus) hungaricus mingens Quensel, 1806
Male adult
male, 31 mm
SW-Russia: Stavropol Krai, near Stavropol city , Russkiy Les, 11.VI-28.IX.2010, Pavluchuk S. leg.
Copyright © Ruslan Panin
Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.
female, 26 mm
Copyright © Ruslan Panin
Carabus (Procechenochilus) kokujewi Semenov, 1898
Carabus (Procechenochilus) kokujewi Semenov, 1898
Female adult
RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
Copyright © Alexander Anichtchenko
Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Female adult
Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
female, 23 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Female adult
NW Caucasus, S Russia, Adygea Rep., Confluence of the river Belaya & Kisha h~700m, 24.V-7.VIII.2010, D.Fominyh leg.
female,32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus wolffii Dejean, 1826 loc.typ.
Carabus (Megodontus) violaceus wolffii Dejean, 1826 loc.typ.
Male+Female adult
SW-Romania: Banat, Moldova Noua, Ravenska viil., 21.V.-5.VI.2010, Krause Rene leg.
female, 32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus wolffii Dejean, 1826 (as pygmaeus Petri, 1912)
Carabus (Megodontus) violaceus wolffii Dejean, 1826 (as pygmaeus Petri, 1912)
Male adult
W Ukraine, NE Carpathians,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range h=2030m, alpine zone, 13-24.VI.2010 Panin R. leg./det.
male, 25 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko
Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
Male+Female adult
UKRAINE,
Kherson Reg., Golaya Pristan distr. near Rybalche vill. 6.IV.2000
long (female) 20mm
Copyright © Ruslan Panin
Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina
Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Male+Female adult


    Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
    Russia, N Burajatia,
    Barguzinskiy Mts., the upper reaches of M.Chiverkuy riv., h=2000m, Vashchenko S. leg.
    female, 21mm/male,19mm
    photo Panin R.



Copyright © Ruslan Panin
Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg.
male, 26mm
Copyright © Ruslan Panin
Carabus (Tachypus) auratus lasserrei Doue, 1855
Carabus (Tachypus) auratus lasserrei Doue, 1855
Male adult
SC-FRANCE, Midi-Pyrуnуes reg. Tarn, Saint Paul cap de Joux, 24.V.2003, A. Laforgue leg.
male, 24mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Male+Female adult
NW France,
Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
female, 28mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult

France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) michailovi Kabak, 1992
Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE-Kazakhstan,
S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
male,18mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2400m, 10.VII.2004
male, 18mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male+Female adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
male,17.5mm/female,19mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Female adult
Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000
female, 21 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya
Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999
2100 m, Brinev A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Male+Female adult
Russia, Sakhalin isl.,
Makarov distr., Lesnaja riv., VII.1992, M. Nesterov leg.
female,23mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians Fischer, 1827
Carabus (Sphodristocarabus) varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm
male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V.Kazaryan leg.
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey,
Artvin Prov., Hopa vers Borcka, Cankutaran Gecidi h=800m, 17.VI.2007 leg. D. Prunier
female, 24mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov
Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996
Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996 Holotype Paratype
Male adult
Holotype
male, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006 Holotype Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004Carabus (Pseudotribax) ferghanicus Breuning, 1933Carabus (Pseudocranion) sackeni sackeni Semenov, 1898Carabus (Pseudocranion) sackeni remondianus Deuve, 1996
Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Male+Female adult
NE KAZAKHSTAN,
S-Altays, Ulbinsky Mt. h=900m., Bobrovka vill., 24.VI.01, A. Klimenko leg.
female,25 mm
Copyright © Ruslan Panin
Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
4.VII.2002 A. & N. Menschikovy leg.
det. Mentschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
SE Siberia, E Sayan, Nizhneudinsk, h=500m, 21-26.VI.2006,
male, 26mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) maurus maurus Adams, 1817
Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia,
Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg.
female, 20mm/male,18mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus hopffgarteni Kraatz, 1878Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Male+Female adult
female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
P.V. Kazaryan leg.
Copyright © Ruslan Panin
Carabus (Procerus) scabrosus tauricus Bonelli, 1811Carabus (Cechenochilus) heydenianus Starck, 1889
Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) adangensis Gottwald, 1983
Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, NW-Caucasus:
Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
female,21mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, W Caucasus,
Karacaj-Cherkessija, Abishira-Akhuba Mnt Range, Area of pass Turii h~2650-2850m, 17.VII.2009, Fominykh D. leg.
male,17 mm
photo & coll. Panin R.
Copyright © Ruslan Panin
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora range., h=2020m. Panin R. leg. 28.VI-9.VIII.2008
male 19 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians,
Transcarpathian Reg., Rakhov Distr., Chornogora Mts Range h=2030m, alpin zone, 13-24.VI.2010 Panin R. leg.
female, 22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine, NW Podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg
L= 22 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) sternbergi Roeschke, 1898
Carabus (Eucarabus) sternbergi Roeschke, 1898
Female adult
Korea, Gyeongi Do,
Bokwang Sa 30.2.1976
female, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) ulrichi s.str. (as volhynicus Lutshnik, 1927)
Carabus (Eucarabus) ulrichi s.str. (as volhynicus Lutshnik, 1927)
Male+Female adult
W Ukraine, NW Podolian,
Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.2008, Panin R. & Solodovnikov I. leg.
Copyright © Ruslan Panin
Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) crassethoracis Kleinfeld, 1999Carabus (Microplectes) riedeli Menetries, 1832Carabus (Microplectes) riedeli Menetries, 1832
Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin
Carabus (Microplectes) convallium convallium Starck, 1889Carabus (Microplectes) convallium convallium Starck, 1889
Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin
Carabus (Microplectes) convallium argonautarum Semenov, 1898Carabus (Pachystus) tamsi tamsi Menetries, 1832
Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male+Female adult
Transcaucasia, Armenia,
Ashotsk, 18.VI.2001, P.V.Kazaryan leg.
female, 26mm//male, 24mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
Male+Female adult
Portugal,
Viseu distr., Gosende, h~1010m, 15.X.2009, D.Prunier leg.
female,26 mm
Copyright © Ruslan Panin
Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Male adult
Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
VI.2001 Sola Claudio leg.
male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornogora Mts. Pietros Mt. h=1850-2000m. alpine zone, 29.VI-28.VII.2007
Panin R. Leg.
Copyright © Safronov
Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
SW-France,
Foret d'Iraty Pyrenees-Atlantiques, d'Occabe, 15.VIII.1988
male,30 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Yamanashi pref., Kodachi, Kawaguchiko-cho, Minamitusru distr.
2.IX.2000 T. Kurihara leg.
male, 29 mm
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Female adult
15.VII.1998 JAPAN, Ichihara, env. Chiba
Copyright © Nil Belonogov
Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
Male adult
Japan, Yamanashi pref., Koganesawa-rindo, Otsuki city,
13.6.1990 K. Kurihara leg.
male, 20 mm
Copyright © Alexander Anichtchenko
Carabus (Megodontus) violaceus fiorii Born, 1901
Carabus (Megodontus) violaceus fiorii Born, 1901
Male+Female adult
NW-Italy,
Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
female, 30 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
Male+Female adult
S France,
Midi-Pyrénées Region, Ariège, Col du Chioula h=1500m, V-VI.2005, Pascal Stefani leg.
photo Panin R.
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Male adult
C.UKRAINE,
Poltava reg., near Novi Sanzhary, bank of Vorskla River, 1-2.VII.07, D.Kovalchuk leg.
male 25mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa, steppe sites, Panin R. leg. 1.V-6.VI.07
male, 26mm
Copyright © Safronov
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823 (as carbonatus Schaufuss, 1882)
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823 (as carbonatus Schaufuss, 1882)
Male adult
W.UKRAINE, Ukrainian Carpathians, Lviv Reg., Skolivski Beskidy Mts., near Skole, h=700m, 20.IX.2009, Panin R. leg. male, 33mm
Copyright © Ruslan Panin
Carabus (Megodontus) schoenherri schoenherri Fischer, 1822 (as sangilenus Obydov, 1997)
Carabus (Megodontus) schoenherri schoenherri Fischer, 1822 (as sangilenus Obydov, 1997)
Male adult
Tyva, Akademika Obrucheva mt.rng., 15km N Cherbi vill.,
Chailag ur., 7.7.1999 A. & N. Menshikovy leg.
male, 26mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) exaratus Quensel, 1806
Carabus (Megodontus) exaratus Quensel, 1806
Male adult
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush,
h=1400 m, 10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov
Carabus (Megodontus) exaratus Quensel, 1806
Carabus (Megodontus) exaratus Quensel, 1806
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg., Krasnodar City, 27.IX-07.X.07, D.D.Fominykh leg.
female,29mm/male, 27mm
Copyright © Ruslan Panin
Carabus (Megodontus) danilevskii Obydov, 1993
Carabus (Megodontus) danilevskii Obydov, 1993
Male adult
Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg.
L=17,0 mm
Copyright © Maksim Smirnov
Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin
Carabus (Pagocarabus) draco Brezina, 1999Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007)Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr., Bandivan, 8-21.X.1991 P.V.Kazaryan leg.
female, 20mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt range h=2020m alpine zone, Panin R. leg. 28.VI-9.VIII.2008
male 19 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
male, 29 mm
photo & collection Panin R.
Copyright © Ruslan Panin
Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male+Female adult
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
male 30mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Male adult
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
Karachaevo-Cherkessia, low cource of Daut river,
16-20.V.2009 Anichtchenko A. leg

Copyright © Alexander Anichtchenko
Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991
Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
Abkhasia,
Valley of river Bzyb near Mt. Achibakhu. h=1500m., A.S. Zamotajlov leg.
female: 33mm, 31mm
coll. R.Panin
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus caelatus Fabricius, 1801
Carabus (Megodontus) caelatus caelatus Fabricius, 1801
Male adult
Bosnia, Trebevic, Vidikovac-Brus, 1200m
21.11.1985 Mihljevic B. leg.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
Male+Female adult
Abkhasia, near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S., Bondarenko A.S. leg.
L=27mm

Copyright © Ruslan Panin
Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991
Zamotajlov & Miroshnikov leg.
Copyright © Sunvas
Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Male+Female adult
Caucasus, E Abkhasia, SW slope of Akiba mt.rng., 1200m, 7.V.-2.VII.1991 Zamotajlov & Miroshnikov leg.
male 37mm, female 38mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Female adult
S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas, 1900m, Khatsavitaya Mt., 4.VI-13.VII.1994
A.Miroshnikov leg.
male 25mm, female 26mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Female adult
NW-Caucasus, Russia,
Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 23 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
NW-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
male, 22mm
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Male+Female adult
W.Caucasus,
Psebai env., Nikitino, 2.V-3.VII.1993, leg. A.S.Zamotajlov
female 33mm
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
NW-Caucasus,Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
L= 42mm.
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg.,Krasnoarmeisk distr., Krasny Les, near Podlesnyi vill.,17-22.IV.2008, D.D.Fominykh leg.
female, 40mm
Copyright © Ruslan Panin
Carabus (Neoplectes) protensus olegi Belousov, 1992
Carabus (Neoplectes) protensus olegi Belousov, 1992 Paratype
Male adult
Caucasus, W Georgia, Svanetia, right bank of riv. Magana, 800m, 12.V.-4.VII.1991, Zamotajlov & Miroshnikov leg.
male, 31mm
photo and collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
male, 29 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
W-Caucasus:Sochi env., Aibga Mt. h=2000m., 06.VII.2000
male, 29mm
Copyright © Ruslan Panin
Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993
Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993 Paratype
Male adult
Abkhasia, Bzybian Mt. Range near Mt. Chibzharga., h=1900-2000m., A.S. Zamotajlov leg.
male, 28mm
photo & coll.Panin R.
Copyright © Ruslan Panin
Carabus (Neoplectes) starcki Heyden, 1885
Carabus (Neoplectes) starcki Heyden, 1885
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mt. Aishkho, Malaya Laba River, h=2200m., 14.VII.2008, D.Fominyh leg.
female-21mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis)
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Male+Female adult
NW Caucasus,
Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
female, 29mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Female adult
Russia,W Caucasus,
Krasnodar reg.,Chernogorie mts.,Chernogor mt. h=1700-1800m. 23.V.-30.VII.1999. A.S.Zamotajlov leg.
female 29mm.
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
male, 28mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus theseus Briansky, 1910
Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
female, 32mm
Copyright © Ruslan Panin
Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887Carabus (Aulonocarabus) canaliculatus rufinus Beheim & Breuning, 1943Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
Male adult
Arctic Russia, NW Siberia,
Yamal, 230 N from Labytnangi, Panta-Yakha Riv., 15-29.VI.2002
male,18mm.
Copyright © Ruslan Panin
Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995Carabus (Carabus) arvensis arvensis Herbst, 1784
Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin
Carabus (Carabus) arvensis arvensis Herbst, 1784
Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis carpathus Born, 1902
Carabus (Carabus) arvensis carpathus Born, 1902
Male+Female adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornohora range, h=1850 -2000m., 29.VI-28.VII.2007 Panin R. leg.
18mm.
Copyright © Ruslan Panin
Carabus (Carabus) granulatus leander Kraatz, 1878
Carabus (Carabus) granulatus leander Kraatz, 1878
Male+Female adult
Russia, C-Caucasus: Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992, Zamotajlov A. leg.
female,22mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W Ukraine,
Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
male, 21mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W-Ukraine, Roztochia, Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, Panin R. leg., 1-20.V.2005, male, 21 mm.
Copyright © Ruslan Panin
Carabus (Carabus) stscheglowi Mannerheim, 1827
Carabus (Carabus) stscheglowi Mannerheim, 1827
Male+Female adult
Central Russia , Voronezh region, near Grafskoe, 12.VIII.1984 long=20mm
Copyright © Ruslan Panin
Carabus (Cathaicus) brandti brandti Faldermann, 1835
Carabus (Cathaicus) brandti brandti Faldermann, 1835
Male adult
China, Beijing, Zhongguanchun, in the flower bed, 2007, Liu ye leg.;
Photo & det. by liuye
Copyright © Ye Liu
Carabus (Platycarabus) fabricii fabricii Panzer, 1812
Carabus (Platycarabus) fabricii fabricii Panzer, 1812
Male adult
NE-Italy,
Trentino-Alto Adige Reg,. Bolzano-Bozen Prov.,Val Aurina – Rottal 2050-2150m, 28.VI.1997, I.Rapuzzi leg.
male, 19mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus elytraviridis Kleinfeld, 2007
Carabus (Coptolabrus) formosus elytraviridis Kleinfeld, 2007 Holotype Paratype
Male+Female adult
China, Sichuan, Aba pref., Nanping Co., Jiuzhaigou (ca. 33.17N/103.54E), 2000 m, 30.5.-6.6.1990, coll. Kleinfeld.
HT 32 mm, PT 36 mm
Copyright © Frank Kleinfeld
Carabus (Carabus) sculpturatus Menetries, 1832
Carabus (Carabus) sculpturatus Menetries, 1832
Male+Female adult
Azerbaijan,
Talysh Mts. Hirkan National Park, Moscow Forest, 8-9.VII.1988, P.V.Kazaryan leg.
female, 22 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822
Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov
Carabus (Hygrocarabus) variolosus nodulosus Creutzer, 1799
Carabus (Hygrocarabus) variolosus nodulosus Creutzer, 1799
Male+Female adult
Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin
Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787
Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787
Male adult
W-Ukraine, NW Podolian,
Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg.
male, 27mm
Copyright © Ruslan Panin
Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787
Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787
Male+Female adult
W-Ukraine,
Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg.
female, 30mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
Male adult
NE-Italy,
Friuli Reg., Udine, Carnia, Pierabech h~1100m, I.Rapuzzi leg.
male, 23 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Male+Female adult
N-Korea: Pyonganam-Do, YangTok County,(39;127), 12-19,VI.2006
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Male+Female adult
W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens x illigeri Dejean, 1826
Carabus (Morphocarabus) praecellens x illigeri Dejean, 1826
Male+Female adult
W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
Male adult
W Slovenia,
Lokve, Trnovski Gozd h=1100m, 17.IV.2006, leg. I.Rapuzzi
male, 26mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT)
Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT) Paratype
Male+Female adult
Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env., VIII.1998, I.Rapuzzi leg.
photo & koll. Panin R.
female, 32 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male+Female adult
NE-Italy, Veneto Reg., Belluno, Forcella Lavardet h=2000m, VII.2002, I.Rapuzzi leg.
female, 24 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) lucepunctus ignipunctatus Rapuzzi, 2009 PT
Carabus (Diocarabus) lucepunctus ignipunctatus Rapuzzi, 2009 PT Paratype
Male+Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
femele, 18 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
Male adult
S Italy, Sicily, Province Palermo, Piana degli Albanesi h~700m, XI.2010, I.Rapuzzi leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
Male adult
N Korea, Ryanggang Prov., Mt. Tongjom Ridge h~1861m, (128.60E;41.20N) I. Rapuzzi leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Female adult
NE China, Liaoning, Anshan City, Mt.Qian Shan, 4-16.VII.2005, I.Rapuzzi leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Female adult
Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male adult
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill., 15.VII.2000, A.& N. Menstschicovy leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Pagocarabus) erwini erwini s.str.Carabus (Meganebrius) arunensis Heinertz, 1980Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Armenia,
Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
female 34mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m
Narusawa vill. 3.V.2005 Kurihara K. leg.
female, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005)Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr. Tsoghamarg vill, .21-22.V.1997, P.V.Kazaryan leg
female, 24 mm
Copyright © Ruslan Panin
Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko
Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Male adult
CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
K. & B. Brezina, M.Brychcin, M. Hackel
Copyright © Boleslav Brezina
Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
Male adult
Russia, Khakassia,
W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill., 11.VII.2000, A.& N. Menstschicovy leg.
L=21 mm
Copyright © Ruslan Panin
Carabus (Homoeocarabus) maeander maeander s.str.
Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Cathoplius) aliai Escalera, 1944Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)Carabus (Cechenotribax) petri Semenov & Znojko, 1932Carabus (Cephalornis) potanini (Semenov, 1887)
Carabus (Cephalornis) potanini (Semenov, 1887)
Male adult
China, S Gansu, 3400m, Minshan mts.,
Wenxian 8.92, J. M. Bousquet
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male+Female adult
S-Italy,
Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
female, 26mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
Female adult
W- FRANCE,
Bretagne Reg., Finistrere, Foret de Coatloc'h, XI.2006
female, 26 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
Male+Female adult
C-FRANCE,
Limousin Reg., Corrиze Dep., Aubazines, X.2005, Pascal Stefani leg.
female,26mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Female adult
NW-FRANCE,
Basse-Normandie Reg., Orne, Forêt de Bellême, 14.XII.2008, Pascal Stefani leg.
female, 26mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Male+Female adult
NW-FRANCE,
Basse-Normandie Reg,, Orne, Foret d' Ecouves, V.2009, Pascal Stefani leg.
male, 25 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens escheri (as laevipennis  Seidlitz, 1891)
Carabus (Chrysocarabus) auronitens escheri (as laevipennis Seidlitz, 1891)
Male adult
W Ukraine, NE Carpathians Mts.
Transcarpathian, Rakhov Distr., Chornogora range, Petros Mt.h=2010m, 9.VI.2009, R.Panin leg
male, 25 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as opacus Haury, 1878)
Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as opacus Haury, 1878)
Male+Female adult
W Ukraine, NE Carpathians Mts.,
Transcarpathian reg., Rakhov Distr., Chornogora range, h=2020m, alpin zone, 28.VI-22.VIII.2009, Panin R. leg.
female, 27mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826	(as barthei Lapouge, 1912)
Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826 (as barthei Lapouge, 1912)
Male+Female adult
SC -France,
Midi-Pyrйnйes reg., Ariиge, Foret de Belesta, XI.2008, J.Nicolas leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
Male+Female adult
SC-FRANCE,
Haute-Garonne, Col de Mentй h~1300m., II.1980. Kretschmer leg.
female, 23mm.
photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
Female adult
NW Italy,
Altare, Savona,15.VI.1995, Cartini A.leg.
female, 33mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) splendens Olivier, 1790
Carabus (Chrysocarabus) splendens Olivier, 1790
Male+Female adult
SE France,
Midi-Pyrйnйes reg., Ariиge, Vaychis, 22.X.2002, A.Laforgue leg.
female, 30mm.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) hispanus Fabricius, 1787
Carabus (Chrysocarabus) hispanus Fabricius, 1787
Male adult
FRANCE,
Tarn, Foret de la Gresigne, h=400m.VII.2005 Laforgue leg.
male-30mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis Mollard, 1999)
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis Mollard, 1999)
Male+Female adult
ANDORRA,
E-Pyrenees, Juberri, h=1350-1400m, 22.VI-18.VII.2007
female, 33mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male+Female adult
Russia, S Siberia, W Sayan, Ergaki Mt range, Nizhnyaya Buyba river h=1700 m, 19.VII.1999, A.& N. Menstschikovy leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Male adult
Russia, Khakassia,
Abakanskii Mt Range, Askiz river vall., 7 km E Kazanovka vill., 15.VI.2006, A.& N. Menstschikovy leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
Male adult
Russia, W Tuva,
Kyzyl-Tayga Mt Range h=1500 m, Ak-Suk riv vall., 21.VI.2006, A.& N. Menstschikovy leg.
L=16 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Male adult
NW Ukrainian Carpathians, Lviv Reg., Staryi Sambor Distr., near Dobromyl , h=500 m., 15-24.VI.2005, Panin R. leg., male, 28mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male+Female adult
Russia,Tuva,
Akademika Obrucheva Mt range, 15 km N Cherbi vill., Chailang ur. 7.VII.1999, A & N. Menstschikovy leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov
Carabus (Hemicarabus) nitens Linne, 1758
Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as  dacicus Csiki, 1906)
Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as dacicus Csiki, 1906)
Female adult
NW Romania , Bistrita Nasaud, Beclean env., 17.VI.2009, S.Budka leg.
female, 31 mm
Copyright © Ruslan Panin
Carabus (Heterocarabus) marietti marietti  (as ativsengueni Schweiger, 1962)Carabus (Goniocarabus) sogdianus Semenov, 1898Carabus (Gnathocarabus) kusnetzovi Semenov, 1903Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Male adult
N Korea, Samjiyon, Ryang Gang Province
6-12.6.2006 leg. Wong
Copyright © Boleslav Brezina
Carabus (Megodontus) kolbei aino Rost, 1908Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, S Russia, Adygea Rep., Lagonaki plateau, Site Murzikal h=1800-2000m, 24.V-18.IX.2010, D.Fominykh leg.
L= 22 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Male adult
Italy, Emilia reg., Modena, Castelnuovo Rangone, 31.XII.1991, Sola C. leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
Female adult
NC-Italy - S-Switzerland, Val Colla – Gazzirola, 22.VI.1999, C. Sola leg.
L=21mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Male adult
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill., 29.V.12.VI.2010, Krause Rene leg./det.Panin R. L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Male adult
SW-Romania:Banat, Bocsa/Coltan, 21.V-5.VI.2010, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Male adult
SE-Bulgaria, Burgas Prov., Primorsko env., 18-30.VI.2000, Krause Rene leg.
L=18 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793  (as slabai Niedl, 1968)
Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
Female adult
Czech, WS Bohemia,
Plzen Reg., Domazlice Distr., Babylon, h=500m, 30.VIII.2008, S.I. Budka leg.
L=24mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus violaceus Linne, 1758
Carabus (Megodontus) violaceus violaceus Linne, 1758
Male adult
S.Bohemia, Hluboka nad Vltavou, h=380 m, 6.VIII.2004, S. Budka leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis deyrollei Gory, 1839
Carabus (Carabus) arvensis deyrollei Gory, 1839
Female adult
NW Spain, Galicia Reg., Pontevedra Prov., A Lama, Xesta, h~650 m, 1.X.1994
L=17 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) victor wiedemanni Menetries, 1836
Carabus (Archicarabus) victor wiedemanni Menetries, 1836
Male adult
SE-Bulgaria:Burgas env. Tsarevo (Michurin), 10-15.VII.1981, J.Slaba leg.
L=22 mm
Copyright © Ruslan Panin
Calosoma (Campalita) auropunctatum auropunctatum Herbst, 1784: 131Calosoma (Campalita) auropunctatum dzungaricum Gebler, 1833: 274
Calosoma (Campalita) auropunctatum dzungaricum Gebler, 1833: 274
Male adult
W Kazakhstan, Emba riv., near Mijaly vill., 02.07.2009, P.Gorbunov leg.; 27 mm.
Copyright © Anton Menstschikov
Carabus (Aulonocarabus) gossarei gossarei Haury, 1879Carabus (Apoplesius) omphreodes Reitter, 1898Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
Male adult
Czech, Bohemia, Plzeň Reg., Domažlice Distr., Babylon, h=500 m, 30.VIII.2008, S.I. Budka leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus germarii Sturm, 1815
Carabus (Megodontus) violaceus germarii Sturm, 1815
Female adult
Croatia, Ivanščica Mts , Ivanec, 6-13.VI.2003, Krause Rene leg.
L=36 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
CE-Greece: Pieria, Stomio (Larissa) Ossa Mt. 7-14.VII.2009, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus banaticus Redtenbacher, 1849
Carabus (Procrustes) coriaceus banaticus Redtenbacher, 1849
Male adult
SW Romania, Banat, Băile Herculane env., Domogled valley, 24-30.V.2008, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin
Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male adult
NE Turkey, Trabzon, Zigana Pass, h~2000m, V. Slovak leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus graniger Palliardi, 1825
Carabus (Tachypus) cancellatus graniger Palliardi, 1825
Male adult
SW-Romania, Banatului Montan, Resita - Valiug, 29.V-9.VI.2005. Krause R. leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Female adult
NW Croatia, Bosanci, 1-12,VI.2004, Baros Slovak leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superbus Kraatz, 1878)
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superbus Kraatz, 1878)
Male+Female adult
SW-Romania,
Banat, Moldova Noua, Ravenska viil., 29.V-12.VI.2006, Krause Rene leg.
female,31mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) linnaei macairei Dejean, 1826
Carabus (Orinocarabus) linnaei macairei Dejean, 1826
Female adult
SW-Romania, Banat, Resita - Valiug, 29.V- 9.VI.2005, Krause R. , Baros, Pittner leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Male+Female adult
NW Spain,
E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
female, 27mm.
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
Female adult
E UKAINE, Lugansk city, Site Luganskoe, 20-30.VII.2010, V.A.Landik leg.
L=21 mm
det.Panin.R.
Copyright © Ruslan Panin
Carabus (Goniocarabus) sogdianus Semenov, 1898
Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
N Tajikistan, SW Pamir-Alay, Zarafshan Range, Fann Mts, Marguzor Lake circ., Padrut vill., 16.V.1997
L=21 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
Male adult
Russia, NW Altay, Barnaul env.,23.07.1998, D.Lomakin leg.

Copyright © Ruslan Panin
Cychrus caraboides Linnaeus, 1758
Cychrus caraboides Linnaeus, 1758
Male adult
L=17,0 mm
RUSSIA, Moscow city, East distr., National park «Losiniy Ostrov», spruce forest. 9.IX.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Male adult
Russia, SE Siberia, Transbaikalia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env.h~1800m., 1.VII.2006, S. Rybalkin leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Male adult
W. UKRAINE,
Transcarpathian reg., near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Female adult
N Tajikistan, SW Pamir-Alay, Fann Mts h=1800m, Padrut vill., 2-3.V.1997
L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri ronayi Csiki, 1906 (? scheidleri)
Carabus (Morphocarabus) scheidleri ronayi Csiki, 1906 (? scheidleri)
Male adult
W.UKRAINE,
Transcarpathian region,Mukachevo Distr., floodplain forest, 9.VII-25.VIII. 2011, Panin R. leg.
L=26 mm

Copyright © Ruslan Panin
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan: Niigata pref., Chayagahara Jyouetsu City,
18.XI.2006 A.Toba leg.
Copyright © Alexander Anichtchenko
Carabus (Eotribax) eous eous Morawitz, 1889
Carabus (Eotribax) eous eous Morawitz, 1889
Male adult
Kirghyzstan, N Terskei Ala Too, Borskaun riv., h=2500 m., 7-14.07.2000
male
L=17 mm
Copyright © Maksim Smirnov
Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill , 26.VII-3.VIII.2011, A. Korshunov leg.
L=29
Copyright © Ruslan Panin
Carabus (Megodontus) obovatus Fischer, 1827
Carabus (Megodontus) obovatus Fischer, 1827
Male adult
SW Siberia, Russia, Kemerevo reg., Krapivino distr., Azhendarovo, forest, 26-31.VII.2011, D. Efimov leg.
L=18 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis faldermanni Dejean, 1829
Carabus (Carabus) arvensis faldermanni Dejean, 1829
Male+Female adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill., 11.VII-1.VIII..2011, A. Korshunov leg.
female, 23 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens (as frivaldszkyi  Kraatz, 1887)
Carabus (Morphocarabus) excellens (as frivaldszkyi Kraatz, 1887)
Female adult
W-UKRAINE, NW Podolian,
Lviv Reg., Zolochiv Distr., near Gologirky vill.,steppe areas, 14-30.V.2008, Panin. R. leg./det.
L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens Fabricius, 1798
Carabus (Morphocarabus) excellens Fabricius, 1798
Male adult
NC Ukraine, Kyiv, Lysa Gora, on the fringes forest, 5-24.VI.2007,Gerasimov R.P.. leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens Fabricius, 1798
Carabus (Morphocarabus) excellens Fabricius, 1798
Male adult
C- UKRAINE, NW Odessa reg., Kodyma distr., near Aleksandrivka vill.,18.V.2003, Demidov G.N. leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nízke Tatry Mts., 4.VI.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Male adult
S Siberia, Burjatia reg., E Sayan Mts., Khugaituma mt., h=1700 m, 1-5. VII.2001
L=14,3 mm

Copyright © Maksim Smirnov
Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
Male adult
S Sakhalin, Susunaj Mt.R. 10.08 - 13.09.2003 D. Kochetkov leg.
L=14,5 mm
Copyright © Maksim Smirnov
Carabus (Deroplectes) arcanus arcanus Semenov, 1898Carabus (Morphocarabus) praecellens sp.
Carabus (Morphocarabus) praecellens sp.
Male adult
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus hopffgarteni (as emgei Ganglbauer, 1888)
Carabus (Procrustes) coriaceus hopffgarteni (as emgei Ganglbauer, 1888)
Male adult
CE-Greece: Larissa reg. Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
Copyright © Ruslan Panin
Carabus (Procerus) gigas gigas Creutzer, 1799
Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
Male adult
SE–France, Var departament, Foret royale -Saint-Paul-en-Foret vill.,18.VI.2002, Thelot J.P. leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Female adult
SE France, Alpes-Maritimes, Ascros, VI.2011
L=30 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) cribratus cribratus Quensel, 1806Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896Carabus (Platycarabus) depressus lucens Schaum, 1856
Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
SE-France: Alpi Marittime, Monte Viso, 14.VII.2010
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
Male adult
NW-Spain, León Province, Acebo, h=1000m, V.2007, Pascal Stefani leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Tachypus) auratus lotharingus Dejean, 1826
Carabus (Tachypus) auratus lotharingus Dejean, 1826
Male adult
SC France: Herault departament, Foret de Saint Gervais, V. 2009 leg. J. Nicolas
L=27mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus nagyagensis Seidlitz, 1888
Carabus (Eucarabus) obsoletus nagyagensis Seidlitz, 1888
Female adult
NW-Romania, Mts Ses (Plopis),Cluj Reg., Bucea env, h=580 m, 8.VI.2011, Sl.Budka leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
C Romania, Târgu Mureş, Ernei, 46°35′471″N 24°37’351″E, h=300 m,20.VIII.2004, Sl.Budka leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
Male+Female adult
NW Romania, Zalău env., 31.VII.2003, Sl.Budka leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nizke Tatry Mts., h=1750 m,.9.VI.2007, V.Slovak leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) aumonti maroccanus Bedel, 1895
Carabus (Macrothorax) aumonti maroccanus Bedel, 1895
Male adult
N Morocco (Spain:Melilla), hipodromo, X.1969, leg. Pardo Alcaide
L=38 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Male adult
E Ukraine, Lugannsk Reg., Sverdlovsk distr., Steppe Reserve “Provalsky", 3.V-3.VI.2011, leg.Glotov S./det.Panin R.
female, 22 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Female adult
C Hungary, Pest reg., Örkény, 9-10.V.1997, S.Budka leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Muntii Metaliferi, Hunedoara reg., near Sacaramb vill., 6.VI.2011, S.Budka leg.
L=37 mm
Copyright © Ruslan Panin
Carabus (Megodontus) vietinghoffi bowringi (as  lazoensis Obydov, 1999)
Carabus (Megodontus) vietinghoffi bowringi (as lazoensis Obydov, 1999)
Male+Female adult
Russia, Far East, Primorsky reg., S. Sichote-Alin Mts.R., Kavalerovo vill. env.,
5-25.VII.1992, S. Kunitzyn leg.
male - 28 mm, female - 30 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male+Female adult
Russia, Far East, Primorsky reg., Khasansky distr.,
Zanadvorovka vill., Gusevsky pit.
M - 19-21.VI.2007, body lenght 29 mm.
F - 15.VII.2008, body lenght 30 mm.
S. Veriga leg.
Copyright © Alexandr Safronov
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male+Female adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
Body lenght 30-33 mm.
Copyright © Alexandr Safronov
Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003
Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003 Paratype
Male+Female adult
W-Tianshan:Karatau Mts.,N-part:Boialdyr River valley,
N of Ken-su, 600-750m, 22-V-2000 М. Danilevsky leg.
male - 20 мм, female - 19 мм.
Copyright © Alexandr Safronov
Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Male+Female adult
Kazakhstan, S. Karatau Mts. R., Mashat vill. env.
23-IV-1993 М. Danilevsky leg.
male - 21 мм, female - 22 мм.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Male+Female adult
Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
5-25.VII.1992, С. Kunitzyn leg.
male - 25 mm, female - 25 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Male+Female adult
Russia, Far East, Primorsky ter., Khasansky distr., Zanadvorovka vill.,
Gusevsky pit, S.Veriga leg.
male - 13.VII.2008, female 19-21.VI.2007
male - 28 мм, female - 28 мм.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male+Female adult
Russia, Far East, Khabarovsk reg., Obluchye env.
M. Khingan Mts. R.
10-20.VII.1992 S. Andreev leg.
male - 26 mm, female - 27 mm.
Copyright © Alexandr Safronov
Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male+Female adult
Kyrgyzstan, Kyrgyzskyi Mts. R., Uzun Bulak Gorge, h=1800 ms.
01-09.VI.2002 loc. coll.
male - 28 mm, female - 33 mm.
Copyright © Alexandr Safronov
Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992Carabus (Cupreocarabus) brezinai Deuve, 1994Carabus (Ctenocarabus) galicianus Gory, 1839
Carabus (Ctenocarabus) galicianus Gory, 1839
Male adult
Spain,
Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
L= 20 mm
Copyright © Alexander Anichtchenko
Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)Carabus (Cryptocarabus) subparallelus Ballion, 1878
Carabus (Cryptocarabus) subparallelus Ballion, 1878
Female adult
Kazakhstan,
near Alma-Ata, V.2009, Pak O.leg.
female, 17mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) adamsi Adams, 1817
Carabus (Sphodristocarabus) adamsi Adams, 1817
Male+Female adult
Russia, N. Caucasus, Stavropol reg., Zheleznovodsk,
Zheleznaya Mt., h=600 m.
5-11.VI.2011, Е. Shankhiza leg.
male - 24 mm, female - 25 mm.
Copyright © Alexandr Safronov
Carabus (Sphodristocarabus) varians Fischer, 1827
Carabus (Sphodristocarabus) varians Fischer, 1827
Male+Female adult
N. Armenia, Idzhevan distr., Acharkut vill. env., h=800 m.
26-28.VII.2008 S. Andreev leg.
male - 22 mm, female - 23 mm.
Copyright © Alexandr Safronov
Carabus (Cryptocarabus) lindemanni Ballion, 1878
Carabus (Cryptocarabus) lindemanni Ballion, 1878
Female adult
SE Kazakhstan, Alma-Ata env., 16.04. 2000
L=16,0 mm
Copyright © Maksim Smirnov
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
SE Kazakhstan, N-Tianshan:N-slopes Ketmen Mt.Rng. h=2250-2500m., near Ketmen vill.,1-5.VI.1990, Dolin V.G. leg.
L=13.5 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Male+Female adult
NE-Kazakhstan, W Altay, Ivanovsky Mt.Rng, near Ridder (Leninogorsk), h-1300-1500m, 4-6.VII.2003
female, 25 mm
det.Panin R.
Copyright © Ruslan Panin
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
male - 30 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831 (as cyanescens Kraatz, 1882, nec Sturm, 1815)
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831 (as cyanescens Kraatz, 1882, nec Sturm, 1815)
Male+Female adult
Kazakhstan, E. Kazakhstan area,
Zyryanovsk env., near Putintzevo vill.
10-VI-2005 M. Danilevsky leg.
male - 24 mm, female - 25 mm.
Copyright © Alexandr Safronov
Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
Male+Female adult
Kyrgyzstan, NE Tian-Shan, E. Terskey Alatau Mts. R.,
Kok-kia River, h=2800 м.
7-8.VI.1997 М. Danilevsky leg.
Body lenght 15-18 mm.
Copyright © Alexandr Safronov
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male+Female adult
Kyrgyzstan, NC Tianshan, Kyrgyz Mt. R., Tuyuk riv., h=2000 м.
25-VI-1997 М. Danilevsky leg.
male - 20 mm, female - 21 mm.
Copyright © Alexandr Safronov
Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 23 mm, female - 24 mm.
Copyright © Alexandr Safronov
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male+Female adult
W. Caucasus, Abkhasia, 8 km NE of Ritza Lake,
Avadkhara valley, h=1700 m
10-VI-2009 A.Safronov leg.
Copyright © Alexandr Safronov
Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 28 mm, female - 31 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male+Female adult
SW Siberia, Kemerovo reg., Krapivino distr.,
8 km SSW Saltymakovo vill., h=150 m.
1-20.VII.2010 D. Efimov leg.
male - 25 mm, female - 26 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male+Female adult
Russia, S. Siberia, Krasnoyarsk reg., W. Sayan,
Sayanogorsk distr., Cheremushki vill. env.,
07-VII-1994 М. Danilevsky leg.
male - 22 mm, female - 25 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) danilevskii Obydov, 1993
Carabus (Megodontus) danilevskii Obydov, 1993
Male+Female adult
Russia, SW Siberia, Krasnoyarsk reg., W. Sayan, near Cheryomushki.
26-29.VII.1996 loc. coll.
male - 18mm, female - 20 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) kantaikensis Gehin, 1885
Carabus (Megodontus) kantaikensis Gehin, 1885
Female adult
Russia, SW Siberia, W. Sayan, Krasnoyarsk reg.,
S of Abakan, near Cheryomushki, Dzhoysky Pass.
07-VII-1994 M. Danilevsky leg.
body lenght 18 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) obovatus Fischer, 1827
Carabus (Megodontus) obovatus Fischer, 1827
Male adult
SW Siberia, Kemerovo reg., Krapivino distr.,
8 km SSW Saltymakovo vill., h=150 m.
20-30.VI.2010 D. Efimov, A. Korshunov leg.
body lenght 18 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) imperialis Fischer, 1823
Carabus (Megodontus) imperialis Fischer, 1823
Male adult
NE Kazakhstan, Zyrjanovsk distr., Putincevo vill. env.
25-VI-2005 M. Danilevsky leg.
male 24 mm, female 26 mm.
Copyright © Safronov
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
CZECH, SE Moravia, Velka nad Velickou. 20-V-2005, V. Slovak leg. male - 26 mm, female - 30 mm.
Copyright © Alexandr Safronov
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male+Female adult
Male - SW Romania, Banat - Bocsa/Coltan,
21.V-05.VI.2010, R. Krause leg., L=29 mm.
Female - SW Romania, Banat - Saska Montana, Stinapari,
25-VI-2010, R. Krause leg., L=31 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) violaceus violaceus Linne, 1758 (as intermarinus Blumenthal & Staven, 1985)
Carabus (Megodontus) violaceus violaceus Linne, 1758 (as intermarinus Blumenthal & Staven, 1985)
Male+Female adult
Poland, Gdansk distr., Gdynya env., Reservat «Cisowa»,
5-9.08.1998 I. Solodovnikov leg.
female - 26 mm.
Copyright © Alexandr Safronov
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860 (as ursuius Lapouge, 1911)
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860 (as ursuius Lapouge, 1911)
Male+Female adult
FRANCE (64), Ursuya Mt.
XII-2009, P. Stefani leg.
male - 30 mm, female - 32 mm.
Copyright © Alexandr Safronov
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Anyos (ANDORRE), h=1450ms,
5-VI-2010, P. Stefani leg.
body lenght - 32 mm.
Copyright © Alexandr Safronov
Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
Male adult
W. Caucasus, Abkhazia, Gudauta distr.,
Duripsh vill. env., Khipsta River, h=700 m,
11-06-2009 A. Safronov leg.
male - 40 mm
Copyright © Alexandr Safronov
Carabus (Neoplectes) polychrous polychrous Rost, 1892
Carabus (Neoplectes) polychrous polychrous Rost, 1892
Female adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
8-11.06.2009 A. Safronov leg.
body lenght 40-43 mm.
Copyright © Alexandr Safronov
Carabus (Sphodristocarabus) janthinus janthinus s.str. (as dvorshaki Mandl, 1975)
Carabus (Sphodristocarabus) janthinus janthinus s.str. (as dvorshaki Mandl, 1975)
Male+Female adult
W. Caucasus, Abkhazia, Gudauta distr..,
Duripsh vill. env., Khipsta riv., h=400-500 м.
03-13.VI.2009 A. Safronov leg.
body lenght 20-25 мм.
Copyright © Alexandr Safronov
Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886Calosoma (Callistenia) luxatus Say, 1823
Calosoma (Callistenia) luxatus Say, 1823
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Calosoma (Calopachys) omiltemium Bates, 1891: 226
Calosoma (Calopachys) omiltemium Bates, 1891: 226
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Calosoma (Blaptosoma) viridisulcatum viridisulcatum Chaudoir, 1863
Calosoma (Blaptosoma) viridisulcatum viridisulcatum Chaudoir, 1863
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Calosoma (Callitropa) externum Say, 1823: 149
Calosoma (Callitropa) externum Say, 1823: 149
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Calosoma (Callistenia) wilkesi Le Conte, 1851: 200
Calosoma (Callistenia) wilkesi Le Conte, 1851: 200
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Calosoma (Carabomimus) laevigatum laevigatum Chaudoir, 1869a: 374
Calosoma (Carabomimus) laevigatum laevigatum Chaudoir, 1869a: 374
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Calosoma (Carabosoma) eremicola Fall, 1910: 91
Calosoma (Carabosoma) eremicola Fall, 1910: 91
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Carabus (Apotomopterus) dechambreianus dechambreianus Deuve & Li, 2000
Carabus (Apotomopterus) dechambreianus dechambreianus Deuve & Li, 2000
Male+Female adult
Gailin, Guanxi prov., CHINA,
17-V-2011 loc. coll.
male - 36 mm, female - 38 mm.
Copyright © Alexandr Safronov
Carabus (Apotomopterus) shun floridus Cavazzuti & Ratti, 1998
Carabus (Apotomopterus) shun floridus Cavazzuti & Ratti, 1998
Male+Female adult
Mts. Nanling Reserve, Guangdong prov., CHINA
1-7.VII.2011 loc. coll.
male - 38 mm, female - 40 mm.
Copyright © Alexandr Safronov
Carabus (Neoplectes) reitteri gagrinus Starck, 1894 (as gegicus Gottwald, 1985)
Carabus (Neoplectes) reitteri gagrinus Starck, 1894 (as gegicus Gottwald, 1985)
Male+Female adult
W.Caucasus, Abkhasia, Gagra distr., Gega riv. valley,
13-15.07.2005 D. Kasatkin. leg.
male - 30 mm, female - 34 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) rothi hampei as telekii Csiki, 1937 (? hampei )
Carabus (Morphocarabus) rothi hampei as telekii Csiki, 1937 (? hampei )
Female adult
CW Romania, Arad distr., Zarandulu Mt Range, near Ilteu vill., h=276 m, 46°66′487″N 22°20’455″E, 25.VI.2005, Sl.Budka leg.
L=30 mm

Copyright © Ruslan Panin
Carabus (Cratocephalus) solskyi Ballion, 1878Carabus (Cratocechenus) jacobsoni Semenov, 1898
Carabus (Cratocechenus) jacobsoni Semenov, 1898
Male adult
S Kazakhstan, Almaty reg., Big Lake, h=2800 m, 25.VI.96
male
L=14 mm
Copyright © Maksim Smirnov
Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis )Carabus (Cratocechenus) akinini akinini Morawitz, 1886Carabus (Cratocarabus) puer Morawitz, 1886Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
Male adult
Russia, Far East, South Primoriye, Kaymanovka village, 10-20.07.2000; P.Gorbunov leg.
Copyright © Anton Menstschikov
Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863Carabus (Coptolabrus) smaragdinus longipennis (as mucdenensis Hauser, 1921)
Carabus (Coptolabrus) smaragdinus longipennis (as mucdenensis Hauser, 1921)
Male adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Alexandr Safronov
Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
Male+Female adult
Hua-Bo-zi-Shan, Huanren County,
Liaoning Province, CHINE
20-30.VI.2008
Copyright © Alexandr Safronov
Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Female adult
Chine, C.Sichuan, Ya'an, 07.2005 Local people
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) pustulifer guerryi Born, 1903
Carabus (Coptolabrus) pustulifer guerryi Born, 1903
Female adult
N Yunnan: Gaoligongshan, h =1300 m, 06.2003 Local people
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) pustulifer cyanelongatus Deuve, 2006
Carabus (Coptolabrus) pustulifer cyanelongatus Deuve, 2006
Female adult
Zhagyab, Quamdu City, Tibet
8-15.VI.2008
Copyright © Alexandr Safronov
Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
Male adult
CW Spain, Cáceres, San Martín de Trevejo,24.VI.2011, A. Diez leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Male adult
SC Spain, Jaen reg., Puente de la Sierra, 10.VII.2011, M.Lopez leg.
L=28 mm
Copyright © Ruslan Panin
Scaphinotus (Scaphinotus) unicolor Fabricius, 1787
Scaphinotus (Scaphinotus) unicolor Fabricius, 1787
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Scaphinotus (Brennus) interruptus Menetries, 1844
Scaphinotus (Brennus) interruptus Menetries, 1844
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Scaphinotus (Maronetus) hubbardi Schwarz, 1895
Scaphinotus (Maronetus) hubbardi Schwarz, 1895
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Scaphinotus (Neocychrus) angulatus Harris, 1839
Scaphinotus (Neocychrus) angulatus Harris, 1839
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Scaphinotus (Nomaretus) cavicollis LeConte, 1859
Scaphinotus (Nomaretus) cavicollis LeConte, 1859
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Scaphinotus (Scaphinotus) viduus Dejean, 1826
Scaphinotus (Scaphinotus) viduus Dejean, 1826
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Scaphinotus (Scaphinotus) petersi petersi s.str.
Scaphinotus (Scaphinotus) petersi petersi s.str.
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Scaphinotus (Scaphinotus) petersi kathleenae Ball, 1966
Scaphinotus (Scaphinotus) petersi kathleenae Ball, 1966
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Scaphinotus (Scaphinotus) petersi grahami Vandyke, 1938
Scaphinotus (Scaphinotus) petersi grahami Vandyke, 1938
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Carabus (Morphocarabus) rothi rothi Dejean, 1829
Carabus (Morphocarabus) rothi rothi Dejean, 1829
Male+Female adult
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m, 19.VIII.2001, S.Budka leg.
L=28 mm
Copyright © Ruslan Panin
Scaphinotus (Steniridia) andrewsii waldensius (Valentine, 1935)
Scaphinotus (Steniridia) andrewsii waldensius (Valentine, 1935)
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Ceroglossus suturalis suturalis Fabricius, 1775
Ceroglossus suturalis suturalis Fabricius, 1775
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
Russia, Far East, Primorskiy reg., Ussuriysk Distr., Kaymanovka vill., 6-15.VIII.2011,
A. Korshunov leg.
L= 21.5 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, forest-steppe,1.VII-20.VIII,2011,M.Ivanov leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Female adult
NE-Turkey: Rize Prov.Ovit Pass (40.37;40.46) h-2700m,15.VII.2008, I.Rapuzzi & Caldon leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, on dry steppe slopes, 7.VIII,2011, M.Ivanov leg.
L=21 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) latus latus Dejean, 1826
Carabus (Mesocarabus) latus latus Dejean, 1826
Female adult
CW Spain, Cáceres, San Martín de Trevejo, 24.VI.2011, A. Diez leg./det.Panin R.
L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Male adult
SC Spain, Jaen reg., Monte Javalcuz h=1436 m, 6.X.2010, M.A. Lopez leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Female adult
S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
L=28 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Male adult
NC-Spain, Soria, Puerto de Oncala, 2.VI.2006
L=24 mm
Copyright © Ruslan Panin
Cychrus attenuatus attenuatus Fabricius, 1792
Cychrus attenuatus attenuatus Fabricius, 1792
Male+Female adult
W.UKRAINE, NE Carpathians,
Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt.Range h=700m, 26.V -1.VII.2010, Panin R. leg.

Copyright © Ruslan Panin
Carabus (Coptolabrus) principalis Bates, 1889
Carabus (Coptolabrus) principalis Bates, 1889
Male+Female adult
Zi-Ding-Shan, h=1406 m, Guiyang County, Binzhou City, Hunan, CHINE
19-29.IV.2008
Copyright © Alexandr Safronov
Carabus (Carabus) stscheglowi Mannerheim, 1827Carabus (Coptolabrus) lafossei tiantai Kleinfeld, 1997Carabus (Coptolabrus) lafossei pseudocoelestis Kleinfeld, 1999Carabus (Coptolabrus) jankowskii jankowskii Oberthur, 1883
Carabus (Coptolabrus) jankowskii jankowskii Oberthur, 1883
Male adult
N.Korea, Chagang Do, Mt. Langlin, 30.07.2005
Copyright © Anton Menstschikov
Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Female adult
CHINA, Jilin prov., Mt. Longgang., 15.07.2005
Copyright © Anton Menstschikov
Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Male adult
NE-China,
Liaoning Prov., Donggou, Mt. Guanmen, 15.VI.2009
male, 36mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus grumorum Semenov, 1898Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898 (as rostandi Deuve, 1991)
Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898 (as rostandi Deuve, 1991)
Male adult
China, N Sichuan province, Venchuan, h=1400 m., 30.VI.2007, A. Gorodinski leg.;
L=33 mm
Copyright © Anton Menstschikov
Carabus (Coptolabrus) elysii pulcher Kleinfeld, 1997
Carabus (Coptolabrus) elysii pulcher Kleinfeld, 1997 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld
Carabus (Coptolabrus) elysii magnificens Kleinfeld, 1999
Carabus (Coptolabrus) elysii magnificens Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld
Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai Kleinfeld, 2000)Carabus (Coptolabrus) elysii elysii Thomson, 1856
Carabus (Coptolabrus) elysii elysii Thomson, 1856
Female adult
China, Sichuan province, Wanyuan; 15.06.2009; Local people leg.;
L=32 mm.
Copyright © Anton Menstschikov
Carabus (Coptolabrus) augustus pustuliferus Kleinfeld, 1998Carabus (Coptolabrus) augustus cristianofonti Deuve & Font, 1998
Carabus (Coptolabrus) augustus cristianofonti Deuve & Font, 1998
Female adult
China,
Guizhou Prov., Sinan, 14.V.2009
L= 41mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey: Artvin:Hopa, Borçka env., h=750 m, 9.VI-7.VII.2011, M.Janalik leg.
female, 42 mm
Copyright © Ruslan Panin
Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Male adult
NE-Turkey: Artvin:Hopa, Borçka env., h=450 m, 29.VI-7.VII.2011, M.Janalik leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Male+Female adult
C Bulgaria:Stara planina Mts, Gabrovo,12-15.VI.2007, M. Janalik leg. female, 31 mm
Copyright © Ruslan Panin
Carabus (Tribax) puschkini ponticus Deyrolle, 1869
Carabus (Tribax) puschkini ponticus Deyrolle, 1869
Male adult
NE-Turkey:Trabzon, Zigana Pass h=1700m,14-24.VI.2010, M.Janalik leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Male adult
SW Romania, Tarcu Mts, vf. Nedeia h=2000m, 12.VI.2008, J.Kynkor leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis carpathus (as pseudonoricus Kenyery, 1983)
Carabus (Carabus) arvensis carpathus (as pseudonoricus Kenyery, 1983)
Male adult
Slovenia: Julske Alpy, Podvelka, h=450m, 24.X.2010, M.Janalik leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
Male adult
NE-Turke,Rize vill., S Ayder h=1740 m,28.VI-6.VII.2011, M.Janalik leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) schrencki dongningensis Deuve & Li, 2005
Carabus (Acoptolabrus) schrencki dongningensis Deuve & Li, 2005
Male adult
NE China, Heilongjiang prov., Dongning, V.2009
L=25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Male adult
N Romania, E Carpathian, Maramureş Reg., Gutâi Mts, cabana Mogosa,4 km N Baia Sprie, 9-17.VI.2010, J.Kynkor leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus nagyagensis (as paranagyagensis Lie, 1990)
Carabus (Eucarabus) obsoletus nagyagensis (as paranagyagensis Lie, 1990)
Male adult
W Romania, Arad reg., Zarand Mts (Muntii Zarandului), Nadas-Barzavah h=142 m, J. Kynkor leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Male adult
NE-Turkey:Trabzon env., Sumela, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Male adult
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=26 mm
Copyright © Ruslan Panin
Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
Male adult
NE-China: Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=27 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as feichtauensis Müller & Sach, 1999)
Carabus (Platycarabus) fabricii fabricii (as feichtauensis Müller & Sach, 1999)
Male adult
NC Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
L=18 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as koralpicus Sokolar, 1910)
Carabus (Platycarabus) fabricii fabricii (as koralpicus Sokolar, 1910)
Female adult
S Austria, Carinthia reg.(Kärnten), Wolfsberg, Koralpe Mt Range,18.VI.2005, Manfred Egger leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
Male adult
Cyprus, Troodos, 17.2.1V.2010,. Manfred Egger leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Female adult
Greece:, N Peloponnes, Xylokastro, Zemeno,h=1000m, 6.VI.1997, Manfred Egger leg.
L=21 mm
Copyright © Ruslan Panin
Carabus (Procrustes) banoni Dejean, 1829
Carabus (Procrustes) banoni Dejean, 1829
Male adult
Greece,W Crete Island, Mili gorge ,12.V.2009, Manfred Egger leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Male adult
N Morocco, El Biutz –Taghramt,20.I.2005, Manfred Egger leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris hoppei Germar, 1824
Carabus (Orinocarabus) alpestris hoppei Germar, 1824
Male adult
W Austria, Tirol, Innsbruck, Hafelekar Mts h=2200m, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) linnaei prope linnaei Panzer, 1812 (aberrant)
Carabus (Orinocarabus) linnaei prope linnaei Panzer, 1812 (aberrant)
Male+Female adult
W Ukraine, NE Carpathians Mts.,Transcarpathian Reg., Rakhiv Distr., Chornogora Mt. Range, Mt Gutyn-Tomnatyk h=2015m, alpin zone, 8.VI-27.VII.2011, Panin R. leg. female, 17 mm; male, 14 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris alpestris Sturm, 1815
Carabus (Orinocarabus) alpestris alpestris Sturm, 1815
Female adult
C Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
Male adult
W-Ukraine, NW podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, 17.II.2007,Panin R. leg.
female, 21 mm
Copyright © Ruslan Panin
Carabus (Megodontus) croaticus croaticus Dejean, 1826
Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
SW Slovenia, Ilirska Bistrica, Sviščaki, Snežnik,17-28.VI.2009, Manfred Egger leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus schreiberi Kraatz, 1887
Carabus (Megodontus) caelatus schreiberi Kraatz, 1887
Male adult
SW Slovenia, Obrov, Poljane, 14-27.VIII.2010, Manfed Egger leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Male adult
NE-Italy, Bolzano, Brenner, Zirog h=2000 m, 26.VII.1992, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as poloniensis Géhin, 1876)
Carabus (Platycarabus) fabricii fabricii (as poloniensis Géhin, 1876)
Female adult
N-Slovakia, Orava Beskids Mts, Mt Babia Góra h=1720 m, 5.VII.20.VIII.2010, Rudolf Kmeco leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Male adult
S-Greece:Peloponnesos: Taygetos Mt.Rng., 17-23.V.2005, V. Sima leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt., 6-17.VII.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus mopsucrenae Peyron, 1858
Carabus (Procrustes) coriaceus mopsucrenae Peyron, 1858
Male adult
SC-Turkey, Adana Prov., Doganbeyli, 8.V.2001, Snizhek M. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Male adult
W Croatia, Ivanščica Mts , Ivanec, 24.VI-7.VII.2006, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi ulrichi (as intercessor Sokolar, 1912)
Carabus (Eucarabus) ulrichi ulrichi (as intercessor Sokolar, 1912)
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Male+Female adult
C Romania, Brașov County, Cristian vill., h=380 m,V-VI.2008, M.Pasek leg.
female, 27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi incompsus (as spectabilis Csiki, 1906)
Carabus (Morphocarabus) rothi incompsus (as spectabilis Csiki, 1906)
Female adult
NC Romania, Bistrita Nasaud reg., Călimani Mts, Colibita vill., mixed forest, meadows h=800m, 15-21.V.2011, M.Pasek leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi incompsus (as mendax Csiki, 1906)
Carabus (Morphocarabus) rothi incompsus (as mendax Csiki, 1906)
Female adult
W Ukraine, NE Carpathians,
Transcarpathian reg., Chornogora Mt Range, h=1100m,13.VI.2011, Panin R. leg./det.
L=33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male+Female adult
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts., VII.2007, I. Marvan leg.
L=29 mm
Copyright © Ruslan Panin
Cychrus semigranosus balcanicus Hopffgarten, 1881
Cychrus semigranosus balcanicus Hopffgarten, 1881
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg.
L=21 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Male adult
SW Romania, S Carpathians,
Banat , Tarcu Mts, Borlova env., 19-31.V.2005, P. Sima leg.
L=24 мм
Copyright © Ruslan Panin
Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Male adult
NE-Turkey:Trabzon, Zigana Pass h=2000m,14-24.VI.2010, V. Slovák leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE-Serbia:Majdanpek, 9-11,V.2010, L.Brhel leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Male adult
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan, 20-29.VI.2009, L. Bieber leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) armeniacus Mannerheim, 1830
Carabus (Sphodristocarabus) armeniacus Mannerheim, 1830
Male adult
NE Turkey, Artvin Prov., Şavşat, Çamlıca,18.VI.1993, M. Janata leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Male adult
Czech, S Moravia, Pouzdrany, 8,VIII.2005, V. Slovak lgt.
L=27 mm
Copyright © Ruslan Panin
Carabus (Megodontus) planicollis verae Csiki, 1905
Carabus (Megodontus) planicollis verae Csiki, 1905
Male+Female adult
♂ C-Romania:S Carpathians, Fagaraş Mts, near Avrig Lake, h=2100 m, 5.VI.2005
♀ C-Romania:S Carpathians, Fagaraş Mts, Saratii-Valley h=1700 m, 6.VI.2005
female, 28 мм
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Male adult
NE-Bulgaria, Dobrich Region, Kavarna env.
L=31 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815 (as prunneri Mallasz, 1901)
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815 (as prunneri Mallasz, 1901)
Male adult
NW Romania, Apuseni Mts, Trascău Mt Rng, Lungesti vill., 25 km W Turda,.VI.2012
L=26 mm
Copyright © Ruslan Panin
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m,Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) stroganowi transfuga Semenov, 1898
Carabus (Megodontus) stroganowi transfuga Semenov, 1898
Male adult
SE-Azerbaijan: Talysh, Kalvaz, 21-30.VI., Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Male adult
S-Kazakhstan:Ili River valley,:near Bakanas,3-5.V.2002
L=21 mm
Copyright © Ruslan Panin
Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
Female adult
Azerbaijan, Talysh Mts., Zuvand, Kalvaz vill., h=1650 m, 15.VII.2003, Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.,12.VII.2006, Pak O. leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Male adult
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m, 10.V.1996, Dolin V.G. leg.
det. Panin R.Yu.
L=22 mm
Copyright © Ruslan Panin
Calosoma (Charmosta) investigator Illiger, 1798Calosoma (Calosoma) inquisitor inquisitor Linne, 1758: 414Calosoma (Callisthenes) panderi Fischer, 1822Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Male adult
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,. 6.VIII.2008
L=27 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Female adult
Tadzhikistan, Pamir, N-slope Peter I.Mt Rng,h=2100m,Ganishou vill., Dolin V.G. leg.
det.Panin R.Yu.
L=24 mm
Copyright © Ruslan Panin
Carabus (Procerus) scabrosus caucasicus Adams, 1817
Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright © Ruslan Panin
Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
Male adult
S-Greece: Peloponnes, Mt.Chelmos, 20.V.2007.
L=46 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Female adult
Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
L=19 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 186
Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 186
Male adult
CW Spain, Province Huelva, Arroyomolinos de León,30.X.2011, T. Alcantara leg.; L=23 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians,
Ivano-Frankivsk reg.,Yaremcha distr., Gorgany Mts., h=1590 m, 22.VI-3.VIII.08, Panin R. leg.
male, 23mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians,
Ivano-Frankivsk reg., Gorgany Mts., h=1590 m, 17.V-26.VI.2011, Panin R. leg.
male, 23mm
Copyright © Ruslan Panin
Carabus (Megodontus) kolbei kolbei Roeschke, 1887
Carabus (Megodontus) kolbei kolbei Roeschke, 1887
Male adult
N-Japan, C-Hokkaido, Hidaka distr., Niikappu env., 27.VIII.1989, Sasaki leg,
L=27 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
Male adult
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan,VI.2007, Hao leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Male adult
N-Japan, C-Hokkaido, Hidaka Mt.Rng, Mt.Apoidak near Samani town, 20.VIII.1982,T. Akaishi leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
Male adult
NE-Korea:North Hamgyong Prov., Sechon-dong, VII.2004, Li leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini lucasi Gaubil, 1849
Carabus (Eurycarabus) famini lucasi Gaubil, 1849
Male adult
N-Tunisia, Bizerte reg., Menzel Bourguiba city,h=5 m, XII.1981, S. Remond leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m,VII.2001, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (?Oreocarabus) titanus Breuning, 1933
Carabus (?Oreocarabus) titanus Breuning, 1933
Male adult
C-China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Male adult
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m, IV.1999, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) kumagai kumagai Kimura & Komiya, 1974
Carabus (Leptocarabus) kumagai kumagai Kimura & Komiya, 1974
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Male adult
SC-USA, New Mexico, Santa Fe,VIII.1999, Sehnal R. leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.
L=21 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
20-25.VII.2010. A.Dergalev leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Female adult
W-UKRAINE, Ivano-Frankivsk reg., Gorgany Mts, Syvula Mt., h =1820 м.,
25.VII.2011,Panin R. leg.
L=27 mm

Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W-Ukraine, Roztochia,
Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, 17.IV-3.V.2010, Panin R. leg.
L=20 mm

Copyright © Ruslan Panin
Carabus (Procrustes) piochardi ? piochardi Gehin, 1883
Carabus (Procrustes) piochardi ? piochardi Gehin, 1883
Male adult
C-Lebanon, Faraya vill., Mt. Djebel Sannin h=1800m, X.2012, C. Reuter leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
Female adult
SE-Iceland:Höfn env.:Hornafjördur, 24-28.VIII.2008, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Male adult
C-Chine, SE-Shaanxi Province, Shahezi, Shangzhou City, 6-18.VII.2010
L=43 mm
Copyright © Ruslan Panin
Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003)
Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003) Paratype
Female adult
C-China, S-Gansu: SSW of Minxian: Qilian Shan: W Kunda, 34:10N/103:47E, h=2800-3000m, forest, 7.VII.2002, Belousov & Kabak leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
China, NC-Sichuan (ZOIGE), LANGMUSI (=DOGCANGLHAMO), 34:03:48N/102:38:08E, h=4100m, alpine meadows, screes, 16-18.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus delerei Mandl, 1955
Carabus (Ulocarabus) theanus delerei Mandl, 1955
Male adult
N-Afghanistan, Hindukush Mt.Rng, NE Salang Pass h=2300 m, 35°27'494"N, 68°58'399"E, IV.2011, C. Reuter leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus theanus Reitter, 1895
Carabus (Ulocarabus) theanus theanus Reitter, 1895
Male adult
Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo, Haidar gorge,1.VI.1984, Dolin V.G. leg./det.Panin R.Yu.
L=21 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus)? theanus theanus Reitter, 1895
Carabus (Ulocarabus)? theanus theanus Reitter, 1895
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) canaliculatus hailinensis Deuve & Li, 2000
Carabus (Aulonocarabus) canaliculatus hailinensis Deuve & Li, 2000
Male adult
NE China, Heilongjiang Prov., Mudanjiang City, Mt.Tai-Gu-Ding-zi h=800m, 15-25.VI.2010
L=29 mm
Copyright © Ruslan Panin
Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
Male adult
CW-Serbia, Zlatar Mts, Nova Varoš, Zlatar Lake env., 1-12.VII.2007, P.Sima leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
Female adult
N-Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, 29.X.2012, C. Reuter leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) kadoudali (Morvan, 1981)
Carabus (Meganebrius) kadoudali (Morvan, 1981)
Male+Female adult
C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
female, 20 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 (ab.)
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 (ab.)
Female adult
C Bulgaria: Stara Planina Mts, Schipka pass h=1450m, 12-15.VI.2007, M.Janalik leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
N- Romania, E-Carpathians, Rodna Mts, h=2000m, 7-8.VI.2012, Cartuz Z.leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) manfredeggeri Müller 2012
Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
Male+Female adult
N-Korea, Chagang Prov., Manpo City, Songhak-ri, 2-16.VII.2011
female, 21 mm
male, 19 mm
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus dalmatinus (as procerus Reitter, 1885)
Carabus (Megodontus) caelatus dalmatinus (as procerus Reitter, 1885)
Male adult
S Bosnia, Potoci env. h=500 m, VII.2006, Bartoloni leg.
L=38 mm
Copyright © Ruslan Panin
Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Male adult
N India.Kashmir, Loran, W part Pir Panchal range,h=2400-2500m, 33°49'"N, 74°20'E
L=25 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus dachangensis Lu & Müller, 2009
Carabus (Coptolabrus) smaragdinus dachangensis Lu & Müller, 2009
Female adult
NE-China, Liaoning Prov., Changhai County, Dachangshan Dao Isl.,12.VI.2011
L=40 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
Male adult
N Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, C. Reuter leg.
L=17.5 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Male adult
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m, 1-12.VII.2007, Pr.Sima leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
C-China, E Tibet, N-Sichuan, Hongyuan,valley 10 km SE Sanggarpar, h=4200m, 32*18’ N; 102*35’E, alpine meadow / scree, 19.VII.1995, B.Brezina leg.
L=21 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Male adult
N-Korea, South Pyongan prov., Mt. Maeng-san, 6.VI.2010
L=25 mm
Copyright © Ruslan Panin
Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
Female adult
China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr. Nan Tian Men h=3200m,
33°55'17"N/ 107°46'53"E, 25-30.VII.2011, Li Wuyang leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus dryas Gistl, 1857
Carabus (Megodontus) violaceus dryas Gistl, 1857
Male adult
NE-Greece, Mt. Olympus, Litochoro env., 20-30.V.2005, Sima leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Male adult
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m.V.2012, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
Male adult
N India, Kashmir, Pahalgam, Aru Valley h=2400m, VII.2007
L=25 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (?Oreocarabus) qinlingensis Imura, 1993
Carabus (?Oreocarabus) qinlingensis Imura, 1993
Male adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E, 25-30.VII.2011, Li Wuyang leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill., 45°97'472"N/23°02'630"E, h=680 m, mixed forest, meadows, 15-20.V.2011, B.Brezina leg.
L=36 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Female adult
E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
L=23 mm

Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ( as jasilkowskii Born,1905)
Carabus (Morphocarabus) excellens ( as jasilkowskii Born,1905)
Male adult
NE Romania, 5 km W Suceava, 47°63′940″N 26°18′795″E,h=340m, agricultural field, 15-20.V.2011, B. Brezina leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) arunensis Heinertz, 1980
Carabus (Meganebrius) arunensis Heinertz, 1980
Male adult
E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
L=27 mm

Copyright © Ruslan Panin
Carabus (Ophiocarabus) ballionis Kraatz, 1879
Carabus (Ophiocarabus) ballionis Kraatz, 1879
Male adult
NW-China, Xinjiang prov., 40 km N Narat, h=1500, 12-12.VI.2004, Benes & Kaiser leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
Male adult
NE-China, E-Liaoning:Anshan:Mt.Qian Shan, 9-14.VI.2011
L=39 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Male adult
China, N-Sichuan, Jiuzhaigou env., 13.VI.2001,E. Kučera leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus pseudopinguis Heinz & Staven, 2000
Carabus (Lamprostus) syrus pseudopinguis Heinz & Staven, 2000
Male adult
NW-Jordan, Ajlun env., oak forests on dry limestone, h=897m, 32°21′33.1″N/ 035°43’47.4″E, 4.V.2010, Schnitter/Schellhorn leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) quinlani smetanai Häckel, 2011
Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Male+Female adult
C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
male, 21mm
female, 23 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus vlasuljensis Apfelbeck, 1894
Carabus (Megodontus) violaceus vlasuljensis Apfelbeck, 1894
Male adult
WC-Serbia, Borak - Gornje Babine, 1-10.VII.2007, M. Sima leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
Female adult
C-China, E Tibet,W-Sichuan prov., Kangding, Zheduo Pass h=4300m 30.VI.2005 , V.Benes leg,
L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Male adult
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar, 19.VII.2007, V.Benes leg.
L=17 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus azurescens Dejean, 1826 (loc.typ.)
Carabus (Megodontus) violaceus azurescens Dejean, 1826 (loc.typ.)
Male adult
W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill., VI.2002, Manfred Egger leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Apoplesius) pseudodepressus Machard, 1988
Carabus (Apoplesius) pseudodepressus Machard, 1988
Female adult
Turkey, Prov. Bitlis, S near Tatvan, 23.IV.2009 leg. M. & S. Nabozhenko
L=19.0 mm
Copyright © Kirill Makarov
Carabus (Rhigocarabus) ? roborowskii roborowskii Semenov, 1887
Carabus (Rhigocarabus) ? roborowskii roborowskii Semenov, 1887
Female adult
C-China, E Tibet, NW- Sichuan prov., Sêrxü, h=4290m, 20.VI.2005,, V.Benes leg.
L=18 мм
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus caelatus  (as malissorum Apfelbeck, 1919)
Carabus (Megodontus) caelatus caelatus (as malissorum Apfelbeck, 1919)
Male adult
Montenegro, National park Durmitor; near Zabljak vill., Mt. Sedlo h=2200 m, 6-22.VIII.2010, Franko Sandel leg.
29 мм
Copyright © Ruslan Panin
Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Male adult
China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus germarii Sturm, 1815
Carabus (Megodontus) violaceus germarii Sturm, 1815
Male adult
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m, 3-13.VI.2010, Manfred Egger leg.
29 mm
Copyright © Ruslan Panin
Calosoma (Carabomorphus) brachycerum (Gersteacker, 1884)
Calosoma (Carabomorphus) brachycerum (Gersteacker, 1884)
Male adult
N Tanzania, 105 km W of Arusha and 5 km S of Kwa Kuchinga vill. 3°41'S 25°59' E 31.VIII-2.IX.2006 leg. A.Sokolov
L=23 mm
Copyright © Kirill Makarov
Cychrus schmidti Chaudoir, 1837
Cychrus schmidti Chaudoir, 1837
Male adult
NW-Slovenia, Julian Alps,Mt. Mangart h=2670m, VIII.1995, Manfred Egger leg.
L=15 mm
Copyright © Ruslan Panin
Cychrus angustatus Hoppe & Hornschuch, 1825
Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
NC-Italy, Bergamo, Mt. Presolana h=1500m, Colere env., 4-15.VI.2010, Manfred Egger leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
W-UKRAINE, Transcarpathian region, Rakhiv distr., Chornogora Mt Range, h=800 m, near Bogdan vill., 10.VI.2012, Panin R. leg.
L= 33 мм
Copyright © Ruslan Panin
Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Male adult
China, E Tibet, N-Sichuan, (Barkam),valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E, mixed/coniferous forest, 22.VII.1995, K.& B.Brezina
L=23 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cristoforii Spence, 1823
Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček
Calosoma (Callisthenes) breviusculus Mannerheim, 1830Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Male adult
Macedonia, Bistra planina Mts.; 13mm
Copyright © Aleš Sedláček
Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Female adult
Macedonia, Bistra planina Mts.; 13mm
Copyright © Aleš Sedláček
Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Male adult
Macedonia, Bistra planina Mts.; 14mm
Copyright © Aleš Sedláček
Calosoma (Calosoma) sycophanta Linne, 1758
Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
CZ; S Moravia, Břeclav; 26mm
Copyright © Aleš Sedláček
Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
Female adult
NC Turkey, Gölköy env., Harcbeli pass, 37mm
Copyright © Aleš Sedláček
Carabus (Oreocarabus) cribratus staneki Sterba, 1931
Carabus (Oreocarabus) cribratus staneki Sterba, 1931
Male adult
NC Turkey, Küre Mts., 60km N Kastamonu; 25mm
Copyright © Aleš Sedláček
Carabus (Oreocarabus) cribratus staneki Sterba, 1931
Carabus (Oreocarabus) cribratus staneki Sterba, 1931
Female adult
NC Turkey, Küre Mts., 60km N Kastamonu; 27mm
Copyright © Aleš Sedláček
Carabus (Tachypus) auratus honnoratii Dejean, 1826
Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male adult
SE France, Carpentras; 23mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Male+Female adult
W-UKRAINE,
Transcarpathian region, Uzhhorod distr., h=250 m, 8.VII-25.VIII.2011
Panin R. leg./det.
female, 33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Male+Female adult
W-UKRAINE,
Transcarpathian region, Uzhhorod distr., 28.IV-23.V.2011
Panin R. leg./det.
female, 33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
L=30 mm
det.Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Female adult
NW Ukrainian Carpathians,
Lviv Region, Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.07, Panin R. leg./det.Solodovnikov I. & Panin R.
female, 35mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Female adult
NW Ukrainian Carpathians
Lviv Region, Staryi Sambor Distr., near Dobromyl , h=500m 9.VI-25.VII.2007, Panin R. leg./det.Solodovnikov I. & Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male adult
W.Ukraine, Ivano-Frankivsk reg.,
Yaremche distr., near Tatariv vill., Gorgany Mts. h=700 m, 16.VI.2011, Panin R. leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male+Female adult
W.UKRAINE, Ukrainian Carpathians
Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt. h=1000m. 9-20.VII.07 Panin R. leg.
female, 35mm.
Copyright © Ruslan Panin
Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013
Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Male adult
China, Xinjiang, Koeksu Bas., Tshon-Aidak, SSE Tekes, 42* 32' 47'' N / 82* 05' 56'' E -- 42* 32' 51'' N / 82* 05' 36'' E, 3330-3550 m, 11.07.2012, Kabak I.I. leg.
Copyright © Ilia Kabak
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male adult
NE Serbia, Majdanpek; male, 33mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Kučevo; female, 33mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Majdanpek; 35mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 24mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 25mm
Copyright © Aleš Sedláček
Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Female adult
as ativsengueni Schweiger, 1962
NW Turkey, Bolu, Abant Mts.; 16mm

Copyright © Aleš Sedláček
Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Male adult
Canarias Isle, NC Tenerife; 22mm
Copyright © Aleš Sedláček
Carabus (Carabus) arvensis venetianus Bernau, 1914
Carabus (Carabus) arvensis venetianus Bernau, 1914
Male adult
N Italy, Veneto, Dolomiten Mts.; 13mm
Copyright © Aleš Sedláček
Carabus (Carabus) arvensis venetianus Bernau, 1914
Carabus (Carabus) arvensis venetianus Bernau, 1914
Female adult
N Italy, Veneto, Dolomiten Mts.; 15mm
Copyright © Aleš Sedláček
Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
Female adult
CZ, C Moravia, Dřevohostice - Líšná, 13mm
Copyright © Aleš Sedláček
Carabus (Procrustes) banoni Dejean, 1829
Carabus (Procrustes) banoni Dejean, 1829
Male adult
Crete, Agia Irini (Sélinos); 31mm
Copyright © Aleš Sedláček
Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Female adult
SE Spain, Murcia reg., P.N. de Calblanque, 1.V.2003, T.Gazurek leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) lafossei coelestis Stewart, 1845
Carabus (Coptolabrus) lafossei coelestis Stewart, 1845
Female adult
CE-China, Shangai, 3.V.2007/ det.Panin R.
L=34 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Male adult
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R.<br /> L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) baguenai Breuning, 1926
Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
SE Spain, Almería Province, Carboneras env., 25.XI.2006, T.Gazurek leg./det. R.Panin <br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Female adult
SW-Portugal, Setúbal Distr., Cercal do Alentejo/ Tanganheira vill. 8.XII.2006, T.Gazurek leg.
L=31 m
Copyright © Ruslan Panin
Carabus (Archicarabus) steuartii Deyrolle, 1852
Carabus (Archicarabus) steuartii Deyrolle, 1852
Female adult
NW-Spain, Pontevedra prov., Serra do Candán Mt range, 1.III.1992, S. Devesa leg./det. S. Sobrino
Copyright © Ruslan Panin
Carabus (Ctenocarabus) galicianus Gory, 1839
Carabus (Ctenocarabus) galicianus Gory, 1839
Female adult
NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Female adult
C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male adult
S Austria, Karnische Alpen; male, 24mm
Copyright © Aleš Sedláček
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Female adult
S Austria, Karnische Alpen; 25mm
Copyright © Aleš Sedláček
Carabus (Mesocarabus) macrocephalus Dejean, 1826
Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m, 25.IX.2004,T.Gazurek leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis deyrollei Gory, 1839
Carabus (Carabus) arvensis deyrollei Gory, 1839
Male adult
NW-Spain,
Cantabria, Udías env., Monte Corona h=100m,14.VI.2002, Javier de Diego leg.
L=16 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris adamellicola Ganglbauer, 1904
Carabus (Orinocarabus) alpestris adamellicola Ganglbauer, 1904
Male adult
N-Italy, Brescia prov., Bagolino, Adamello Mts, Passo del Maniva env., h=2000m, VIII.2011, M.Monzini leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
Male adult
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m, 7.II.2009, T.Gazurek leg./det.R. Panin.
L=24 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
Female adult
W Bulgaria, Cerovo, 31 mm
Copyright © Aleš Sedláček
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Female adult
NW Slovenia, Trnovski Gozd; 24 mm
Copyright © Aleš Sedláček
Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
Male adult
N Italy, Alpi Graie; 20 mm
Copyright © Aleš Sedláček
Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
Female adult
W-Ireland, County Mayo, Letterneevoge (R312),16-30.VI.2013, J.Stypula leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) fairmairei heteromorphus Daniel, 1896
Carabus (Orinocarabus) fairmairei heteromorphus Daniel, 1896
Female adult
NW-Italy, Aosta region, Champorcher env., Mt.Dondenaz, VIII.2008, V.Monzini leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Female adult
SE-France, Var, near Le Muy, VIII.1980, Michon leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847 (loc.typ.)
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847 (loc.typ.)
Male adult
S-Ukraine, near Dnipropetrovsk, 6-15.IX.2013, A. Sukhenko leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Female adult
N-Japan, SW-Hokkaido, Chitose env.,VI.2011.
L=34 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) planatus Chaudoir, 1843
Carabus (Macrothorax) planatus Chaudoir, 1843
Male adult
S Italy, N Sicily, Nebrodi; 33 mm
Copyright © Aleš Sedláček
Carabus (Orinocarabus) fairmairei fenestrellanus Beuthin, 1899
Carabus (Orinocarabus) fairmairei fenestrellanus Beuthin, 1899
Male adult
NW-Italy, Piemonte prov., Cottian Alps, Mt Albergian h=2450 m.
L=20 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
Female adult
SE-France, Var, Fréjus env., 26.II.1997,
L=31 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus ?harcyniae (as procedens Csiki, 1927)
Carabus (Mesocarabus) problematicus ?harcyniae (as procedens Csiki, 1927)
Male adult
W-Ireland, County Mayo, Clare Isl., 17-28.VI.2002, S. McCormack leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus ?harcyniae (as procedens Csiki, 1927)
Carabus (Mesocarabus) problematicus ?harcyniae (as procedens Csiki, 1927)
Female adult
N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg.
L=27mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
Male adult
W-FRANCE, Brittany , Finistère, Botvarec Forest, 19.XI.2006, Pascal Stefani leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Male adult
S Greece, Taygetos Mts.; 13mm
Copyright © Aleš Sedláček
Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Female adult
SW Romania, Banat; 15mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male adult
SW Romania, Banat - Eibenthal; 29 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Female adult
SW Romania, Banat - Eibenthal; 33 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE Serbia, Majdanpek; 23 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Female adult
NE Serbia, Majdanpek; 29 mm
Copyright © Aleš Sedláček
Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male+Female adult
KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992/ det.Panin R.
female 19mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896 (as carpathicus Kolbe, 1913)
Carabus (Tachypus) cancellatus durus Reitter, 1896 (as carpathicus Kolbe, 1913)
Male+Female adult
Ukrainian Carpathians,
Ivano-Frankivsk reg., Gorgany Mts., 20km SW Nadvorna, 22.VI-3.VIII.20Panin R. leg.
female, 26mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896 (as ungensis Csiki, 1905)
Carabus (Tachypus) cancellatus durus Reitter, 1896 (as ungensis Csiki, 1905)
Male+Female adult
W Ukraine,
Transcarpathian reg., Vinogradov distr., near Malaya Kopanya vill.,7-18.VII.2008, Panin R. leg.
female, 32mm.
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896 (? hybrid)
Carabus (Tachypus) cancellatus durus Reitter, 1896 (? hybrid)
Female adult
W. UKRAINE, NE Carpathians,
Marmarosh Mts h=450m, Rakhiv distr., near Dilove vill.,19-23.VII.2010, Panin R. leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis kodaricola Deuve, 1998
Carabus (Aulonocarabus) truncaticollis kodaricola Deuve, 1998
Female adult
SE-Siberia:N-Chita reg., Kodar Mt.Rng.,h=1800m, Leprindo vill., 1.VII.2006,
L=17 mm
Copyright © Ruslan Panin
Carabus (Cratocephalus) solskyi Ballion, 1878
Carabus (Cratocephalus) solskyi Ballion, 1878
Female adult
SE-KAZAKHSTAN, Ketmen Mt Rng, h=1800m, Tujuk vill.,15.V.2004
L=36 mm
Copyright © Ruslan Panin
Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Male adult
N-Kyrgyzstan, Kyungey Ala-Too Mt Rng, Cholpon-Ata env., h=2500m, 19.VI.2000,
L=19 mm
Copyright © Ruslan Panin
Cychrus attenuatus attenuatus s.str.
Cychrus attenuatus attenuatus s.str.
Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 13mm
Copyright © Aleš Sedláček
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Male adult
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env, h=1100m, 24.V.1998, /det. Panin R.
22 mm

Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg./det. R.Panin
L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Male adult
NW-Hungary::Komarom distr., Tardos, Gerecse Mts., 20.-27.V.2011, Krause Rene leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Male adult
NW Hungary: Fejer: Vértes Mt Rng, Tatabanya, 28.VII-2.VIII.1996
L=29 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus tenuiorithorax Deuve & Li, 2006
Carabus (Coptolabrus) augustus tenuiorithorax Deuve & Li, 2006
Male+Female adult
SE-China, SW-Jiangxi, Mt Guan, V.2009, local coll.
female, 47 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
Male adult
C-China, S-Gansu prov., Wenxian County, Mt.Guanzigou,1-14.V.2010, local coll. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens Fabricius, 1798 (as moldaviensis Born, 1903)
Carabus (Morphocarabus) excellens Fabricius, 1798 (as moldaviensis Born, 1903)
Male adult
E-Romania, Vaslui reg.,15 km SE Vaslui, Crasna env., V-VI,2013, M,Pasek leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
Male adult
N-Korea, North Pyongan Prov., Tongnim County, 5-25.V.2010
29 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
Male adult
NC-Korea, Ryanggang Prov., Puksubaek-san, 4-10.V.2010
28 mm

Copyright © Ruslan Panin
Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924Carabus (Tachypus) vagans Olivier, 1795
Carabus (Tachypus) vagans Olivier, 1795
Female adult
SE-France, Var: Le Muy env., 30.VII.2008., A. Paulian leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
Female adult
NW-Morocco, betw.Meknes/Kenitra, Foret de Mamora, II.2008
L= 27 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
Male adult
SE-China, Jiangxi prov.,Yushan County, Mt Wangjing, VI.2012
40 мм
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
Male adult
NC-Spain, Sierra de Cantabria , Sierra de Toloño Rng, Puerto de Herrera , h=1100m, 19.I,2007 25 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) pustulifer cyanelongatus Deuve, 2006
Carabus (Coptolabrus) pustulifer cyanelongatus Deuve, 2006
Male adult
C-Chine, Tibet, Qamdo, Gonjo County, Ke Ri Xiang env., VI.2012, Alain Mollard leg.
L=43 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Male adult
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m, X.2010, A. Mollard leg.<br /> L=23 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Male adult
CW-Spain: Salamanca, Brincones env., h=750m, X.2003<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Male adult
SE-China, Jiangxi prov., Yushan County, Mt Wangjing, VI,2012
L=38 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Male adult
Luxembourg, Bollendorf-Pont env., 3.III.1999 /det.R Panin
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Male adult
NE Spain, Catalonia, Girona Prov., San Hilario Sacalm env., 20.II.1988, A. Mollard leg.<br /> L=25 m
Copyright © Ruslan Panin
Carabus (Orinocarabus) bertolinii Kraatz, 1878
Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Dolomites-Alps, Trentino prov., Rolle Pass env., Malga Fosse h=2000m, 8-28.VI.2013, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
Female adult
C-China, S-Shaanxi: Shangzhou City, Shahezi,1-14.VI.,2013, local coll. leg.
L=39 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus dolichognathus Deuve, 1995
Carabus (Coptolabrus) smaragdinus dolichognathus Deuve, 1995
Male adult
NW-Korea, North Pyongan Province, Mt. Donggorim, 1-20.VI.2013, local coll. leg.
L=37 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus augustus Bates, 1888
Carabus (Coptolabrus) augustus augustus Bates, 1888
Male adult
SE-China, Jiangxi prov., Yudu County, Ganzhou city, Yun Fengzang h=1000m, 5-20.V.2011
L=43 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
Male adult
NE-Italy, South Tyrol, Dolomites-Alps, 500 m left of Würzjoch, Gömaalm - at the foot Mt.Peitlerkofel h=2100m, 28.VII-17.VIII.2013, Manfred Egger leg.
l=18 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) monticola roccae Born, 1923
Carabus (Archicarabus) monticola roccae Born, 1923
Male adult
NW-Italy, Liguria:Savona env., Monte San Giorgio, 30.X.1977, Monauzzi leg.
L=23 mm

Copyright © Ruslan Panin
Carabus (Orinocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Carabus (Orinocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Female adult
W Ukraine, NW-Podolian, near Lviv city, Vynnykivskyi forest, 20.IV.-13.V.2011, R. Panin leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) linnaei prope linnaei Panzer, 1812
Carabus (Orinocarabus) linnaei prope linnaei Panzer, 1812
Male+Female adult
W-Ukraine, Transcarpathian reg., Chornogora Mt Rng, 8 km NE Bohdan vill., left-bank terrace the Bohdan river h=800m, 8.VII-24.VIII.2012, Panin R. leg. female, 21 mm
Copyright © Ruslan Panin
Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
Male adult
W-Caucasus: Abkhasia, Gudauta distr., E slope Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
25 mm

Copyright © Ruslan Panin
Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Male adult
Spain, Canarias Isl.,Teneriffe: NW-part:Teno Massif, Erjos env., h=1000m, 18-21.III.2013, V.Slovak leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
Female adult
C Romania, Sibiu reg., left bank Olt River, Avrig env., 12.VI.2013, V. Slovak leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Male adult
N-Japan, Hokkaido, Katō District,Kamishihoro, Horoka near Maruyama Bridge,19.VI.2013, T. Hirabayashi leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Female adult
NW-Caucasus, Karachay-Cherkessia, Bol'shaya Marka River h=2500-2600m, 12.VIII.2010, I.A. Solodovnikov leg./det.
Copyright © Ruslan Panin
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Male adult
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
L=25 mm
Copyright © Ruslan Panin
Carabus (Tribax) agnatus agnatus Ganglbauer, 1889Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
NС-Caucasus, Karachaevo-Cherkessia, Abishir-Akhuba Mt.Range h=2250-2600 m, right bank Pkhiya river, 10-13.VI.2013, S. Pavluchuk leg.
L=25 mm
Copyright © Ruslan Panin
Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Male adult
NW-USA, Washington, Thurston County, Douglas fir forest, 15.X.2005, A. Laforgue leg. L=21mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
Male adult
NW-USA, Washington, Thurston County, Mima Mound Prairie, 15.IX.2005, A. Laforgue leg. L=23 мм
Copyright © Ruslan Panin
Carabus (Eucarabus) catenulatus   fontanellae Reitter, 1896
Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Cres island,VIII.1994, G. Zannna leg. L=33 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) cartereti omisanensis Lassalle, 1999
Carabus (Eucarabus) cartereti omisanensis Lassalle, 1999
Male adult
South Korea, Gangwon Province, Kyebang-san, h=1100m, VI.1998 L=24 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Male adult
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812Calosoma (Ctenosta) guineense Imhoff, 1843
Calosoma (Ctenosta) guineense Imhoff, 1843
Male adult
Rep. Centre Africaine: Bangui env. IV.1974 28 mm
Copyright © Ruslan Panin
Calosoma (Calosoma) frigidum Kirby, 1837: 19
Calosoma (Calosoma) frigidum Kirby, 1837: 19
Female adult
SC-Canada, Manitoba prov., Winnipeg, King’s Park, 23.VI.2008, Todd Lawton leg. L=24 mm
Copyright © Ruslan Panin
Calosoma (Campalita) chinense Kirby, 1819: 379
Calosoma (Campalita) chinense Kirby, 1819: 379
Female adult
North Korea, Ryanggang Province, Mt P'urun-bong, 24.VIII.1994 L=33 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., between Chagari-Zbarazhki & Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859Carabus (Carabus) granulatus yezoensis Bates, 1883Carabus (Carabus) menetriesi menetriesi Hummel, 1827Carabus (Diocarabus) aurocinctus Motschulsky, 1846Carabus (Diocarabus) opaculus opaculus Putzeys, 1875Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)Carabus (Morphocarabus) henningi henningi Fischer, 1817Carabus (Morphocarabus) venustus venustus Morawitz, 1862Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as laevipennis Seidlitz, 1891)
Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as laevipennis Seidlitz, 1891)
Male adult
W Ukraine, NE Carpathians,Transcarpathian Region, Rakhiv Distr., Chornogora Mts Range, h=2000 m, alpin zone, V-28.VI..2014, Panin R. leg.
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Region,Chornogora Mt Range, h=2000 m, alpin zone, 28.VI-11.IX..2014, Panin R. leg. L=22 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) reitteri reitteri Retowski, 1885Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
Male adult
SC-Turkey: Alania (Irmasan), rte 695 au km 25 Akseki, h= 1225 m, 02.X.2003
Copyright © Igor Solodovnikov
Carabus (Imaibius) wittmerorum Heinertz, 1978
Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult
N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg.S.Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
Male adult
N-Pakistan: Mansehra prov., Hazara, Kaghan valley: Shocgran env., h = 2750 m L= 42 mm
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides fortunei Adams, 1861
Carabus (Damaster) blaptoides fortunei Adams, 1861
Male adult
Japan: C Honshu: Niigata pref., Mysato: Sakae Abiko-City; Iwafune distr., Awashima Isl.
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides fortunei Adams, 1861
Carabus (Damaster) blaptoides fortunei Adams, 1861
Female adult
Japan: C Honshu: Niigata pref., Mysato: Sakae Abiko-City; Iwafune distr., Awashima Isl.
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides capito Lewis, 1881
Carabus (Damaster) blaptoides capito Lewis, 1881
Male adult
Japan: Niigato pref., Sado Isl., Sado, Yamato
Copyright © Igor Solodovnikov
Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
Male adult
Kyrgyzstan, W Kungei Ala-Too Mt Rng, Chon-Kemin riv., Kyzyl-Bairak vill., 25.IV.2010. L= 18 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) pumilio Kuster, 1846
Carabus (Mimocarabus) pumilio Kuster, 1846
Male adult
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m, 9.VI.2014, L=16 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) imitator imitator Reitter, 1883Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male+Female adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) edithae Reitter, 1893
Carabus (Neoplectes) edithae Reitter, 1893
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Scaphinotus (Scaphinotus) viduus Dejean, 1826
Scaphinotus (Scaphinotus) viduus Dejean, 1826
Male adult
E-USA, West Virginia, Pendleton Co., Allegheny Mts. Mt. Spruce Knob, 20-27.VI.2012, T. Lawton leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) staehlini Adams, 1817
Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
S Russia, E-Caucasus, Dagestan, Nukatl Mt Rng h=2600m, 25.VII.1997, V. Savizky leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Female adult
SE-Spain, Prov.Alicante, Port de la Carrasqueta, 26.XII.1987, M.Toribio leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens (as rareulensis Born, 1907)
Carabus (Morphocarabus) excellens (as rareulensis Born, 1907)
Male adult
NE-Romania, Bucovina, Rarau Mts. h-1300m, VI.2014, J. Kynkor leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Imaibius) boysi boysi Tatum, 1851
Carabus (Imaibius) boysi boysi Tatum, 1851
Male adult
NW-India, Himachal Pradesh, Manikaran env., Pulga vill., 4.VII.2008. L=26 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Holotype Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Eucarabus) obsoletus nagyagensis (as fossulifer Fleischer, 1893)
Carabus (Eucarabus) obsoletus nagyagensis (as fossulifer Fleischer, 1893)
Male adult
NW-Romania, Bihor, Mts Seș (Plopis), 20 km N Aleșd, Pădurea Neagră env., VI.2014, J.Kynkor leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=32 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
Female adult
China, Tibet AR, Qamdo, Gonjo County, Kè Rì Xiāng vill.(Kerri Township), h=3100m, VI.2012, A. Mollard leg. L=38 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
Male adult
SE France, Alpes-Maritimes, Isola env., Colle Santa-Anna h=2309 m, 44*13',127" N, 07*06',242"E, 31-7.VIII..2012. L=21 mm
Copyright © Ruslan Panin
Carabus (Iniopachus) pyrenaeus pyrenaeus  (as cephalicus Csiki, 1927)
Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
NC-Spain, Cantabria, Santander, Cueto env.XII.2013 L=28 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 (ab. holochrysum)
Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 (ab. holochrysum)
Female adult
SC-FRANCE, Tarn, Le Linas h=550 m, XI.2008, Pascal Stefani leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus mixtus (as neomixtus Tarrier, 1975)
Carabus (Megodontus) violaceus mixtus (as neomixtus Tarrier, 1975)
Male adult
L=29mm SE-France, Drôme, Foret de Saou, VII.2014, Pascal Stefani leg.
Copyright © Ruslan Panin
Ceroglossus chilensis villaricensis Kraatz-Koschlau, 1885
Ceroglossus chilensis villaricensis Kraatz-Koschlau, 1885
Male adult
CHILE, Villarica, local collector leg. 26 мм.
Copyright © Sergey Kakunin
Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) phoenix Lapouge, 1924
Carabus (Archicarabus) phoenix Lapouge, 1924
Male+Female adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. female,23mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
Male adult
Russsia, Altay, Ukok upland, Kaldzhin-Kul Lake h=2400-2550 m, 49 *19*N/87*26*E, 16-27.VII.2008, R.Dudko leg. L=17.5 mm
Copyright © Ruslan Panin
Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Male adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Male adult
NE-Turkey:NE of Artvin:Dalis Dağ ,h=1600-1800m, 8-18.VII.2014, C. Reuter leg. L= 25 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
Female adult
W.UKRAINE, NE Carpathians, Lviv Reg., Skole distr., Zelemyanka Mt.Range h=700m, 26.V-1.VII.2010, Panin R. leg. L=43 mm
Copyright © Ruslan Panin
Calosoma (Orinodromus) deckeni (Gersteacker, 1867)
Calosoma (Orinodromus) deckeni (Gersteacker, 1867)
Male adult
04/1912, Tanzania, Mount Kilimanjaro, Bismarck Hügel, 2700m. Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
Copyright © - - -
Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
Male adult
Spain, Jaen prov., Despeñaperros, Arroyo de los Arcos, 38.381847, -3.515651, 1-2.05.2014, Anichtchenko A. leg. & det.; 18 mm
Copyright © Alexander Anichtchenko
Carabus (Limnocarabus) clathratus clathratus Linne, 1761Carabus (Euleptocarabus) porrecticollis Bates, 1883
Carabus (Euleptocarabus) porrecticollis Bates, 1883
Female adult
Atsushiokanomachikano, Kitakata City, Fukushima Pref., Japan, 29.III.2014, T. Nakamura leg.
Copyright © Alexander Anichtchenko
Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 3-12..III.2015, B.Vas’ko leg./det. Panin R.
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 13.XII.2012, Vas’ko, leg./det. Panin R. L=30мм
Copyright © Ruslan Panin
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Male adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
Female adult
SC-China, Guangxi Reg., Pingle County, Mt. Shizishan h=1290 m,5-20,VII.2011, local coll. leg. L=43 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cavernosus variolatus Costa, 1839
Carabus (Pachystus) cavernosus variolatus Costa, 1839
Male adult
C-Italy:Abruzzo reg., Gran Sasso d'Italia Massif, h~1600 m, VIII.1997, Seichert leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Carabus) stscheglowi Mannerheim, 1827
Carabus (Carabus) stscheglowi Mannerheim, 1827
Male adult
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka, L=27 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
Male+Female adult
E-Netherlands, Gelderland, Winterswijk Woold env.,15.II.2015, H.de Haan leg.
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
Copyright © Noël Mal
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Habitat
Andorra, Anyos, ii.2012, J. Nicolas Leg. & Photos
Copyright © Noël Mal
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Andorra, Leg. Gerts, vi.1966, 32 mm
Copyright © Noël Mal
Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997
Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Paratype
Male adult
Chine, C-Fujian: SE of Yong’an, Mt.Tianbaoyan, 25°58’ N / 117°22’ E, h = 1100 m, VII.1997, leg. Wong
Copyright © Igor Solodovnikov
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
Copyright © Noël Mal
Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Female adult
Portugal, Ribatejo, Coruche, 13.x.1974, A. Zuzarte leg., 25 mm, Coll. N. Mal
Copyright © Noël Mal
Calosoma (Calosoma) sycophanta Linne, 1758
Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
Turkey : Antalya, Yukarikaraman 3.iv.2000 - M. Grotz Leg. (Size : 33 mm) Coll. N. Mal
Copyright © Noël Mal
Calosoma (Calosoma) sycophanta Linne, 1758
Calosoma (Calosoma) sycophanta Linne, 1758
Female adult
Turkey : Antalya, Yukarikaraman 3.iv.2000 - M. Grotz Leg. (Size : 33 mm) Coll. N. Mal
Copyright © Noël Mal
Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Female adult
NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis carpathus Born, 1902
Carabus (Carabus) arvensis carpathus Born, 1902
Male adult
W Ukraine, NE Carpathians,Transcarpathian reg., Vihorlat, near Strychava vill., Mt Knyaginytsya h= 500 m, 18.V-22.VI. 2014, Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis arvensis Herbst, 1784
Carabus (Carabus) arvensis arvensis Herbst, 1784
Female adult
NW Ukraine, Polesia, Rovno reg., Rokytne distr., near Bil’sk vill., 4-6.VII.2015, Panin R. leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Female adult
China : Guizhoh, Sinan vi.2006 (Size : 35 mm) Coll. N. Mal
Copyright © Noël Mal
Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Female adult
China : Shaanxi, Shazhou City, Shahezi 10.vii.2007 - J. Li Leg. (Size : 42 mm), Coll. N. Mal
Copyright © Noël Mal
Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812
Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812
Male adult
Hungary, Budapest, Mt Buda, Harsbokor-hill,16-19.V.1992, I.Retezár leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő), 8.VI.1992, leg./det. I.Retezár leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
Male adult
NC-Hungary: E- part of Budapest, Cinkota,28.VIII-17.IX.1980, I.Retezár leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
Female adult
W-Hungary, NE-Alps, Mts. Kőszeg, Velem env., Borha-forrás, 26.VII.1981, Gy. Somorijai leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus kvirensis Belousov, 1992
Carabus (Pachycarabus) swaneticus kvirensis Belousov, 1992
Male adult
SC-Caucasus: NW-Georgia, Mingrelia, Egrissky Mt.Rng., h~1800-2000 m, 23-26.VII.1990, I.Retezár leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013
Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013 Paratype
Male adult
Romania, Apuseni : Trascău Mts. L=27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Male adult
C-Slovakia, Detva Distr., Stara Huta env. h=700-800m, 3.VI.-5.VII.2012, V.Slovák lgt. L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
Male+Female adult
NC-Hungary:Borzsony Mts, Nograd-Diosjeno,1-14.V.2004, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Female adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env., h~1800 m., L=21 mm
Copyright © Ruslan Panin
Calosoma (Ctenosta) grandidieri Maindron, 1900
Calosoma (Ctenosta) grandidieri Maindron, 1900
Female adult
MADAGASCAR, Tulear Prov., nr. Tsimanampetsotsa NP, P. Schüle det.
Copyright © Peter Schüle
Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966, loc.typ.
Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966, loc.typ.
Male adult
NW-Turkey:Bolu Prov., Abant Lake, 25.VIII.1990 L=35 mm
Copyright © Ruslan Panin
Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
Female adult
NE Pakistan, Azad Kashmir, Bagh Distr., Sudhan-Gali h~2300 m,15.VIII.1994, S Dacatra leg. L=37 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
Female adult
S-China, Guangxi, Jinxiu Co., Mt. Dayao,28.VII.2007, L=40 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Male adult
NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor lepontinus Born, 1908
Carabus (Orinocarabus) concolor lepontinus Born, 1908
Male adult
NW-Italy, Piemonte (VB), Lepontine Alps, Mt. Zeda , 6.VII.1998, L Diotti leg., L=20 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Female adult
NW-Italy, Breuil-Cervinia h~2500 m, 2.IX.1971, Georgeon leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
Male adult
NW-Turkey,Istanbul (Asian part),Polonezköy env., 2.VI.2009. S. Dacatra leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
C China, Qinghai, Zadoi County, Toramarkog h~4600 m, VI.2001, P. Cavazzuti leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) pedemontanus germanae Casale & Cavazzuti, 1977
Carabus (Orinocarabus) pedemontanus germanae Casale & Cavazzuti, 1977
Male adult
NW-Italy, Piemonte, Imperia prov., Mt.Saccarello h~2000m, 30.V.1986. L=21 mm
Copyright © Ruslan Panin
Calosoma (Castrida) leleuporum (Basilewsky, 1968)
Calosoma (Castrida) leleuporum (Basilewsky, 1968) Holotype
Male adult
Picture: Camille Locatelli, Copyright: RBINS / DIGIT-3 Belspo Licence: CC BY NC ND http://virtualcollections.naturalsciences.be
Copyright © - - -
Carabus (Orinocarabus) fairmairei pelvicus Cavazzuti, 1973
Carabus (Orinocarabus) fairmairei pelvicus Cavazzuti, 1973
Male adult
NW-Italy, Cuneo, Maira Vall., Acceglio env., La Colletta h~2800 m, 12.VII.2007, R. Monguzzi leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
S-China, N-Guangxi, Mao'er Shan, h~1600 m, V.2009, L= 26 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) italicus rostagnoi Luigioni, 1904
Carabus (Eucarabus) italicus rostagnoi Luigioni, 1904
Female adult
C-Italy, Lazio, Fondi env.,10.II.1979, L = 25 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) victor victor Fischer, 1836
Carabus (Archicarabus) victor victor Fischer, 1836
Female adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m, 23.VII.2001, S. Dacatra leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus giresuni Breuning, 1964 (as paululus Heinz & Korge, 1964)
Carabus (Lamprostus) torosus giresuni Breuning, 1964 (as paululus Heinz & Korge, 1964)
Male adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m,24.VII.2001, S. Dacatra leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri baldensis Schaum, 1857
Carabus (Platycarabus) creutzeri baldensis Schaum, 1857
Male adult
NC-Italy: Trento, Sega di Ala, IX.1999, A. Vicini leg. , L=24 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) rossii Dejean, 1826
Carabus (Archicarabus) rossii Dejean, 1826
Female adult
NC Italy, Emilia reg., Bologna, San Martino, 29.V.2016, G..L.Agnoli leg. L=30 mm
Copyright © Ruslan Panin
Cychrus (Cychrus) morawitzi morawitziCarabus (Orinocarabus) linnaei folgariacus Born, 1913
Carabus (Orinocarabus) linnaei folgariacus Born, 1913
Male adult
NE-Italy, Trentino-Alto Adige (Südtirol), N of vill. San Michele vill., 13-17.VI.2016, R. Janak leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens kraussi Lapouge, 1898
Carabus (Chrysocarabus) auronitens kraussi Lapouge, 1898
Female adult
C-Austria, Styria, Ennstal Alps, Totes Gebirge Mt Rng, Mt Loser h~1100 m, 16-20.VII.2014, R.Janak leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) adamsi Adams, 1817
Carabus (Sphodristocarabus) adamsi Adams, 1817
Male adult
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env., 20.VI.2015, F. Grycz leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) ignimitella guangxicola Deuve, 1998
Carabus (Coptolabrus) ignimitella guangxicola Deuve, 1998
Male adult
S-China, Guangxi prov., Jingxiu, Mt. Dayao Shan, 1-17.V.2006, L=47 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) robustus triumphans Cavazzuti, 1987
Carabus (Lamprostus) robustus triumphans Cavazzuti, 1987
Male adult
NE-Turkey:Trabzon, Tonya env., VIII.2004. L=41 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) latreilleanus Csiki, 1927
Carabus (Orinocarabus) latreilleanus Csiki, 1927
Female adult
NW-Italy, Piemonte, Biella, Biellese Alps Rng, Mt.Camino, 25.VI.1997, V.Monzini leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
Male adult
NE-Turkey:Ordu prov.,3 km N Gölköy h~1100 m, VII.2006, L=37 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus lampedusae Born, 1925
Carabus (Macrothorax) morbillosus lampedusae Born, 1925
Male adult
S Italy, Isl. Lampedusa, Baia dei Conigli, VI.2003, L=29 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Female adult
Abkhazia, Bzybsky Mt.Rng, right bank of river Reshevie (left tributary of river Bzyb) h~670-720 м, 13-14.V.2010, D.Fominykh & I.Solodovnikov leg. L=41 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) breuningi Csiki, 1927
Carabus (Mesocarabus) breuningi Csiki, 1927
Female adult
NW-Spain, Prov. Zamora, San Martín de Castañeda, 5.VII.1984, M.Toribio leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Male adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg., 17 mm
Copyright © Ruslan Panin
Synonyms
Countries and regions
Museums
Description
Biotope
Genotype
Level down
Login:
Password:
Register | Forgot your password?
ВКонтакте Facebook Google Accounts Twitter Yandex Mail.ru OpenID WebMoney

Support our efforts

This project help increase the availability of scientific knowledge worldwide. Contributions at any level help sustain our work. Thank you for your support.
Sitemap
Powered by CMS Eleanor © 2017
Page generated in 1.438 seconds. Make queries: 14. Memory: 2.095 MB